Neornithischian
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Neornithischian
Neornithischia ("new ornithischians") is a clade of the dinosaur order Ornithischia. It is the sister group of the Thyreophora within the clade Genasauria. Neornithischians are united by having a thicker layer of asymmetrical enamel on the inside of their lower teeth. The teeth wore unevenly with chewing and developed sharp ridges that allowed neornithischians to break down tougher plant food than other dinosaurs. Neornithischians include a variety of basal forms historically known as "hypsilophodonts", including the Parksosauridae; in addition, there are derived forms classified in the groups Marginocephalia and Ornithopoda. The former includes clades Pachycephalosauria and Ceratopsia, while the latter typically includes ''Hypsilophodon'' and the more derived Iguanodontia. Classification Neornithischia was first named by Cooper in 1985 and defined as "all genasaurians more closely related to '' Parasaurolophus walkeri'' than to ''Ankylosaurus magniventris'' or ''Stegosaurus st ...
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Kulindadromeus
''Kulindadromeus'' was a herbivorous dinosaur, a basal neornithischian from the Middle Jurassic. The first ''Kulindadromeus'' fossil was found in Russia. Its feather-like integument is evidence for protofeathers being basal to Ornithischia and possibly Dinosauria as a whole, rather than just to Coelurosauria, as previously suspected. Discovery and naming In 2014, the type species ''Kulindadromeus zabaikalicus'' was named and described by Pascal Godefroit, Sofia Sinitsa, Danielle Dhouailly, Yuri Bolotsky, Alexander Sizov, Maria McNamara, Michael Benton, and Paul Spagna. The generic name is derived from the Kulinda sites and Classical Greek δρομεύς, ''dromeus'', "runner". The specific name refers to the Zabaykalsky Krai. The holotype, INREC K3/109, was found in a layer of the Ukureyskaya Formation dating from the Middle to Late Jurassic, Bajocian-Tithonian, with the specific bed likely dating to the Bathonian stage of the Middle Jurassic, between 168 and 166 million ye ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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Cerapoda
Cerapoda ("ceratopsians" and "ornithopods") is a clade of the dinosaur order Ornithischia, that includes pachycephalosaurs, ceratopsians and ornithopods Classification Cerapoda is divided into two groups: Ornithopoda ("bird-foot") and Marginocephalia ("fringed heads"). The latter group includes the Pachycephalosauria ("thick-headed lizards") and Ceratopsia ("horned faces"). The following taxonomy follows Richard J. Butler, Paul Upchurch and David B. Norman, 2008 (and Butler ''et al.'', 2011) unless otherwise noted. Cerapoda was first named by Sereno in 1986 and defined by him as "'' Parasaurolophus walkeri'' Parks, 1922, '' Triceratops horridus'' Marsh, 1889, their most recent common ancestor and all descendants". In 2021, Cerapoda was given a formal definition under the PhyloCode: "The smallest clade containing ''Iguanodon bernissartensis'' Boulenger in Beneden, 1881, ''Pachycephalosaurus wyomingensis'' (Gilmore, 1931), and '' Triceratops horridus'' Marsh, 1889."Madzia D ...
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Parasaurolophus Walkeri
''Parasaurolophus'' (; meaning "near crested lizard" in reference to ''Saurolophus)'' is a genus of herbivorous hadrosaurid ornithopod dinosaur that lived in what is now North America and possibly Asia during the Late Cretaceous Period, about 76.5–73 million years ago. It was a herbivore that walked both as a biped and as a quadruped. Three species are universally recognized: ''P. walkeri'' (the type species), ''P. tubicen'', and the short-crested ''P. cyrtocristatus''. Additionally, a fourth species, ''P. jiayinensis'', has been proposed, although it is more commonly placed in the separate genus ''Charonosaurus''. Remains are known from Alberta (Canada), New Mexico and Utah (United States), and possibly Heilongjiang (China). The genus was first described in 1922 by William Parks from a skull and partial skeleton found in Alberta. ''Parasaurolophus'' was a hadrosaurid, part of a diverse family of Cretaceous dinosaurs known for their range of bizarre head adornments whi ...
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Iguanodontia
Iguanodontia (the iguanodonts) is a clade of herbivorous dinosaurs that lived from the Middle Jurassic to Late Cretaceous. Some members include ''Camptosaurus'', ''Dryosaurus'', ''Iguanodon'', ''Tenontosaurus'', and the hadrosaurids or "duck-billed dinosaurs". Iguanodontians were one of the first groups of dinosaurs to be found. They are among the best known of the dinosaurs, and were among the most diverse and widespread herbivorous dinosaur groups of the Cretaceous period. Classification Iguanodontia is often listed as an infraorder within a suborder Ornithopoda, though Benton (2004) lists Ornithopoda as an infraorder and does not rank Iguanodontia. Traditionally, iguanodontians were grouped into the superfamily Iguanodontoidea and family Iguanodontidae. However, phylogenetic studies show that the traditional "iguanodontids" are a paraphyletic grade leading up to the hadrosaurs (duck-billed dinosaurs). Groups like Iguanodontoidea are sometimes still used as unranked clades ...
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Ceratopsia
Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, ''Yinlong downsi'', lived between 161.2 and 155.7 million years ago.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> The last ceratopsian species, ''Triceratops prorsus'', became extinct during the Cretaceous–Paleogene extinction event, . ''Triceratops'' is by far the best-known ceratopsian to the general public. It is traditional for ceratopsian genus names to end in "''-ceratops''", although this is not always the case. One of the first named genera was ''Ceratops'' itself, which lent its name to the group, although it is considered a '' nomen dubium'' today as its fossil remains have no distinguishing charact ...
