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''Taxus baccata'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
tree in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Taxaceae, native to western, central and southern Europe (including Britain and Ireland), northwest Africa, northern Iran, and southwest Asia.Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins . It is the tree originally known as yew, though with other related trees becoming known, it may now be known as common yew, English yew, or European yew. It is primarily grown as an ornamental. Most parts of the plant are poisonous, with toxins that can be absorbed through inhalation and through the skin; consumption of even a small amount of the foliage can result in death.


Taxonomy and naming

The word ''yew'' is from
Proto-Germanic Proto-Germanic (abbreviated PGmc; also called Common Germanic) is the reconstructed proto-language of the Germanic branch of the Indo-European languages. Proto-Germanic eventually developed from pre-Proto-Germanic into three Germanic bran ...
''*īwa-'', possibly originally a loanword from Gaulish ''*ivos'', compare Breton ''ivin,'' Irish '' ēo'', Welsh ''ywen'', French '' if'' (see
Eihwaz Eiwaz or Eihaz is the reconstructed Proto-Germanic name of the rune , coming from a word for "yew". Two variants of the word are reconstructed for Proto-Germanic, ''*īhaz'' (''*ē2haz'', from Proto-Indo-European '), continued in Old English as ...
for a discussion). In German it is known as ''Eibe''. ''Baccata'' is Latin for ''bearing berries''. The word ''yew'' as it was originally used seems to refer to the colour brown. The yew (μίλος) was known to
Theophrastus Theophrastus (; grc-gre, Θεόφραστος ; c. 371c. 287 BC), a Greek philosopher and the successor to Aristotle in the Peripatetic school. He was a native of Eresos in Lesbos.Gavin Hardy and Laurence Totelin, ''Ancient Botany'', Routle ...
, who noted its preference for mountain coolness and shade, its evergreen character and its slow growth. Most Romance languages, with the notable exception of French (''if''), kept a version of the Latin word ''taxus'' (Italian ''tasso'', Corsican ''tassu'', Occitan ''teis'', Catalan ''teix'', Gasconic ''tech'', Spanish ''tejo'', Asturian ''texu'', Portuguese ''teixo'', Galician ''teixo'' and Romanian ''tisă'') from the same root as ''toxic''. In Slavic languages, the same root is preserved: Polish ''cis'', Ukrainian, Slovakian and Russian ''tis'' (''тис''), Serbian-Croatian-Bosnian-Montenegrin ''tisa''/''тиса'', ''tis'', Slovenian ''tisa''. Albanian borrowed it as ''tis''. In
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
, the tree is known as ''sorkhdār'' ( fa, سرخدار, literally "the red tree"). The common yew was one of the many species first described by
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
. It is one of around 30 conifer species in seven genera in the family
Taxaceae Taxaceae (), commonly called the yew family, is a coniferous family which includes six extant and two extinct genera, and about 30 species of plants, or in older interpretations three genera and 7 to 12 species. Description They are many-bran ...
, which is placed in the order
Pinales The order Pinales in the division Pinophyta, class Pinopsida, comprises all the extant conifers. The distinguishing characteristic is the reproductive structure known as a cone produced by all Pinales. All of the extant conifers, such as cedar, ...
.


Description

Yews are small to medium-sized evergreen trees, growing (exceptionally up to ) tall, with a trunk up to (exceptionally ) in diameter. The bark is thin, scaly brown, and comes off in small flakes aligned with the stem. The leaves are flat, dark green, long, broad, and
arranged In music, an arrangement is a musical adaptation of an existing composition. Differences from the original composition may include reharmonization, melodic paraphrasing, orchestration, or formal development. Arranging differs from orchest ...
spirally on the stem, but with the leaf bases twisted to align the leaves in two flat rows on either side of the stem, except on erect leading shoots where the spiral arrangement is more obvious. The leaves are poisonous.Mitchell, A. F. (1972). ''Conifers in the British Isles''. Forestry Commission Booklet 33. The
seed cones A conifer cone (in formal botanical usage: strobilus, plural strobili) is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in conifers an ...
are modified, each cone containing a single seed, which is long, and partly surrounded by a fleshy scale which develops into a soft, bright red berry-like structure called an aril. The aril is long and wide and open at the end. The arils mature 6 to 9 months after pollination, and with the seed contained, are eaten by
thrushes The thrushes are a passerine bird family, Turdidae, with a worldwide distribution. The family was once much larger before biologists reclassified the former subfamily Saxicolinae, which includes the chats and European robins, as Old World flyca ...
,
waxwing The waxwings are three species of passerine birds classified in the genus ''Bombycilla''. They are pinkish-brown and pale grey with distinctive smooth plumage in which many body feathers are not individually visible, a black and white eyestripe, ...
s and other birds, which disperse the hard seeds undamaged in their droppings. Maturation of the arils is spread over 2 to 3 months, increasing the chances of successful seed dispersal. The seeds themselves are poisonous and bitter, but are opened and eaten by some bird species, including hawfinches, greenfinches, and
great tit The great tit (''Parus major'') is a passerine bird in the tit family Paridae. It is a widespread and common species throughout Europe, the Middle East, Central Asia and east across the Palearctic to the Amur River, south to parts of North Af ...
s. The aril is not poisonous; it is gelatinous and very sweet tasting. The male cones are globose, in diameter, and shed their pollen in early spring. Yews are mostly dioecious, but occasional individuals can be variably monoecious, or change sex with time.Dallimore, W., & Jackson, A. B. (1966). ''A Handbook of Coniferae and Ginkgoaceae'' 4th ed. Arnold.


