''Kakuru'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

from the early

Cretaceous Period The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of t ...

. The only described species, ''Kakuru kujani'', is known primarily from evidence of a single

tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...

, which had been fossilised through a rare process in which the bone through

hydration Hydration may refer to: * Hydrate, a substance that contains water * Hydration enthalpy, energy released through hydrating a substance * Hydration reaction, a chemical addition reaction where a hydroxyl group and proton are added to a compound * ...

turned to

opal Opal is a hydrated amorphous form of silica (SiO2·''n''H2O); its water content may range from 3 to 21% by weight, but is usually between 6 and 10%. Due to its amorphous property, it is classified as a mineraloid, unlike crystalline forms ...

. The bone was dug up at the

fields of Andamooka,

South Australia South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...

. The opalised tibia was exhibited by a gem shop in 1973 and by chance brought to the attention of paleontologist Neville Pledge. The owner at the time, a certain A. Fleming, allowed pictures and two casts to be made but eventually the specimen was sold at an auction to an anonymous buyer. It was presumed lost to science. In 2004, however, the

South Australian Museum The South Australian Museum is a natural history museum and research institution in Adelaide, South Australia, founded in 1856 and owned by the Government of South Australia. It occupies a complex of buildings on North Terrace in the cultu ...

succeeded in procuring the fossil for $22,000. ''Kakuru'' was formally named in 1980 by Pledge and

Ralph Molnar Ralph E. Molnar is a paleontologist who had been Curator of Mammals at the Queensland Museum and more recently associated with the Museum of Northern Arizona. He is also a research associate at the Texas natural Science Centre. He co-authored descr ...

.R. E. Molnar and N. S. Pledge, 1980, "A new theropod dinosaur from South Australia", ''Alcheringa'' 4: 281-287 The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Kakuru kujani''. The generic name is that of a

Rainbow Serpent The Rainbow Serpent or Rainbow Snake is a common deity often seen as the creator God, known by numerous names in different Australian Aboriginal languages by the many different Aboriginal peoples. It is a common motif in the art and religion ...

Australian Aboriginal mythology Australian Aboriginal religion and mythology is the sacred spirituality represented in the stories performed by Aboriginal Australians within each of the Aboriginal Australian languages, language groups across Australia in their Aboriginal ce ...

. The specific name is that of the local aboriginal tribe, the Kujani or Guyani. Kakuru phalanx

One of the casts is the plastoholotype, SAM P17926. The specimen was discovered in the marine Bulldog Shale of the Marree Subgroup, dating to the

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...

. Apart from the tibia, the find included some small probable fibula fragments. Later a foot digit was referred that might have come from the same species, specimen SAM P18010, but the assignment is dubious. The tibia is broken into about ten larger pieces and roughly 33 centimetres long. It is very slender in build and shows the impression of the ascending process of the

astragalus ''Astragalus'' is a large genus of over 3,000 species of herbs and small shrubs, belonging to the legume family Fabaceae and the subfamily Faboideae. It is the largest genus of plants in terms of described species. The genus is native to tempe ...

, an ankle bone itself lost. The process seems to have been very long and narrow. ''Kakuru'' is believed to have been carnivorous, was bipedal and about two to three meters in length. This small dinosaur seems to have had long, slender legs. Due to the paucity of the remains it has been difficult to establish the

phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...

position of ''Kakuru''. Molnar and Pledge gave no more precise determination than a

Theropoda Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ca ...

incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...

''. The tibia provides two indications: its gracile form and the tall height and narrow width of the astragalar process. Both seems to point to the

Coelurosauria Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyrann ...

, perhaps the

Oviraptorosauria Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or wit ...

. However, in 2005

Oliver Rauhut Oliver may refer to: Arts, entertainment and literature Books * ''Oliver the Western Engine'', volume 24 in ''The Railway Series'' by Rev. W. Awdry * '' Oliver Twist'', a novel by Charles Dickens Fictional characters * Ariadne Oliver ...

pointed out that the

Abelisauroidea Abelisauroidea is typically regarded as a Cretaceous group, though the earliest abelisauridae remains are known from the Middle Jurassic of Argentina (classified as the species Eoabelisaurus mefi) and possibly Madagascar (fragmentary remains of ...

also have a high ascending process of the astragalus.

References

External links

Kakuru kujani
on Alphalink.com.au by ''Molnar and Pledge, 1980''.

on ''Kakuru'' Prehistoric maniraptorans Aptian life Early Cretaceous dinosaurs of Australia Paleontology in South Australia Fossil taxa described in 1980 Taxa named by Ralph Molnar {{theropod-stub