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Autogamy
Autogamy or self-fertilization refers to the Cell fusion, fusion of two gametes that come from one individual. Autogamy is predominantly observed in the form of self-pollination, a Reproduction, reproductive mechanism employed by many flowering plants. However, species of protists have also been observed using autogamy as a means of reproduction. Flowering plants engage in autogamy regularly, while the protists that engage in autogamy only do so in stressful environments. Occurrence Protists ''Paramecium aurelia'' ''Paramecium aurelia'' is the most commonly studied protozoan for autogamy. Similar to other unicellular organisms, ''Paramecium aurelia'' typically reproduce asexually via Fission (biology), binary fission or sexually via cross-fertilization. However, studies have shown that when put under nutritional stress, ''Paramecium aurelia'' will undergo meiosis and subsequent fusion of Gamete, gametic-like nuclei. This process, defined as hemixis, a chromosomal rearrangement pro ...
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Paramecium
''Paramecium'' ( , , plural "paramecia" only when used as a Common name, vernacular name) is a genus of eukaryotic, unicellular ciliates, widespread in freshwater, brackish, and Ocean, marine environments. Paramecia are often abundant in stagnant basins and ponds. Because some species are readily cultivated and easily induced to sexual conjugation, conjugate and divide, they have been widely used in classrooms and laboratories to study biological processes. ''Paramecium'' species are commonly studied as model organisms of the ciliate group and have been characterized as the "laboratory rat, white rats" of the phylum Ciliophora. Historical background ''Paramecium'' were among the first ciliates to be observed by microscopy, microscopists, in the late 17th century. They were most likely known to the Dutch pioneer of protozoology, Antonie van Leeuwenhoek, and were clearly described by his contemporary Christiaan Huygens in a letter from 1678. The earliest known illustration of a ...
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Ciliate
The ciliates are a group of alveolates characterized by the presence of hair-like organelles called cilia, which are identical in structure to flagellum, eukaryotic flagella, but are in general shorter and present in much larger numbers, with a different wikt:undulating, undulating pattern than flagella. Cilia occur in all members of the group (although the peculiar Suctoria only have them for part of their biological life cycle, life cycle) and are variously used in swimming, crawling, attachment, feeding, and sensation. Ciliates are an important group of protists, common almost anywhere there is water—in lakes, ponds, oceans, rivers, and soils, including anoxic and oxygen-depleted habitats. About 4,500 unique free-living species have been described, and the potential number of extant species is estimated at 27,000–40,000. Included in this number are many Ectosymbiosis, ectosymbiotic and endosymbiotic species, as well as some Obligate parasite, obligate and Facultative paras ...
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Reproduction
Reproduction (or procreation or breeding) is the biological process by which new individual organisms – "offspring" – are produced from their "parent" or parents. There are two forms of reproduction: Asexual reproduction, asexual and Sexual reproduction, sexual. In asexual reproduction, an organism can reproduce without the involvement of another organism. Asexual reproduction is not limited to unicellular organism, single-celled organisms. The cloning of an organism is a form of asexual reproduction. By asexual reproduction, an organism creates a genetically similar or identical copy of itself. The evolution of sexual reproduction is a major puzzle for biologists. The two-fold cost of sexual reproduction is that only 50% of organisms reproduce and organisms only pass on 50% of their genes.John Maynard Smith ''The Evolution of Sex'' 1978. Sexual reproduction typically requires the sexual interaction of two specialized reproductive cells, called gametes, which contain half t ...
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Self-pollination
Self-pollination is a form of pollination in which pollen arrives at the stigma of a flower (in flowering plants) or at the ovule (in gymnosperms) of the same plant. The term cross-pollination is used for the opposite case, where pollen from one plant moves to a different plant. There are two types of self-pollination: in autogamy, pollen is transferred to the stigma of the same flower; in geitonogamy, pollen is transferred from the anther of one flower to the stigma of another flower on the same flowering plant, or from microsporangium to ovule within a single ( monoecious) gymnosperm. Some plants have mechanisms that ensure autogamy, such as flowers that do not open ( cleistogamy), or stamens that move to come into contact with the stigma. The term selfing that is often used as a synonym is not limited to self-pollination, but also applies to other types of self-fertilization. Occurrence Few plants self-pollinate without the aid of pollen vectors (such as wind or ...
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Geitonogamy
Geitonogamy (from Greek ''geiton'' (γείτων) = neighbor + ''gamein'' (γαμεῖν) = to marry) is a type of self-pollination. Geitonogamous pollination is sometimes distinguished from the fertilizations that can result from it, geitonogamy. If a plant is self-incompatible, geitonogamy can reduce seed production. Geitonogamy is when pollen is exported using a vector (pollinator or wind) out of one flower but only to another flower on the same plant. It is a form of self-fertilization. In flowering plants, pollen is transferred from a flower to another flower on the same plant, and in animal pollinated systems this is accomplished by a pollinator visiting multiple flowers on the same plant. Geitonogamy is also possible within species that are wind-pollinated, and may actually be a quite common source of self-fertilized seeds in self-compatible species. It also occurs in monoecious gymnosperms. Although geitonogamy is functionally cross-pollination involving a pollinating ag ...