Distribution and habitat

''T. baccata'' is native to all countries of Europe (except Iceland), the Caucasus, and beyond from Turkey eastwards to northern Iran. Its range extends south to Morocco and Algeria in North Africa. A few populations are also present in the archipelagos of the Azores and Madeira. The limit of its northern
Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, Norway, and Swe ...
n distribution is its sensitivity to frost, with
global warming In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
predicted to allow its spread inland. It has been introduced elsewhere, including the United States. ''T. baccata''s richest central European populations are in Swiss yew-beech woodlands, on cool, steep marl slopes up to in elevation in the Jura Mountains and
Alpine foothills The Alpine foothills, or Prealps (german: Voralpen; french: Préalpes; it, Prealpi; ), may refer generally to any foothills at the base of the Alps in Europe. They are the transition zone between the High Alps and the Swiss Plateau and the Bavar ...
. In
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
it grows best in steep slopes of the chalk downs, forming extensive stands invading the grassland outside the beech woods. In more continental climates of Europe it fares better in
mixed forests Temperate broadleaf and mixed forest is a temperate climate terrestrial habitat type defined by the World Wide Fund for Nature, with broadleaf tree ecoregions, and with conifer and broadleaf tree mixed coniferous forest ecoregions. These fo ...
, of both coniferous and mixed broadleaf-conifer composition. Under its evergreen shade, no other plants can grow. ''T. baccata'' prefers steep rocky
calcareous Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an ad ...
slopes. It rarely develops beyond saplings on acid soil when under a forest canopy, but is tolerant of soil pH when planted by humans, such as their traditional placement in churchyards and cemeteries, where some of the largest and oldest trees in northwestern Europe are found. It grows well in well-drained soils, tolerating nearly any soil type, typically humus and
base-rich In ecology, base-richness is the level of Base (chemistry), chemical bases in water or soil, such as calcium or magnesium ions. Many organisms prefer base-rich environments. Chemical bases are alkalis, hence base-rich environments are either pH, ...
soils, but also on
rendzina Rendzina (or ''rendsina'') is a soil type recognized in various soil classification systems, including those of Britain and Germany as well as some obsolete systems. They are humus-rich shallow soils that are usually formed from carbonate- or occ ...
and
sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural class o ...
soils given adequate moisture. They can survive temporary flooding and moderate droughts. Roots can penetrate extremely compressed soils, such as on rocky terrain and vertical cliff faces. ''T. baccata'' normally appears individually or in small groups within the
understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abo ...
, but also forms stands throughout its range, such as in sheltered calcareous sites. ''T. baccata'' is extremely shade-tolerant, with the widest temperature range for
photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...
among European trees, able to photosynthesize in winter after deciduous trees have shed their leaves. It can grow under partial canopies of beech and other
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
broad-leafed trees, though it only grows into large trees without such shade. In centuries past ''T. baccata'' was exterminated from many woodlands as a poisonous hazard to the cattle and horses that often grazed in the woods. Rabbits and deer however have a level of immunity to the poisonous alkaloids, and the seeds are dispersed by birds, with
thrushes The thrushes are a passerine bird family, Turdidae, with a worldwide distribution. The family was once much larger before biologists reclassified the former subfamily Saxicolinae, which includes the chats and European robins, as Old World flyca ...
greatly enjoying the fruit. It also regenerates readily from stumps and roots, even when ancient and hollow.


