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Polypteriformes
Bichirs and the reedfish comprise Polypteridae , a family of archaic ray-finned fishes and the only family in the order Polypteriformes .Helfman GS, Collette BB, Facey DE, Bowen BW. 2009. The Diversity of Fishes. West Sussex, UK: Blackwell Publishing. 720 p. All the species occur in freshwater habitats in tropical Africa and the Nile River system, mainly swampy, shallow floodplains and estuaries. Cladistia, polypterids and their fossil relatives, are considered the sister group to all other extant ray-finned fishes ( Actinopteri).Dai Suzuki, Matthew C. Brandley, Masayoshi Tokita: ''CORRECTION: The mitochondrial phylogeny of an ancient lineage of ray-finned fishes (Polypteridae) with implications for the evolution of body elongation, pelvic fin loss, and craniofacial morphology in Osteichthyes.'' BMC Evolutionary Biology. Bd. 10, Art.-Nr. 209, 2010, They likely diverged from Actinopteri at least 330 million years ago. A closely related group, the Scanilepiformes, a ...
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Actinopterygii
Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class (biology), class of Osteichthyes, bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fish fin, fins made of webbings of skin supported by radially extended thin bony spine (zoology), spines called ''lepidotrichia'', as opposed to the bulkier, fleshy lobed fins of the sister taxon, sister clade Sarcopterygii (lobe-finned fish). Resembling folding fans, the actinopterygian fins can easily change shape and wetted area, providing superior thrust-to-weight ratios per movement compared to sarcopterygian and chondrichthyian fins. The fin rays attach directly to the proximal or basal skeletal elements, the radials, which represent the articulation (anatomy), articulation between these fins and the internal skeleton (e.g., pelvic and pectoral girdles). The vast majority of actinopterygians are teleosts. By species count, they domi ...
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Cladistia
Cladistia is a subclass of bony fishes whose only living members are the bichirs of tropical Africa. Their major synapomorphies are a heterocercal tail in which the dorsal fin has independent rays, and a posteriorly elongated parasphenoid. Cladistia are the earliest diverging branch of living Actinopterygii, and are thought to have diverged from the Actinopteri, the group which includes all other living ray finned fish, by the Carboniferous. However, the fossil range for the only extant order ( Polypteriformes) is comparatively young, only reaching as far back as the mid-Cretaceous of South America and Africa, and the two extant genera of bichir only diverged around the Miocene. Aside from bichirs, other extinct fish groups thought to be members of the group include the Scanilepiformes, known from Triassic (and possibly Permian) of the Northern Hemisphere. The Guildayichthyiformes of Carboniferous North America are also sometimes considered cladistians, but this is thought ...
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Scanilepiformes
Cladistia is a subclass of bony fishes whose only living members are the bichirs of tropical Africa. Their major synapomorphies are a heterocercal tail in which the dorsal fin has independent rays, and a posteriorly elongated parasphenoid. Cladistia are the earliest diverging branch of living Actinopterygii, and are thought to have diverged from the Actinopteri, the group which includes all other living ray finned fish, by the Carboniferous. However, the fossil range for the only extant order (Polypteriformes) is comparatively young, only reaching as far back as the mid-Cretaceous of South America and Africa, and the two extant genera of bichir only diverged around the Miocene. Aside from bichirs, other extinct fish groups thought to be members of the group include the Scanilepiformes, known from Triassic (and possibly Permian) of the Northern Hemisphere. The Guildayichthyiformes of Carboniferous North America are also sometimes considered cladistians, but this is thought to ...
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Polypterus Bichir
''Polypterus'' is a genus of freshwater fish in the bichir family (biology), family (Polypteridae) of order (biology), order Polypteriformes. The type species is the Polypterus bichir, Nile bichir (''P. bichir''). Fish in this genus live in various areas in Africa. ''Polypterus'' is the only known vertebrate to have lungs, but no trachea. The etymology of the genus name derives from a combination of the Greek language, Greek Prefix (linguistics), prefix πολυ-, ''poly-'' (many) and the root word πτερον, ''pteron'' (wing or fin) – "many fins". Taxonomy Despite the ancient origins of the Polypteriformes, the earliest fossils that can be confidently assigned as being of the ''Polypterus'' lineage are from the Middle Eocene (Lutetian) of Libya. In addition, studies exclusively using phylogenetic inferences have found that ''Polypterus'' may have only diverged from ''Erpetoichthys'' during the Neogene. Species The following species are known: * ''Polypterus ansorg ...
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Polypterus
''Polypterus'' is a genus of freshwater fish in the bichir family ( Polypteridae) of order Polypteriformes. The type species is the Nile bichir (''P. bichir''). Fish in this genus live in various areas in Africa. ''Polypterus'' is the only known vertebrate to have lungs, but no trachea. The etymology of the genus name derives from a combination of the Greek prefix πολυ-, ''poly-'' (many) and the root word πτερον, ''pteron'' (wing or fin) – "many fins". Taxonomy Despite the ancient origins of the Polypteriformes, the earliest fossils that can be confidently assigned as being of the ''Polypterus'' lineage are from the Middle Eocene (Lutetian) of Libya. In addition, studies exclusively using phylogenetic inferences have found that ''Polypterus'' may have only diverged from '' Erpetoichthys'' during the Neogene. Species The following species are known: * '' Polypterus ansorgii'' Boulenger, 1910 (Guinean bichir) * ''Polypterus bichir'' Lacépède, 1803 (Nile ...