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Paramecium Aurelia
''Paramecium aurelia'' are unicellular organisms belonging to the genus ''Paramecium'' of the phylum Ciliophora. They are covered in cilia which help in movement and feeding.''Paramecium'' can reproduce sexually, asexually, or by the process of endomixis. ''Paramecium aurelia'' demonstrate a strong "sex reaction" whereby groups of individuals will cluster together, and emerge in conjugant pairs. This pairing can last up to 12 hours, during which the micronucleus of each organism will be exchanged. In ''Paramecium aurelia'', a cryptic species complex was discovered by observation. Since then, some have tried to decode this complex using genetic data. Taxonomy ''Paramecium aurelia'' is a species complex composed of 15 known species (syngens), which are *'' Paramecium primaurelia'' *'' Paramecium biaurelia'' *'' Paramecium triaurelia'' *'' Paramecium tetraurelia'' *'' Paramecium pentaurelia'' *'' Paramecium sexaurelia'' *'' Paramecium septaurelia'' *'' Paramecium octau ...
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Protozoa
Protozoa (: protozoan or protozoon; alternative plural: protozoans) are a polyphyletic group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic debris. Historically, protozoans were regarded as "one-celled animals". When first introduced by Georg Goldfuss, in 1818, the taxon Protozoa was erected as a class within the Animalia, with the word 'protozoa' meaning "first animals", because they often possess animal-like behaviours, such as motility and predation, and lack a cell wall, as found in plants and many algae. This classification remained widespread in the 19th and early 20th century, and even became elevated to a variety of higher ranks, including phylum, subkingdom, kingdom, and then sometimes included within the paraphyletic Protoctista or Protista. By the 1970s, it became usual to require that all taxa be monophyletic (derived from a common ancestor that would also be regarded as protozo ...
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Allogromia Laticollaris
''Allogromia'' is a genus of Foraminifera Foraminifera ( ; Latin for "hole bearers"; informally called "forams") are unicellular organism, single-celled organisms, members of a phylum or class (biology), class of Rhizarian protists characterized by streaming granular Ectoplasm (cell bio .... Species: References Foraminifera genera Radiolarian genera {{foram-stub ...
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Foraminifera
Foraminifera ( ; Latin for "hole bearers"; informally called "forams") are unicellular organism, single-celled organisms, members of a phylum or class (biology), class of Rhizarian protists characterized by streaming granular Ectoplasm (cell biology), ectoplasm for catching food and other uses; and commonly an external shell (called a "Test (biology), test") of diverse forms and materials. Tests of chitin (found in some simple genera, and ''Textularia'' in particular) are believed to be the most primitive type. Most foraminifera are marine, the majority of which live on or within the seafloor sediment (i.e., are benthos, benthic, with different sized species playing a role within the macrobenthos, meiobenthos, and Benthos, microbenthos), while a smaller number float in the water column at various depths (i.e., are planktonic), which belong to the suborder Globigerinina. Fewer are known from freshwater or brackish conditions, and some very few (nonaquatic) soil species have been id ...
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Nuclear Dimorphism
Nuclear dimorphism is a term referred to the special characteristic of having two different kinds of nuclei in a cell. There are many differences between the types of nuclei. This feature is observed in protozoan ciliates, like ''Tetrahymena'', and some foraminifera. Ciliates contain two nucleus types: a macronucleus that is primarily used to control metabolism, and a micronucleus which performs reproductive functions and generates the macronucleus. The compositions of the nuclear pore complexes help determine the properties of the macronucleus and micronucleus. Nuclear dimorphism is subject to complex epigenetic controls. Nuclear dimorphism is continuously being studied to understand exactly how the mechanism works and how it is beneficial to cells. Learning about nuclear dimorphism is beneficial to understanding old eukaryotic mechanisms that have been preserved within these unicellular organisms but did not evolve into multicellular eukaryotes. Key components The ciliated ...
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Macronucleus
A macronucleus (formerly also meganucleus) is the larger type of nucleus in ciliates. Macronuclei are polyploid and undergo direct division without mitosis. It controls the non-reproductive cell functions, such as metabolism Metabolism (, from ''metabolē'', "change") is the set of life-sustaining chemical reactions in organisms. The three main functions of metabolism are: the conversion of the energy in food to energy available to run cellular processes; the co .... During conjugation, the macronucleus disintegrates, and a new one is formed by karyogamy of the micronuclei. Macronuclei contain hundreds to thousands of chromosomes, each present in many copies. There is no mechanism to precisely partition this complex genome equally during nuclear division; thus, how the cell manages to maintain a balanced genome after generations of divisions is unknown. See also * Micronucleus References . Cell nucleus Ciliate biology {{Ciliate-stub ...
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Cell Fusion
Cell fusion is an important cellular process in which several uninucleate cells (cells with a single nucleus) combine to form a multinucleate cell, known as a syncytium. Cell fusion occurs during differentiation of myoblasts, osteoclasts and trophoblasts, during embryogenesis, and morphogenesis. Cell fusion is a necessary event in the maturation of cells so that they maintain their specific functions throughout growth. History In 1839, Theodor Schwann, in his '' Microscopical Researches,'' expanded upon the theory that all living organisms are composed of cells when he added that discrete cells are the basis of life. Schwann observed that in certain cells the walls and cavities of the cells coalesce ('' verschmelzen'') together. This observation provided the first hint that cells fuse. It was not until 1960 that cell biologists deliberately fused cells for the first time. To fuse the cells, biologists combined isolated mouse cells and induced fusion of their outer membrane ...
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