Longevity

''Taxus baccata'' can reach 400 to 600 years of age. Some specimens live longer but the age of yews is often overestimated. Ten yews in Britain are believed to predate the 10th century. The potential age of yews is impossible to determine accurately and is subject to much dispute. There is rarely any wood as old as the entire tree, while the boughs themselves often become hollow with age, making ring counts impossible. Evidence based on growth rates and archaeological work of surrounding structures suggests the oldest yews, such as the
Fortingall Yew The Fortingall Yew is an ancient European yew (''Taxus baccata'') in the churchyard of the village of Fortingall in Perthshire, Scotland. Considered one of the oldest trees in Britain, modern estimates place its age at an average of 5,000 years ...
in Perthshire, Scotland, may be in the range of 2,000 years,Harte, J. (1996). How old is that old yew? ''At the Edge'' 4: 1–
online
Kinmonth, F. (2006). Ageing the yew – no core, no curve? ''International Dendrology Society Yearbook'' 2005: 41–46.Lewington, A., & Parker, E. (1999). ''Ancient Trees: Trees that Live for a Thousand Years''. London: Collins & Brown Ltd. placing them among the oldest plants in Europe. One characteristic contributing to yews' longevity is that unlike most other trees they are able to split under the weight of advanced growth without succumbing to disease in the fracture. Another is their ability to give rise to new epicormic and
basal shoot Basal shoots, root sprouts, adventitious shoots, and suckers are words for various kinds of shoots that grow from adventitious buds on the base of a tree or shrub, or from adventitious buds on its roots. Shoots that grow from buds on the base ...
s from cut surfaces and low on their trunks, even in old age.


Significant trees

The
Fortingall Yew The Fortingall Yew is an ancient European yew (''Taxus baccata'') in the churchyard of the village of Fortingall in Perthshire, Scotland. Considered one of the oldest trees in Britain, modern estimates place its age at an average of 5,000 years ...
in Perthshire, Scotland, has the largest recorded trunk girth in Britain and experts estimate it to be 2,000 to 3,000 years old, although it may be a remnant of a post-Roman Christian site and around 1,500 years old. The
Llangernyw Yew The Llangernyw Yew () is an ancient yew (''Taxus baccata'') in the village of Llangernyw, Conwy, North Wales. The tree is fragmented and its core part has been lost, leaving several enormous offshoots. The girth of the tree at the ground level is ...
in Clwyd, Wales, can be found at an early saint site and is about 1,500 years old. Other well known yews include the Ankerwycke Yew, the Balderschwang Yew, the
Caesarsboom Caesarsboom (Caesar's Tree) is a very old tree whose precise age is unknown but is believed to be over 2000 years in age. The ancient tree grows in Lo, a town in Lo-Reninge, a municipality of the province of West-Vlaanderen of Belgium. Its spec ...
, the Florence Court Yew, and the Borrowdale Fraternal Four, of which poet
William Wordsworth William Wordsworth (7 April 177023 April 1850) was an English Romantic poet who, with Samuel Taylor Coleridge, helped to launch the Romantic Age in English literature with their joint publication '' Lyrical Ballads'' (1798). Wordsworth's ' ...
wrote. The Kingley Vale National Nature Reserve in West Sussex has one of Europe's largest yew woodlands. The oldest specimen in Spain is located in Bermiego,
Asturias Asturias (, ; ast, Asturies ), officially the Principality of Asturias ( es, Principado de Asturias; ast, Principáu d'Asturies; Galician-Asturian: ''Principao d'Asturias''), is an autonomous community in northwest Spain. It is coextensiv ...
. It is known as in the
Asturian language Asturian (; ,Art. 1 de lLey 1/1998, de 23 de marzo, de uso y promoción del bable/asturiano aw 1/93, of March 23, on the Use and Promotion of the Asturian Language/nowiki> formerly also known as ) is a West Iberian languages, West Iberian ...
. It stands tall with a trunk diameter of and a crown diameter of . It was declared a Natural Monument on April 27, 1995, by the Asturian Government and is protected by the Plan of Natural Resources. Contains Word document "Monumento Natural Teixu de Bermiego". A unique forest formed by ''Taxus baccata'' and European box (''
Buxus sempervirens ''Buxus sempervirens'', the common box, European box, or boxwood, is a species of flowering plant in the genus ''Buxus'', native to western and southern Europe, northwest Africa, and southwest Asia, from southern England south to northern Morocco ...
'') lies within the city of Sochi, in the
Western Caucasus The Western Caucasus is a western region of the Caucasus in Southern Russia, extending from the Black Sea to Mount Elbrus. World Heritage Site The Western Caucasus includes a natural UNESCO World Heritage Site comprising the extreme western e ...
. The oldest Irish Yew (''Taxus baccata'' 'Fastigiata'), the Florence Court Yew, still stands in the grounds of the
Florence Court Florence Court is a large 18th-century house and estate located 8 miles south-west of Enniskillen, County Fermanagh, Northern Ireland. It is set in the foothills of Cuilcagh Mountain. The nearby village is distinguished by the one-word name F ...
estate in
County Fermanagh County Fermanagh ( ; ) is one of the thirty-two counties of Ireland, one of the nine counties of Ulster and one of the six counties of Northern Ireland. The county covers an area of 1,691 km2 (653 sq mi) and has a population of 61,805 ...
,
Northern Ireland Northern Ireland ( ga, Tuaisceart Éireann ; sco, label= Ulster-Scots, Norlin Airlann) is a part of the United Kingdom, situated in the north-east of the island of Ireland, that is variously described as a country, province or region. Nort ...
. The Irish Yew has become ubiquitous in cemeteries across the world, and it is believed that all known examples are from cuttings from this tree.


