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Caatinga (, ) is a type of semi-arid tropical vegetation, and an
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas o ...
characterized by this vegetation in interior northeastern Brazil. The name "Caatinga" is a Tupi word meaning "white forest" or "white vegetation" (''caa'' = forest, vegetation, ''tinga'' = white). The Caatinga is a xeric shrubland and thorn forest, which consists primarily of small, thorny trees that shed their leaves seasonally. Cacti, thick-stemmed plants, thorny brush, and arid-adapted grasses make up the ground layer. Most vegetation experiences a brief burst of activity during the three-month long rainy season. Caatinga falls entirely within earth's tropical zone and is one of 6 major ecoregions of Brazil. It covers 850,000 km², nearly 10% of Brazil's territory. It is home to 26 million people and over 2000 species of plants, fish, reptiles, amphibians, birds, and mammals. The Caatinga is the only exclusively Brazilian biome, which means that a large part of its biological heritage cannot be found anywhere else on the planet.


Geography

The Caatinga covers the interior portion of northeastern Brazil bordering the Atlantic seaboard (save for a fringe of Atlantic Forest), extending across nine states: Maranhão, Piauí, Ceará, Rio Grande do Norte, Paraíba,
Pernambuco Pernambuco () is a state of Brazil, located in the Northeast region of the country. With an estimated population of 9.6 million people as of 2020, making it seventh-most populous state of Brazil and with around 98,148 km², being the ...
,
Alagoas Alagoas (, ) is one of the 27 federative units of Brazil and is situated in the eastern part of the Northeast Region. It borders: Pernambuco (N and NW); Sergipe (S); Bahia (SW); and the Atlantic Ocean (E). Its capital is the city of Maceió. ...
, Sergipe,
Bahia Bahia ( , , ; meaning "bay") is one of the 26 states of Brazil, located in the Northeast Region of the country. It is the fourth-largest Brazilian state by population (after São Paulo, Minas Gerais, and Rio de Janeiro) and the 5th-largest ...
, and parts of
Minas Gerais Minas Gerais () is a state in Southeastern Brazil. It ranks as the second most populous, the third by gross domestic product (GDP), and the fourth largest by area in the country. The state's capital and largest city, Belo Horizonte (literall ...
. Altogether, the Caatinga comprises 850,000 km², about 10% of the surface area of Brazil. By comparison, it is over nine times the surface area of Portugal, whence came Brazil's early European settlers. Located between 3°S 45°W and 17°S 35°W, the Caatinga experiences irregular winds from all directions. Rainfall is thus intermittent but intense, totalling on average. Although the climate is typically hot and semi-arid, the Caatinga includes several enclaves of humid tropical forest, with trees tall. To the northwest, the Caatinga is bounded by the Maranhão Babaçu forests; to the west and southwest, the Atlantic dry forests and Cerrado savannas; to the east, the humid Atlantic coastal forests; and to the north and northeast, the Atlantic Ocean.


Climate

During the dry winter periods there is no foliage or undergrowth, as plants try to conserve water. Roots protrude through the surface of the stony soil, to absorb water before it is evaporated. Leaves fall off the trees to reduce transpiration. With all the foliage and undergrowth dead during the drought periods and all the trees having no leaves the Caatinga has a yellow-grey, desert-like look. During the peak periods of drought the Caatinga's soil can reach temperatures of up to 60 °C. The drought usually ends in December or January, when the rainy season starts. Immediately after the first rains, the grey, desert-like landscape starts to transform and becomes completely green within a few days. Small plants start growing in the now moist soil and trees grow back their leaves. Rivers that are mostly dry during the past 6 or 7 months start to fill up and streams begin to flow again.


Ecology

Caatinga harbors a unique biota, with thousands of endemic species. Caatinga contains over 1,000 vascular plant species in addition to 187 bees, 240 fish species, 167 reptiles and amphibians, 516 birds, and 148 mammal species, with endemism levels varying from 9 percent in birds to 57 percent in fishes.


Vegetation

The Caatinga does not correspond to a single type of vegetation, but rather a broad mosaic. Nonetheless, all vegetative structure is adapted to the xeric climate. Succulent and crassulaceous species dominate; non-succulents exhibit small, firm leaves and intense branching at the base, akin to shrubs. Palm stands usually contain carnaúba or babaçu palms, but occasionally tucumã and macaúba. The Caatinga has enough endemic species to constitute a floristic province. Most authors divide the Caatinga into two different subtypes: dry ("sertão") and humid ("agreste"), but categorizations vary to as many as eight different vegetative regimes.