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Teleost
Teleostei (; Ancient Greek, Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts (), is, by far, the largest group of ray-finned fishes (class Actinopterygii), with 96% of all neontology, extant species of fish. The Teleostei, which is variously considered a Division (zoology), division or an infraclass in different taxonomic systems, include over 26,000 species that are arranged in about 40 order (biology), orders and 448 family (biology), families. Teleosts range from giant oarfish measuring or more, and ocean sunfish weighing over , to the minute male anglerfish ''Photocorynus spiniceps'', just long. Including not only torpedo-shaped fish built for speed, teleosts can be flattened vertically or horizontally, be elongated cylinders or take specialised shapes as in anglerfish and seahorses. The difference between teleosts and other bony fish lies mainly in their jaw bones; teleosts have a movable premaxilla and corresponding modifications ...
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Sarcopterygii
Sarcopterygii (; )—sometimes considered synonymous with Crossopterygii ()—is a clade (traditionally a class (biology), class or subclass) of vertebrate animals which includes a group of bony fish commonly referred to as lobe-finned fish. These vertebrates are characterised by prominent muscular limb buds (lobes) within their fish fin, fins, which are supported by articulated appendicular skeletons. This is in contrast to the other clade of bony fish, the Actinopterygii, which have only skin-covered lepidotrichia, bony spines supporting the fins. The tetrapods, a mostly terrestrial animal, terrestrial clade of vertebrates, are now recognized as having evolved from sarcopterygian ancestors and are most closely related to lungfishes. Their paired pectoral fins, pectoral and pelvic fins evolved into limb (anatomy), limbs, and their lung bud, foregut diverticulum eventually evolved into air-breathing lungs. Cladistics, Cladistically, this would make the tetrapods a subgroup within ...
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Tetrapod
A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetrapoda (). Tetrapods include all Neontology#Extant taxa versus extinct taxa, extant and Extinction, extinct amphibians and amniotes, with the latter in turn Evolution, evolving into two major clades, the Sauropsida, sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (extinct pelycosaur, "pelycosaurs", therapsids and all extant mammals, including Homo sapiens, humans). Hox gene mutations have resulted in some tetrapods becoming Limbless vertebrate, limbless (snakes, legless lizards, and caecilians) or two-limbed (cetaceans, sirenians, Bipedidae, some lizards, kiwi (bird), kiwis, and the extinct moa and elephant birds). Nevertheless, they still qualify as tetrapods through their ancestry, and some retain a pair of ves ...
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Ganoid
A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as well as possible hydrodynamic advantages. The term ''scale'' derives from the Old French , meaning a shell pod or husk. Scales vary enormously in size, shape, structure, and extent, ranging from strong and rigid armour plates in fishes such as shrimpfishes and boxfishes, to microscopic or absent in fishes such as eels and anglerfishes. The morphology (biology), morphology of a scale can be used to identify the species of fish it came from. Scales originated within the jawless ostracoderms, ancestors to all jawed fishes today. Most bony fishes are covered with the cycloid scales of salmon and carp, or the ctenoid scales of perch, or the ganoid scales of sturgeons and gars. Cartilaginous fishes (sharks and rays) are covered with placoid sca ...
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Dorsal Fin
A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found in most fish, in mammals such as whales, and in extinct ancient marine reptiles such as ichthyosaurs. Most have only one dorsal fin, but some have two or three. Wildlife biologists often use the distinctive nicks and wear patterns which develop on the dorsal fins of whales to identify individuals in the field. The bones or cartilages that support the dorsal fin in fish are called pterygiophores. Functions The main purpose of the dorsal fin is usually to stabilize the animal against rolling and to assist in sudden turns. Some species have further adapted their dorsal fins to other uses. The sunfish uses the dorsal fin (and the anal fin Fins are moving appendages protruding from the body of fish that interact with water to ge ...
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Fish
A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can be grouped into the more basal (phylogenetics), basal jawless fish and the more common jawed fish, the latter including all extant taxon, living cartilaginous fish, cartilaginous and bony fish, as well as the extinct placoderms and acanthodians. In a break to the long tradition of grouping all fish into a single Class (biology), class (Pisces), modern phylogenetics views fish as a paraphyletic group. Most fish are ectotherm, cold-blooded, their body temperature varying with the surrounding water, though some large nekton, active swimmers like white shark and tuna can hold a higher core temperature. Many fish can communication in aquatic animals#Acoustic, communicate acoustically with each other, such as during courtship displays. The stud ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Aus ...
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