Toxicity

The entire yew bush is poisonous, with the exception of the aril (the red flesh of the berry covering the seed). Yews contain numerous toxic compounds, including "at least ten alkaloids, nitriles (
cyanogenic glycoside In chemistry, a glycoside is a molecule in which a sugar is bound to another functional group via a glycosidic bond. Glycosides play numerous important roles in living organisms. Many plants store chemicals in the form of inactive glycosides. ...
esters),
ephedrine Ephedrine is a central nervous system (CNS) stimulant that is often used to prevent low blood pressure during anesthesia. It has also been used for asthma, narcolepsy, and obesity but is not the preferred treatment. It is of unclear benefit in ...
", and their essential oil, but the most important toxins are
taxine alkaloids Taxine alkaloids, which are often named under the collective title of taxines, are the toxic chemicals that can be isolated from the yew tree. The amount of taxine alkaloids depends on the species of yew, with ''Taxus baccata'' and ''Taxus cuspi ...
,
cardiotoxic Cardiotoxicity is the occurrence of heart dysfunction as electric or muscle damage, resulting in heart toxicity. The heart becomes weaker and is not as efficient in pumping blood. Cardiotoxicity may be caused by chemotherapy (a usual example is th ...
chemical compounds which act via calcium and sodium channel antagonism. If any leaves or seeds of the plant are ingested, urgent medical attention is recommended as well as observation for at least 6 hours after the point of ingestion. The European yew is one of the most toxic species in the genus, along with the Japanese yew, ''T. cuspidata''. Yew poisonings are relatively common in both domestic and wild animals which consume the plant accidentally, resulting in "countless fatalities in livestock". Taxines are also absorbed efficiently via the skin. ''Taxus'' species should thus be handled with care and preferably with gloves. "The lethal dose for an adult is reported to be 50 g of yew needles. Patients who ingest a lethal dose frequently die due to cardiogenic shock, in spite of resuscitation efforts." There are currently no known antidotes for yew poisoning, but drugs such as atropine have been used to treat the symptoms. Taxine remains in the plant all year, with maximal concentrations appearing during the winter. Dried yew plant material retains its toxicity for several months and even increases its toxicity as the water is removed. Fallen leaves should therefore also be considered toxic. Poisoning usually occurs when leaves of yew trees are eaten, but in at least one case a victim inhaled sawdust from a yew tree.


Allergenic potential

Male yews are extremely allergenic, blooming and releasing abundant amounts of pollen in the spring, with an OPALS allergy scale rating of 10 out of 10. Completely female yews have an OPALS rating of 1, the lowest possible, trapping pollen while producing none. The pollen, like most species', easily passes through window screens. While yew pollen does not contain sufficient taxine alkaloids to cause poisoning, its allergenic potential has been implicated in adverse reactions to
paclitaxel Paclitaxel (PTX), sold under the brand name Taxol among others, is a chemotherapy medication used to treat a number of types of cancer. This includes ovarian cancer, esophageal cancer, breast cancer, lung cancer, Kaposi's sarcoma, cervical canc ...
treatment.


