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Cynodonts
Cynodontia () is a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extinct ancestors and close relatives ( Mammaliaformes), having evolved from advanced probainognathian cynodonts during the Late Triassic. Non-mammalian cynodonts occupied a variety of ecological niches, both as carnivores and as herbivores. Following the emergence of mammals, most other cynodont lines went extinct, with the last known non-mammaliaform cynodont group, the Tritylodontidae, having its youngest records in the Early Cretaceous. Description Early cynodonts have many of the skeletal characteristics of mammals. The teeth were fully differentiated and the braincase bulged at the back of the head. Outside of some crown-group mammals (notably the therians), all cynodonts probably laid eggs. The temporal fenestrae were much larger than th ...
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Brasilitherium Riograndensis
''Brasilodon'' ("tooth from Brazil") is an extinct genus of small, mammal-like cynodonts that lived in what is now Brazil during the Norian age of the Late Triassic epoch, about 225.42 million years ago. While no complete skeletons have been found, the length of ''Brasilodon'' has been estimated at . Its dentition shows that it was most likely an insectivore. The genus is monotypic, containing only the species ''B. quadrangularis''. ''Brasilodon'' belongs to the family Brasilodontidae, whose members were some of the closest relatives of mammals, the only cynodonts alive today. Two other brasilodontid genera, ''Brasilitherium'' and ''Minicynodon'', are now considered to be junior synonyms of ''Brasilodon''. Discovery and naming The first three specimens referred to ''Brasilodon quadrangularis'' were found at the Linha São Luiz site, a quarry near the town of Faxinal do Soturno in the state of Rio Grande do Sul. The rocks where ''Brasilodon'' was found belong to the upper part ...
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Permian–Triassic Extinction Event
The Permian–Triassic extinction event (also known as the P–T extinction event, the Late Permian extinction event, the Latest Permian extinction event, the End-Permian extinction event, and colloquially as the Great Dying,) was an extinction event that occurred approximately 251.9 Million years ago, million years ago (mya), at the boundary between the Permian and Triassic geologic periods, and with them the Paleozoic and Mesozoic eras. It is Earth's most severe known extinction event, with the extinction of 57% of Family (biology), biological families, 62% of genera and 81% of marine biology, marine species, (previous estimates of 90–96% marine species extinction were due to historical conflation with the End-Capitanian mass extinction event, end-Capitanian mass extinction which occurred 7–10 million years earlier) and 70% of terrestrial vertebrate species. It is also the greatest known mass extinction of insects. It is the greatest of the "Big Five" mass extinctions of t ...
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Eutheriodont
Eutheriodontia is a clade of therapsids which appear during the Middle Permian and which includes therocephalians and cynodonts, this latter group including mammals and related forms. With the dicynodonts, they form one of two lineages of therapsids that survived the End-Permian extinction and which diversified again during the Triassic, before the majority of them disappeared before or during the Triassic–Jurassic extinction, except for a lineage of cynodonts that later gave rise to mammals. Classification The clade was named in 1986 by James Allen Hopson and Herbert Richard Barghusen, the name meaning the "true Theriodontia". Within Hopson's system, the Eutheriodontia are the sister group of the Gorgonopsia within the Theriodontia. A close relationship between therocephalians and cynodonts had been recognized for many years. In 2001 the Eutheriodontia were defined as the least inclusive clade including Mammalia and '' Bauria''.James A. Hopson and James W. Kitching, 2 ...
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Mammal
A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three Evolution of mammalian auditory ossicles, middle ear bones. These characteristics distinguish them from reptiles and birds, from which their ancestors Genetic divergence, diverged in the Carboniferous Period over 300 million years ago. Around 6,640 Neontology#Extant taxon, extant species of mammals have been described and divided into 27 Order (biology), orders. The study of mammals is called mammalogy. The largest orders of mammals, by number of species, are the rodents, bats, and eulipotyphlans (including hedgehogs, Mole (animal), moles and shrews). The next three are the primates (including humans, monkeys and lemurs), the Artiodactyl, even-toed ungulates (including pigs, camels, and whales), and the Carnivora (including Felidae, ...
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Tritylodontidae
Tritylodontidae ("three-knob teeth", named after the shape of their cheek teeth) is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, with several mammalian traits including erect limbs, endothermy, and some details of the skeleton. They were the last-known family of the non-mammaliaform synapsids, persisting into the Early Cretaceous. Most tritylodontids are thought to have been herbivorous, feeding on vegetation such as stems, leaves, and roots, although at least one may have had a more omnivorous diet. Tritylodontid fossils are found in the Americas, South Africa, and Eurasia – they appear to have had an almost global distribution, including Antarctica. Description The skull of tritylodontids had a high sagittal crest. They retained the primitive condition of the joint between the quadrate bone of the skull and the articular bone of the lower jaw – the retention of the joint is one of the reasons they are technically regarded to not ...
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Epicynodontia
Epicynodontia is a clade of cynodont therapsids that includes most cynodonts, such as galesaurids, thrinaxodontids, and Eucynodontia (including mammals). It was erected as a stem-based taxon by Hopson and Kitching (2001) and defined as the most inclusive clade containing Mammalia and excluding ''Procynosuchus'', a Late Permian genus that is one of the most basal cynodonts. Below is a cladogram from Pusch ''et al.'' (2023) showing one hypothesis of cynodont relationships: References

