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The corsac fox (''Vulpes corsac''), also known simply as a corsac, is a medium-sized
fox Foxes are small to medium-sized, omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull, upright, triangular ears, a pointed, slightly upturned snout, and a long bushy tail (or ''brush''). Twelve sp ...
found in
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate grasslands, ...
s,
semi-desert A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi-ar ...
s and
desert A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...
s in Central Asia, ranging into Mongolia and northern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. Since 2004, it has been classified as '' least concern'' by
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
, but populations fluctuate significantly, and numbers can drop tenfold within a single year. It is also known as the steppe fox. The word "corsac" is derived from the Russian name for the animal, ''"korsák"'' (корса́к), derived ultimately from Turkic "karsak".


Description

The Corsac fox is a medium-sized fox, with a head and body length of , and a tail long. Adults weigh from . It has grey to yellowish fur over much of the body, with paler underparts and pale markings on the mouth, chin, and throat. During the winter, the coat becomes much thicker and silkier in texture, and is straw-grey in colour, with a darker line running down the back. For a fox, it has small teeth and a wide skull. One source claims that this species can climb trees and has been domesticated in the past. It is reported to have keen eyesight and hearing and an acute sense of smell. It has a number of scent glands, some of which produce pungent odors, although not so extreme as those found in some other ''Vulpes'' species. The glands are found in the anal region, above the base of the tail, and on the paws and cheeks. Corsac foxes are reported to bark during hunting or when threatening rivals, and to use higher pitch yelps or chirps as alarm calls or social greetings.


Distribution and habitat

Corsac foxes live in the
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate grasslands, ...
s and semidesert of central and northeast Asia. They are found throughout Kazakhstan, Uzbekistan, and Turkmenistan, and through all except the northernmost regions of Mongolia. In the south, their range extends into the more northern parts of Iran, Tajikistan, Kyrgyzstan, Afghanistan, and
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, and they can also be found in neighbouring regions of Russia. Three subspecies are currently recognised: * ''Vulpes corsac corsac'' - northern Kazakhstan, southern Siberia * ''V. c. kalmykorum'' - northern Uzbekistan, Caucasus * ''V. c. turkmenicus'' - southern Uzbekistan, Turkmenistan, China, Mongolia, and neighbouring regions These foxes inhabit open grassy steppes and semideserts, and avoid dense vegetation and mountainous regions. True deserts with drifting sands are also avoided, as are snowfields more than about deep. Corsac foxes generally stay far away from human disturbances.


Ecology and behavior

As an adaption to the arid climate in which they live, corsac foxes can forego food and water for extended periods of time. The corsac fox is an opportunistic forager and hunter. Its diet varies throughout its range, but consists foremost of small and medium-sized vertebrates, insects and small rodents, such as voles, gerbils, jerboas, hamsters, and ground squirrels. It also feeds opportunistically on larger prey including
hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores, and live solitarily or in pairs. They nest in slight depressions called forms, and their young are able to fend for themselves shortly after birth. The ge ...
s and pikas. It scavenges for
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
and human refuse as well. Although predominantly carnivorous, it occasionally eats fruit and other vegetation, especially during winter when availability of animal prey is low. Natural predators of the corsac fox include
gray wolf The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly ...
, Eurasian eagle-owl,
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, just ...
s, and (rarely, for pups) the upland buzzard. Golden eagles are major predators, with 38% of remains found belonging to the corsac fox at a golden eagle nest in Mongolia.Ellis, D.H., Tsengeg, P., Whitlock, P. & Ellis, M.H. 2000. ''Predators as prey at a Golden Eagle ''Aquila chrysaetos'' eyrie in Mongolia''. Ibis, 142: 139 – 141. The Corsac fox is a nocturnal and nomadic hunter of the steppes. It does not have a defended territory, and unlike some foxes, sometimes forms packs. Diurnal activity is more common at times when kits need to be fed and when food is scarce, as during the winter. Because it cannot hunt in deep snow, it either shelters in dens during harsh weather, or, in the northern parts of its range, it migrates up to south in the winter. It has been reported to follow herds of local antelope, relying on them to compress the snow as they pass. It digs its own dens, which are generally shallow, but also takes over the burrows of other animals, such as marmots, ground squirrels, or badgers. Dens may have several entrances, but are usually less than deep. The burrow is shared between the social packs, with several dens and connecting holes, which resemble "corsac cities". They are excellent climbers, but are rather slow runners and could be caught easily by a dog. While they are reported to be
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
in the wild, in captivity they are very active during the day.


Reproduction

The mating season starts in January and ends in March. Males will initially fight for access to females, but eventually establish a
monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...
bond, and assist in the raising of their young. The mother initially creates a birthing den, which is sometimes shared with other pregnant females, but moves her young to new burrows several times after they are born. Typically, two to six young are born after a
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregna ...
period of 52 to 60 days, although cases of ten kits being born in a single litter have been reported. Newborn kits weigh around , and have fluffy, light brown fur that turns yellowish as they age. They are born blind, and open their eyes at around two weeks of age; they begin to eat meat at four weeks, and emerge from the den shortly after. Corsac foxes reach sexual maturity within 9 to 10 months and reproduce in the second year of life. They live up to 9 years in the wild.


Evolution

The corsac fox is one species within a holarctic
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
of foxes that also includes the
red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the Order (biology), order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe ...
, the swift fox and the Arctic fox, all of which it resembles. However, the closest related species to the corsac fox is probably the
Tibetan sand fox The Tibetan fox (''Vulpes ferrilata''), also known as the Tibetan sand fox, is a species of true fox endemic to the high Tibetan Plateau, Ladakh plateau, Nepal, China, Sikkim, and Bhutan, up to elevations of about . It is listed as Least Concer ...
. The immediate ancestor of the corsac fox is believed to be the
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
species ''Vulpes praecorsac'', which lived in central Europe during the early Pleistocene. Fossils of corsac foxes date back to the mid-Pleistocene, and show the species once reached as far west as Switzerland, and as far south as Crimea. Very recent subfossil remains dating to 3000 to 1000 B.C.E. have in found in the Crimea and 0 to 1500 B.C.E. in the
Donev River Donev is a Bulgarian surname. Notable people with the surname include: *Asparuh Nikodimov, Asparuh "Paro" Donev Nikodimov (born 1945), former Bulgarian football player and coach *Doncho Donev, retired Bulgarian professional footballer *Donyo Donev, ...
Basin in the northern Black Sea region.


Threats

The major threat posed to the corsac fox is poaching, as it is a valuable fur-bearing species and has been harvested by humans since the Bronze Age for subsistence and commercial purposes. It is a slow runner and therefore easily caught by hunters; the population has been reduced in areas where it has been heavily hunted for its fur. Traditionally, the corsac fox is often hunted with trained domestic dogs, saker falcons, and golden eagles, as well as caught with traps set at burrow entrances, leghold traps, and firearms. In the late 19th century, up to 10,000 foxes were killed annually for pelt trade. The general population remains healthy, however, as the corsac fox has proven to be able to withstand great hunting pressures, and their habitats remain intact due to the low human population density in its range. Other threats include overgrazing by livestock and landscape development; the decline of marmots may also impact the species in some areas, as it often uses marmot burrows as daytime resting locations. The other main threat is natural disasters, which can cause the numbers of foxes to drop 90% in some areas, but the population often recovers quickly. , the corsac fox is listed as least concern in the IUCN Red List.


References


External links

* {{Taxonbar, from=Q187024 corsac fox Mammals of Afghanistan Mammals of Mongolia Mammals of China Mammals of Central Asia corsac fox Taxa named by Carl Linnaeus