Traditions


Historic suicides

In the ancient Celtic world, the yew tree (''*eburos'') had extraordinary importance; a passage by
Caesar Gaius Julius Caesar (; ; 12 July 100 BC – 15 March 44 BC), was a Roman general and statesman. A member of the First Triumvirate, Caesar led the Roman armies in the Gallic Wars before defeating his political rival Pompey in a civil war, an ...
narrates that
Cativolcus Cativolcus or Catuvolcus (died 53 BC) was king of half of the country of the Eburones, a people between the Meuse and Rhine rivers, united with Ambiorix, the other king, in the insurrection against the Romans in 54 BC; but when Julius Caesar in t ...
, chief of the
Eburones The Eburones (Greek: ) were a Gallic- Germanic tribe dwelling in the northeast of Gaul, in what is now the southern Netherlands, eastern Belgium and the German Rhineland, in the period immediately preceding the Roman conquest of the region. Thou ...
, poisoned himself with yew rather than submit to Rome (''Gallic Wars'' 6: 31). Similarly, Florus notes that when the Cantabrians were under siege by the legate Gaius Furnius in 22 BC, most of them took their lives either by sword, fire, or a poison extracted ''ex arboribus taxeis'', that is, from the yew tree (2: 33, 50–51). In a similar way, Orosius notes that when the
Astures The Astures or Asturs, also named Astyrs, were the Hispano-Celtic inhabitants of the northwest area of Hispania that now comprises almost the entire modern autonomous community of Principality of Asturias, the modern province of León, and the ...
were besieged at ''Mons Medullius'', they preferred to die by their own swords or by yew poison rather than surrender (6, 21, 1).


Place names

The word ''
York York is a cathedral city with Roman origins, sited at the confluence of the rivers Ouse and Foss in North Yorkshire, England. It is the historic county town of Yorkshire. The city has many historic buildings and other structures, such as a ...
'' ( non,
Jórvík Scandinavian York ( non, Jórvík) Viking Yorkshire or Norwegian York is a term used by historians for the south of Northumbria (modern-day Yorkshire) during the period of the late 9th century and first half of the 10th century, when it was do ...
) is derived from the
Brittonic Brittonic or Brythonic may refer to: *Common Brittonic, or Brythonic, the Celtic language anciently spoken in Great Britain *Brittonic languages, a branch of the Celtic languages descended from Common Brittonic *Britons (Celtic people) The Br ...
name ( Latinised variously as , , or ), a combination of "yew-tree" (compare
Old Irish Old Irish, also called Old Gaelic ( sga, Goídelc, Ogham script: ᚌᚑᚔᚇᚓᚂᚉ; ga, Sean-Ghaeilge; gd, Seann-Ghàidhlig; gv, Shenn Yernish or ), is the oldest form of the Goidelic/Gaelic language for which there are extensive writt ...
"yew-tree" (
Irish Irish may refer to: Common meanings * Someone or something of, from, or related to: ** Ireland, an island situated off the north-western coast of continental Europe ***Éire, Irish language name for the isle ** Northern Ireland, a constituent unit ...
, , ;
Scottish Gaelic Scottish Gaelic ( gd, Gàidhlig ), also known as Scots Gaelic and Gaelic, is a Goidelic language (in the Celtic branch of the Indo-European language family) native to the Gaels of Scotland. As a Goidelic language, Scottish Gaelic, as well ...
), Welsh "alder buckthorn", Breton "alder buckthorn") and a suffix of
appurtenance An appurtenance is something subordinate to or belonging to another larger, principal entity, that is, an adjunct, satellite or accessory that generally accompanies something else.Xavier Delamarre, ''Dictionnaire de la langue gauloise'', éditions errance 2003, p. 159. meaning either "place of the yew trees" ( in Welsh, Old Irish "grove of yew trees, place with one or more yew trees", in Irish Gaelic and in Scottish Gaelic; the city itself is called (Irish) and in those languages, from the Latin ); or alternatively, "the settlement of (a man named) " (a Celtic personal name is mentioned in different documents as , , and and, when combined with the Celtic possessive suffix , could be used to denote his property). The name became the Anglian in the 7th century: a compound of , from the old name, and "a village", probably by conflation of the element with a Germanic root ('boar'); by the 7th century the
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlers in the mid-5th c ...
for 'boar' had become . When the
Danish Danish may refer to: * Something of, from, or related to the country of Denmark People * A national or citizen of Denmark, also called a "Dane," see Demographics of Denmark * Culture of Denmark * Danish people or Danes, people with a Danish a ...
army conquered the city in 866, its name became . The
Old French Old French (, , ; Modern French: ) was the language spoken in most of the northern half of France from approximately the 8th to the 14th centuries. Rather than a unified language, Old French was a linkage of Romance dialects, mutually intellig ...
and Norman name of the city following the
Norman Conquest The Norman Conquest (or the Conquest) was the 11th-century invasion and occupation of England by an army made up of thousands of Norman, Breton, Flemish, and French troops, all led by the Duke of Normandy, later styled William the Con ...
was recorded as (modern Norman ) in works such as Wace's Roman de Rou. , meanwhile, gradually reduced to ''York'' in the centuries after the Conquest, moving from the
Middle English Middle English (abbreviated to ME) is a form of the English language that was spoken after the Norman conquest of 1066, until the late 15th century. The English language underwent distinct variations and developments following the Old English ...
in the 14th century through in the 16th century to ''Yarke'' in the 17th century. The form ''York'' was first recorded in the 13th century. Many company and place names, such as the Ebor race meeting, refer to the Latinised Brittonic, Roman name. The 12th‑century chronicler
Geoffrey of Monmouth Geoffrey of Monmouth ( la, Galfridus Monemutensis, Galfridus Arturus, cy, Gruffudd ap Arthur, Sieffre o Fynwy; 1095 – 1155) was a British cleric from Monmouth, Wales and one of the major figures in the development of British historiography ...
, in his fictional account of the prehistoric kings of Britain, , suggests the name derives from that of a pre-Roman city founded by the legendary king Ebraucus. The
Archbishop of York The archbishop of York is a senior bishop in the Church of England, second only to the archbishop of Canterbury. The archbishop is the diocesan bishop of the Diocese of York and the metropolitan bishop of the province of York, which covers th ...
uses ''Ebor'' as his surname in his signature. The area of
Ydre Ydre Municipality (''Ydre kommun'') is a municipality in Östergötland County, southeast Sweden. The municipal seat is located in the town of Österbymo. The municipality was created through the local government reform of 1952, when six former ...
in the South Swedish highlands is interpreted to mean "place of yews". Two localities in particular, Idhult and Idebo, appear to be further associated with yews.