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Pachycephalosauria
Pachycephalosauria (; from Greek παχυκεφαλόσαυρος for 'thick headed lizards') is a clade of ornithischian dinosaurs. Along with Ceratopsia, it makes up the clade Marginocephalia. With the exception of two species, most pachycephalosaurs lived during the Late Cretaceous Period, dating between about 85.8 and 65.5 million years ago. They are exclusive to the Northern Hemisphere, all of them being found in North America and Asia. They were all bipedal, herbivorous/omnivorous animals with thick skulls. Skulls can be domed, flat, or wedge-shaped depending on the species, and are all heavily ossified. The domes were often surrounded by nodes and/or spikes. Partial skeletons have been found of several pachycephalosaur species, but to date no complete skeletons have been discovered. Often isolated skull fragments are the only bones that are found. Candidates for the earliest known pachycephalosaur include ''Ferganocephale adenticulatum'' from Middle Jurassic Period strata ...
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Ornithopoda
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous world, dominating the North American land. Their major evolutionary advantage was the progressive development of a chewing apparatus that became the most sophisticated ever developed by a non-avian dinosaur, rivaling that of modern mammals such as the domestic cow. They reached their apex of diversity and ecological dominance in the hadrosaurids (colloquially known as 'duck-bills'), before they were wiped out by the Cretaceous–Paleogene extinction event along with all other non-avian dinosaurs. Members are known from all seven continents, though they are generally rare in the Southern Hemisphere. History of research In 1870, Thomas Henry Huxley listed Iguanodontidae (coined by Cope a year earlier) as one of his three families of dinosaurs ( ...
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Marginocephalia
Marginocephalia (/mär′jə-nō-sə-făl′ē-ən/ Latin: margin-head) is a clade of ornithischian dinosaurs that is characterized by a bony shelf or margin at the back of the skull. These fringes were likely used for display. There are two clades included in Marginocephalia: the thick-skulled Pachycephalosauria and the horned Ceratopsia. All members of Marginocephalia were primarily herbivores. They basally used gastroliths to aid in digestion of tough plant matter until they convergently evolved tooth batteries in Neoceratopsia (or "new Ceratopsia") and Pachycephalosauria. Marginocephalia first evolved in the Jurassic Period and became more common in the Cretaceous. They are basally small facultative quadrupeds while derived members of the group are large obligate quadrupeds. Primitive marginocephalians are found in Asia, but the group migrated upwards into North America. Pachycephalosaurs, or "thick-headed reptiles", have primitive features that include basally small sized bodi ...
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Parksosauridae
Thescelosauridae is a clade of neornithischians from the Cretaceous of Asia, North America and possibly South America. The group was originally used as a name by Charles M. Sternberg in 1937, but was not formally defined until 2013, where it was used by Brown and colleagues as the group uniting ''Thescelosaurus'' and ''Orodromeus'', based on their phylogenetic results. During a phylogenetic revision of neornithischians by Clint Boyd in 2015, the authorship of Thescelosauridae was given to Brown and colleagues, which meant that the similar name Parksosauridae, informally defined in 2002 by Buchholz, would have had priority over Thescelosauridae. The two clades had slightly different definitions, with Parksosauridae referring to all animals closer to ''Parksosaurus'' than ''Hypsilophodon'', but they contained the same taxa so Boyd used Parksosauridae under the assumption it had priority. However, in formalizing the clade following the regulations of the PhyloCode, Madzia, Boyd, and c ...
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Hypsilophodont
Hypsilophodontidae (or Hypsilophodontia) is a traditionally used family of ornithopod dinosaurs, generally considered invalid today. It historically included many small bodied bipedal neornithischian taxa from around the world, and spanning from the Middle Jurassic until the Late Cretaceous. This inclusive status was supported by some phylogenetic analyses from the 1990s and mid 2000s, although there have also been many finding that the family is an unnatural grouping which should only include the type genus, ''Hypsilophodon'', with the other genera being within clades like Thescelosauridae and Elasmaria. A 2014 analysis by Norman recovered a grouping of ''Hypsilophodon'', Rhabdodontidae and ''Tenontosaurus'', which he referred to as Hypsilophodontia. All other analyses from around the same time have instead found these latter taxa to be within Iguanodontia. Linnaean usage Hypsilophodontidae was named originally in 1882 by Louis Dollo, as a family to include ''Hypsilophodon'' a ...
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Genasauria
Genasauria is a clade of extinct beaked, primarily herbivorous dinosaurs. Paleontologist Paul Sereno first named Genasauria in 1986. The name Genasauria is derived from the Latin word ''gena'' meaning ‘cheek’ and the Greek word ''saúra'' (σαύρα) meaning ‘lizard.’ It is hypothesized that Genasauria had diverged from ''Lesothosaurus'' by the Early Jurassic. Cranial features that characterize Genasauria include a medial offset of the maxillary dentition, a sprout-shaped mandibular symphysis, moderately sized coronoid process, and an edentulous (without teeth) anterior portion of the premaxilla. A distinguishing postcranial feature of Genasauria is a pubic peduncle of the ilium that is less robust than the ischial peduncle. Genasauria is commonly divided into Neornithischia and Thyreophora. Neornithischia is characterized by asymmetrical distributions of enamel covering the crowns of the cheek teeth, an open acetabulum, and a laterally protruding ischial peduncle of the ...
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