Epicynodontia, Cynodontia Lopingian first appearances Extant Permian first appearances Tetrapod unranked clades Taxa named by James Hopson Taxa named by James Kitching {{Cynodont-stub ...
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Mammal
A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three Evolution of mammalian auditory ossicles, middle ear bones. These characteristics distinguish them from reptiles and birds, from which their ancestors Genetic divergence, diverged in the Carboniferous Period over 300 million years ago. Around 6,640 Neontology#Extant taxon, extant species of mammals have been described and divided into 27 Order (biology), orders. The study of mammals is called mammalogy. The largest orders of mammals, by number of species, are the rodents, bats, and eulipotyphlans (including hedgehogs, Mole (animal), moles and shrews). The next three are the primates (including humans, monkeys and lemurs), the Artiodactyl, even-toed ungulates (including pigs, camels, and whales), and the Carnivora (including Felidae, ...
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Megazostrodon Rudnerae
''Megazostrodon'' is an extinct genus of basal mammaliaforms belonging to the order Morganucodonta. It is approximately 200 million years old.Fur and Fangs: Mammal Origins
. Palaeobiology and Biodiversity Research Group, University of Bristol.
Two species are known: ''M. rudnerae'' from the of Lesotho and South Africa, and ''M. chenali'' from the of France.


Discovery

The

Probainognathia
Probainognathia is one of the two major subgroups of the clade Eucynodontia, the other being Cynognathia. The earliest forms were carnivorous and insectivorous, though some groups eventually also evolved herbivorous diets. The earliest and most basal probainognathian is the Middle Triassic (Anisian) aged ''Lumkuia'', from South Africa, though probainognathians would not become prominent until the mid Norian stage of the Late Triassic. Three groups survived the Triassic–Jurassic extinction event, extinction at the end of Triassic: Tritheledontidae and Tritylodontidae, which both survived until the Jurassic—the latter even into the Cretaceous (''Montirictus'' and ''Xenocretosuchus'')—and Mammaliaformes, which includes the mammals. Phylogeny Below is a cladogram from Ruta, Botha-Brink, Mitchell and Benton (2013) showing one hypothesis of cynodont relationships: Cladogram from Stefanello ''et al.'' (2023): See also * Evolution of mammals * List of prehistoric mammals Refe ...
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Dvinia
''Dvinia'' is an extinct genus of cynodonts found in the Salarevo Formation of Sokolki on the Northern Dvina River near Kotlas in Arkhangelsk Oblast, Russia. It is the only known member of the family Dviniidae. Its fossil remains date from the Late Permian and were found with ''Inostrancevia'', ''Scutosaurus'' and '' Vivaxosaurus''. ''Dvinia'' was a small omnivore possessing a large temporal opening typical of advanced therapsids, with a thin postorbital bar separating the eye from the muscle attachment. As a cynodont, it was closely related to mammals. The dentition consisted of a set of small incisors followed by 2 canines and 10-14 postcanines. See also * List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all Genus, genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also gene ... References Further reading * Patricia ...
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Madysaurus
''Madysaurus'' (meaning " Madygen reptile") is an extinct genus of cynodonts which existed in Kyrgyzstan. It was first named by Leonid Petrovich Tatarinov in 2005. ''Madysaurus'' is known from the Madygen Formation, a Triassic Lagerstätte that also includes well-preserved remains of insects and small reptiles like '' Sharovipteryx'' and ''Longisquama ''Longisquama'' is a genus of extinct reptile. There is only one species, ''Longisquama insignis'', known from a poorly preserved skeleton and several incomplete fossil impressions from the Middle Triassic, Middle to Late Triassic Madygen Formati ...''. ''Madysaurus'' is one of the most primitive cynodonts and is placed in its own family, Madysauridae. References Cynodontia Middle Triassic synapsids of Asia Late Triassic synapsids of Asia Carnian genera Fossils of Kyrgyzstan Madygen Formation Fossil taxa described in 2005 Taxa named by Leonid Petrovich Tatarinov {{Paleo-cynodont-stub ...
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Dvinia Prima
''Dvinia'' is an extinct genus of cynodonts found in the Salarevo Formation of Sokolki on the Northern Dvina River near Kotlas in Arkhangelsk Oblast, Russia. It is the only known member of the family Dviniidae. Its fossil remains date from the Late Permian and were found with ''Inostrancevia'', ''Scutosaurus'' and '' Vivaxosaurus''. ''Dvinia'' was a small omnivore possessing a large temporal opening typical of advanced therapsids, with a thin postorbital bar separating the eye from the muscle attachment. As a cynodont, it was closely related to mammals. The dentition consisted of a set of small incisors followed by 2 canines and 10-14 postcanines. See also * List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all Genus, genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also gene ... References Further reading * Patricia ...
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