Religion

The yew is traditionally and regularly found in churchyards in England, Wales, Scotland, Ireland, and Northern France (particularly
Normandy Normandy (; french: link=no, Normandie ; nrf, Normaundie, Nouormandie ; from Old French , plural of ''Normant'', originally from the word for "northman" in several Scandinavian languages) is a geographical and cultural region in Northwestern ...
). Some examples can be found in La Haye-de-Routot or
La Lande-Patry La Lande-Patry () is a commune in the Orne department in north-western France. The commune is known for its ''Ifs millénaires'' (thousand year old yews), designated as an intangible cultural heritage in France. See also *Communes of the Orne d ...
. It is said up to 40 people could stand inside one of the La-Haye-de-Routot yew trees, and the Le Ménil-Ciboult yew is probably the largest, with a girth of 13 m. Yews may grow to become exceptionally large (over 5 m diameter) and may live to be over 2,000 years old. Sometimes monks planted yews in the middle of their
cloister A cloister (from Latin ''claustrum'', "enclosure") is a covered walk, open gallery, or open arcade running along the walls of buildings and forming a quadrangle or garth. The attachment of a cloister to a cathedral or church, commonly against a ...
, as at Muckross Abbey (Ireland) or abbaye de Jumièges (Normandy). Some ancient yew trees are located at St. Mary the Virgin Church, Overton-on-Dee in Wales. In the
Septuagint The Greek Old Testament, or Septuagint (, ; from the la, septuaginta, lit=seventy; often abbreviated ''70''; in Roman numerals, LXX), is the earliest extant Greek translation of books from the Hebrew Bible. It includes several books beyond ...
rendering of the
Book of Nahum The Book of Nahum is the seventh book of the 12 minor prophets of the Hebrew Bible. It is attributed to the prophet Nahum, and was probably written in Jerusalem in the 7th century BC. Background Josephus places Nahum during the reign of Jotham ...
, 1:10, Nineveh and other deemed enemies of the biblical God are foretold to "be laid bare even to its foundation, and…devoured as a twisted yew." In Asturian tradition and culture, the yew tree was considered to be linked with the land, people, ancestors, and ancient religion. It was tradition on
All Saints' Day All Saints' Day, also known as All Hallows' Day, the Feast of All Saints, the Feast of All Hallows, the Solemnity of All Saints, and Hallowmas, is a Christian solemnity celebrated in honour of all the saints of the church, whether they are kn ...
to bring a branch of a yew tree to the tombs of those who had died recently so they would be guided in their return to the Land of Shadows. The yew tree has been found near chapels, churches, and cemeteries since ancient times as a symbol of the transcendence of death. They are often found in the main squares of villages where people celebrated the open councils that served as a way of general assembly to rule village affairs. It has been suggested that the
sacred tree A sacred tree is a tree which is considered to be sacred, or worthy of spiritual respect or reverence. Such trees appear throughout world history in various cultures including the ancient Greek, Hindu mythology, Celtic and Germanic mythologies. T ...
at the
Temple at Uppsala The Temple at Uppsala was a religious center in the ancient Norse religion once located at what is now Gamla Uppsala (Swedish "Old Uppsala"), Sweden attested in Adam of Bremen's 11th-century work '' Gesta Hammaburgensis ecclesiae pontificum'' and ...