Fauna

The Caatinga is home to nearly 50
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found els ...
species of birds, including Lear's macaw ''(Anodorhynchus leari)'', Spix's macaw ''(Cyanopsitta spixii)'',
moustached woodcreeper The moustached woodcreeper (''Xiphocolaptes falcirostris'') is a Vulnerable species of bird in the subfamily Dendrocolaptinae of the ovenbird family Furnariidae. It is endemic to Brazil. Taxonomy and systematics The moustached woodcree ...
''(Xiphocolaptes falcirostris)'', Caatinga parakeet,
Caatinga antwren The Caatinga antwren (''Radinopsyche sellowi'') is a species of bird in the family Thamnophilidae. It is the only member of the monotypic genus ''Radinopsyche''. Prior to 2022, it was classified in the genus ''Herpsilochmus'', but was reclassi ...
, Sao Francisco black tyrant and Caatinga cacholote. Endemic mammal species include: *eleven rodents -
Caatinga vesper mouse The caatinga vesper mouse (''Calomys expulsus'') (also known as the caatinga laucha or rejected vesper mouse) is a rodent species in the family Cricetidae from South America. It is endemic to eastern Brazil, where it is found in open savanna (cer ...
, ''
Wiedomys pyrrhorhinos The red-nosed mouse (''Wiedomys pyrrhorhinos'') is an arboreal rodent species endemic to Brazil. It is found in caatinga and cerrado habitat in southeast Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portug ...
'', ''
Trinomys yonenagae Yonenaga's Atlantic spiny-rat (''Trinomys yonenagae'') or torch-tail spiny rat is a spiny rat species endemic to Brazil. Locally, it is known as ''rabo de facho''. Named for Yatiyo Yonenaga-Yassuda, a cytogenetics researcher, it is considered a ...
'', ''
Trinomys albispinus The white-spined Atlantic spiny rat (''Trinomys albispinus'') is a spiny rat species endemic to Brazil. Description This is one of the smaller spiny rats, with a head-body length of , and a tail long. Adult weight can be anything from . The fu ...
'' minor, ''Trinomys albispinus sertonius'', '' Thylamys karimii'', '' Dasyprocta sp. n.'', '' Oryzomys sp. n.'', '' Oxymycterus sp. n.'', '' Rhipidomys sp. n. ssp. 1'', and '' Rhipidomys sp. n. ssp. 2'' *one primate - '' Callicebus barbarabrownae'' *two bats - ''
Xeronycteris vieirai Vieira's long-tongued bat (''Xeronycteris vieirai'') is a species of bat from northeastern Brazil, discovered in 2005 by Gregorin and Ditchfield. It is the only species in the genus ''Xeronycteris''. References Phyllostomidae Bats of Braz ...
'' and '' Chiroderma sp. n''


Possible anthropogenic origins

Based on radiocarbon dating of potsherds, proponents of historical ecology such as William Denevan and William Balee have suggested that large sections of the Caatinga region may be of anthropogenic origin. Over 1000 years ago, native peoples may have unintentionally created the environment of the modern-day Caatinga through constant slash-and-burn agriculture, thereby stymying plant succession and preventing major rainforests from growing within the region. Conversely, fossil evidence suggests that the Caatinga may historically have been part of a much larger dry belt.


Conservation

The Caatinga is poorly represented in the Brazilian Conservation Area network, with only 1% in Integral Protection Conservation Areas and 6% in Sustainable Use Conservation Areas. Protected areas include Chapada Diamantina National Park, Serra da Capivara National Park, and
Serra das Confusões National Park The Serra das Confusões National Park ( pt, Parque Nacional da Serra das Confusões) is a national park in the state of Piauí, Brazil. Geography The Serra das Confusões National Park is in the municipalities of Alvorada do Gurguéia, Brejo ...
. Economic developed has fragmented the native biome. Estimates on the amount of Caatinga transformed affected by economic development range 25-50%, making Caatinga the most degraded ecosystem in Brazil, following the Atlantic Forest, which has lost over 80% of its original cover.


Economic exploitation

The local population lives in extreme poverty, and many rely on extraction of natural resources for a livelihood. There are few drinkable water sources, and harvesting is difficult because of the irregular rainfall.


Agriculture

Native plants are used in local agriculture, much of it slash-and-burn. ''
Pilocarpus jaborandi ''Pilocarpus jaborandi'' is a species of flowering plant in the family Rutaceae, native to northeast Brazil. It is a source of the drug pilocarpine Pilocarpine is a medication used to reduce pressure inside the eye and treat dry mouth. As ey ...
'' appears to exhibit medicinal properties. The fruits of
umbú ''Spondias tuberosa'', commonly known as imbu, , Brazil plum, or umbu, is a plant native to northeast Brazil, where it grows in the Caatinga, the chaparral scrub that grows wild across dry lands. The round fruit is light yellow to red in colou ...
and mangabá are used as food directly, and other species are used for forage. Local palms produce commercial-grade lauric and
oleic Oleic acid is a fatty acid that occurs naturally in various animal and vegetable fats and oils. It is an odorless, colorless oil, although commercial samples may be yellowish. In chemical terms, oleic acid is classified as a monounsaturated omega ...
oils, which undergirds much of the economy of northeast Brazil. Meliponiculture is also a well-developed and traditional activity in the region. One of the most productive species, '' Melipona subnitida'', known locally as jandaíra, produces up to 6 liters of honey a year, resulting in economic profit for the population. Irrigation along the São Francisco River promises to turn the region into a breadbasket. The soil is very fertile, and existing irrigation infrastructure already supports the export of
grape A grape is a fruit, botanically a berry (botany), berry, of the deciduous wood