Yandusaurus
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''Yandusaurus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of herbivorous basal
neornithischian Neornithischia ("new ornithischians") is a clade of the dinosaur order Ornithischia. It is the sister group of the Thyreophora within the clade Genasauria. Neornithischians are united by having a thicker layer of asymmetrical enamel on the insi ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the
Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Strat ...
age (middle
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
, approximately 168 to 162 Ma) of
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
.


Discovery and naming

In January 1973 the Museum of the Salt Industry at
Zigong Zigong (, ), formed by the merger of the two former towns of Ziliujing (Tzuliuching, literally "self-flow well") and Gongjing (Kungching, literally "offering well"), is a prefecture-level city in Sichuan, southwestern China. Demographics Accord ...
in
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
was warned that during construction activities with a composter at Jinzidang near the Honghe dam inadvertently a dinosaur skeleton had been processed. A team of the museum managed to salvage some heavily damaged remains. Though locally this animal was at first referred to as "Yubasaurus" or "Honghesaurus", in 1979
Beijing } Beijing ( ; ; ), alternatively romanized as Peking ( ), is the capital of the People's Republic of China. It is the center of power and development of the country. Beijing is the world's most populous national capital city, with over 21 ...
professor He Xinlu named and described it as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
''Yandusaurus hongheensis''. The generic name is derived from ''Yandu'', the ancient name for Zigong. This name is a contraction of ''yan'', "salt", and ''du'', "capital", occasioned by the fact that Zigong was historically the centre of Chinese salt mining. In this way ''Yandusaurus'' indirectly also refers to the Salt Museum. The specific name refers to the Honghe river.He X., 1979, "A newly discovered ornithopod dinosaur ''Yandusaurus'' from Zigong, Sichuan". In: ''Contributions to International Exchange of Geology. Part 2. Stratigraphy and paleontology''. Geol. publishing House, Beijing: pp. 116-123 The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen, GCC V20501, had been dug up in a layer of the
Lower Shaximiao Formation The Shaximiao Formation () is a Middle Jurassic, Middle to Late Jurassic aged geological formation in Sichuan, China, most notable for the wealth of dinosaurs fossils that have been excavated from its strata. The Shaximiao Formation is exposed in ...
. It consists of a partial skeleton with skull. Elements of most parts of the body have been preserved, the skull, vertebral column, shoulder girdle, frontlimbs and hindlimbs but all are very limited and/or damaged. The lower jaws, pelvis and the end of the tail had probably been entirely destroyed by the machine.He, X. & Cai, K., 1984, ''The Middle Jurassic dinosaurian fauna from Dashampu, Zigong, Sichuan. Vol. 1: The ornitopod dinosaurs''. Sichuan Scientific and Technological Publishing House, Chengdu, Sichuan, pp. 1-71 In 1983 He and Cai Kaiji identified a second species: ''Yandusaurus multidens'', known from two nearly complete skeletons and nine partial skeletons. In 1992 this was by Guangzhao Peng reassigned to the genus ''
Agilisaurus ''Agilisaurus'' (; 'agile lizard') is a genus of ornithischian dinosaur from the Middle Jurassic Period of what is now eastern Asia. It was about 3.5–4 ft (1.2-1.7 m) long, 2 ft (0.6 m) in height and 40 kg in weight. It has ...
'' as a ''A. multidens''. In 1996
Gregory S. Paul Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dino ...
renamed it as ''
Othnielia ''Nanosaurus'' ("small or dwarf lizard") is the name given to a genus of neornithischian dinosaur that lived about 155 to 148 million years ago, during the Late Jurassic-age. Its fossils are known from the Morrison Formation of the south-wes ...
multidens''. In 2005 it was by
Paul Barrett Paul Franklyn "Legs" Barrett (14 December 1940 – 20 January 2019) was a UK agent and manager of 1950s style Rock and Roll artistes, an author and previously a singer, songwriter and film actor. Barrett is the discoverer, mentor and first ma ...
e.a. placed in its own genus, ''
Hexinlusaurus ''Hexinlusaurus'' is a genus of basal ornithischian dinosaur from the Middle Jurassic of China. The holotype (ZDM T6001, Zigong Dinosaur Museum, Dashanpu, People's Republic of China), consists of an almost complete, articulated skull and some ...
''.


Description

A fast-moving biped, ''Yandusaurus'' had four toes on each foot and five fingers on each hand. It had large eyes as shown by the curved
jugal The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anatomy ...
. The teeth, showing a unique pattern of parallel vertical ridges compared by Chinese researchers to the fingers of the hand of Buddha statues, are very asymmetrical in that the inner side is strongly worn down. The fifteen
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
ry teeth are largest in the middle and overlap. When indicating the size of ''Yandusaurus'' many sources give that of ''Hexinlusaurus'', the previous ''Y. multidens'', the (all juvenile) specimens of which are sixty centimetres (two feet) to 1.6 metres (5 ft) in length, and weighed around seven
kilogram The kilogram (also kilogramme) is the unit of mass in the International System of Units (SI), having the unit symbol kg. It is a widely used measure in science, engineering and commerce worldwide, and is often simply called a kilo colloquially ...
s (fifteen lb). ''Yandusaurus hongheensis'' was however considerably larger than these exemplars: He estimated the body length at three metres, Peng at 3.2 metres; in 2010 G.S. Paul gave an estimation of 3.8 metres length and a weight of hundred forty kilogrammes.Paul, G.S., 2010, ''The Princeton Field Guide to Dinosaurs'', Princeton University Press p. 274


Phylogeny

''Yandusaurus'' was assigned by He to the Hypsilophodontidae. Today however Hypsilophodontidae is considered an unnatural (
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
) group and ''Yandusaurus'' is viewed as taking an unresolved position within
Neornithischia Neornithischia ("new ornithischians") is a clade of the dinosaur order Ornithischia. It is the sister group of the Thyreophora within the clade Genasauria. Neornithischians are united by having a thicker layer of asymmetrical enamel on the inside ...
.


Notes

{{Portal bar, Dinosaurs, China Fossil taxa described in 1979 Middle Jurassic dinosaurs of Asia Paleontology in Sichuan Ornithischian genera