was an ancient yew tree. The Christian church commonly found it expedient to take over existing pre-Christian sacred sites for churches. It has also been suggested that yews were planted at religious sites as their long life was suggestive of eternity, or because, being toxic when ingested, they were seen as trees of death. Another suggested explanation is that yews were planted to discourage farmers and drovers from letting animals wander onto the burial grounds, the poisonous foliage being the disincentive. A further possible reason is that fronds and branches of yew were often used as a substitute for palms on
Palm Sunday Palm Sunday is a Christian moveable feast that falls on the Sunday before Easter. The feast commemorates Christ's triumphal entry into Jerusalem, an event mentioned in each of the four canonical Gospels. Palm Sunday marks the first day of Hol ...
. King
Edward I of England Edward I (17/18 June 1239 – 7 July 1307), also known as Edward Longshanks and the Hammer of the Scots, was King of England and Lord of Ireland from 1272 to 1307. Concurrently, he ruled the duchies of Aquitaine and Gascony as a vas ...
ordered yew trees planted in churchyards to protect the buildings. Some yews existed before their churches, as preachers held services beneath them when churches were unavailable. Due to the ability of their branches to root and sprout anew after touching the ground, yews became symbols of death, rebirth, and therefore immortality. In interpretations of
Norse cosmology Norse cosmology is the study of the cosmos (cosmology) as perceived by the ancient North Germanic peoples. The topic encompasses concepts from Norse mythology, such as notions of time and space, cosmogony, personifications, anthropogeny, and e ...
, the tree ''
Yggdrasil Yggdrasil (from Old Norse ), in Norse cosmology, is an immense and central sacred tree. Around it exists all else, including the Nine Worlds. Yggdrasil is attested in the ''Poetic Edda'' compiled in the 13th century from earlier traditional ...
'' has traditionally been interpreted as a giant ash tree. Some scholars now believe errors were made in past interpretations of the ancient writings, and that the tree is most likely a European yew (''Taxus baccata''). In the '' Crann Ogham''—the variation on the ancient Irish ''
Ogham Ogham ( Modern Irish: ; mga, ogum, ogom, later mga, ogam, label=none ) is an Early Medieval alphabet used primarily to write the early Irish language (in the "orthodox" inscriptions, 4th to 6th centuries AD), and later the Old Irish langu ...
'' alphabet which consists of a list of trees—yew is the last in the main list of 20 trees, primarily symbolizing death. There are stories of people who have committed suicide by ingesting the foliage. As the ancient Celts also believed in the
transmigration of the soul Reincarnation, also known as rebirth or transmigration, is the philosophical or religious concept that the non-physical essence of a living being begins a new life in a different physical form or body after biological death. Resurrection is a ...
, there is in some cases a secondary meaning of the eternal soul that survives death to be reborn in a new form.


Uses

Yew wood was historically important, finding use in the
Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the post-classical period of global history. It began with the fall of the Western Roman Empire ...
in items such as musical instruments, furniture, and
longbows A longbow (known as warbow in its time, in contrast to a hunting bow) is a type of tall bow that makes a fairly long draw possible. A longbow is not significantly recurved. Its limbs are relatively narrow and are circular or D-shaped in cross ...
. The species was felled nearly to extinction in much of Europe. In the modern day it is not considered a commercial crop due to its very slow growth, but it is valued for hedging and
topiary Topiary is the horticultural practice of training perennial plants by clipping the foliage and twigs of trees, shrubs and subshrubs to develop and maintain clearly defined shapes, whether geometric or fanciful. The term also refers to plants w ...
.


Medical

Certain compounds found in the bark of yew trees were discovered in 1967 to have efficacy as anti-cancer agents. The precursors of the
chemotherapy Chemotherapy (often abbreviated to chemo and sometimes CTX or CTx) is a type of cancer treatment that uses one or more anti-cancer drugs ( chemotherapeutic agents or alkylating agents) as part of a standardized chemotherapy regimen. Chemothe ...
drug
paclitaxel Paclitaxel (PTX), sold under the brand name Taxol among others, is a chemotherapy medication used to treat a number of types of cancer. This includes ovarian cancer, esophageal cancer, breast cancer, lung cancer, Kaposi's sarcoma, cervical canc ...
(taxol) were later shown to be synthesized easily from extracts of the leaves of European yew, which is a much more renewable source than the bark of the Pacific yew (''
Taxus brevifolia ''Taxus brevifolia'', the Pacific yew or western yew, is a species of tree in the yew family Taxaceae native to the Pacific Northwest of North America. It is a small evergreen conifer, thriving in moisture and otherwise tending to take the form ...
'') from which they were initially isolated. This ended a point of conflict in the early 1990s; many environmentalists, including
Al Gore Albert Arnold Gore Jr. (born March 31, 1948) is an American politician, businessman, and environmentalist who served as the 45th vice president of the United States from 1993 to 2001 under President Bill Clinton. Gore was the Democratic no ...
, had opposed the destructive harvesting of Pacific yew for paclitaxel cancer treatments.
Docetaxel Docetaxel (DTX or DXL), sold under the brand name Taxotere among others, is a chemotherapy medication used to treat a number of types of cancer. This includes breast cancer, head and neck cancer, stomach cancer, prostate cancer and non-small-cell ...
can then be obtained by semi-synthetic conversion from the precursors.


Woodworking

Wood from the yew is classified as a closed-pore softwood, similar to cedar and pine. Easy to work, yew is among the hardest of the softwoods, yet it possesses a remarkable elasticity, making it ideal for products that require springiness, such as bows. The wood is esteemed for
cabinetry A cabinet is a case or cupboard with shelves and/or drawers for storing or displaying items. Some cabinets are stand alone while others are built in to a wall or are attached to it like a medicine cabinet. Cabinets are typically made of wood (s ...
and tool handles. The hard, slow-growing wood also finds use in gates, furniture, parquet floors, and
paneling Panelling (or paneling in the U.S.) is a millwork wall covering constructed from rigid or semi-rigid components. These are traditionally interlocking wood, but could be plastic or other materials. Panelling was developed in antiquity to make ro ...
. Its typical
burl A burl (American English) or burr (British English) is a tree growth in which the grain has grown in a deformed manner. It is commonly found in the form of a rounded outgrowth on a tree trunk or branch that is filled with small knots from dor ...
s and contorted growth, with intricate multicolored patterns, make it attractive for carving and
woodturning Woodturning is the craft of using a wood lathe with hand-held tools to cut a shape that is symmetrical around the axis of rotation. Like the potter's wheel, the wood lathe is a simple mechanism that can generate a variety of forms. The operator ...
, but also make the wood unsuited for construction. It is good firewood and is sometimes burnt as incense. Due to all parts of the yew and its volatile oils being poisonous and cardiotoxic, a mask should be worn if one comes in contact with sawdust from the wood. One of the world's oldest surviving wooden artifacts is a
Clactonian The Clactonian is the name given by archaeologists to an industry of European flint tool manufacture that dates to the early part of the interglacial period known as the Hoxnian, the Mindel- Riss or the Holstein stages (c. 400,000 years ago). C ...
yew spear head, found in 1911 at
Clacton-on-Sea Clacton-on-Sea is a seaside town in the Tendring District in the county of Essex, England. It is located on the Tendring Peninsula and is the largest settlement in the Tendring District with a population of 56,874 (2016). The town is situated ...
, in
Essex Essex () is a county in the East of England. One of the home counties, it borders Suffolk and Cambridgeshire to the north, the North Sea to the east, Hertfordshire to the west, Kent across the estuary of the River Thames to the south, and Grea ...
, UK. Known as the Clacton Spear, it is estimated to be over 400,000 years old.


Longbows

Yew is also associated with Wales and England because of the
longbow A longbow (known as warbow in its time, in contrast to a hunting bow) is a type of tall bow that makes a fairly long draw possible. A longbow is not significantly recurved. Its limbs are relatively narrow and are circular or D-shaped in cross ...
, an early weapon of war developed in northern Europe, and as the English longbow the basis for a medieval tactical system. The oldest surviving yew longbow was found at Rotten Bottom in Dumfries and
Galloway Galloway ( ; sco, Gallowa; la, Gallovidia) is a region in southwestern Scotland comprising the historic counties of Wigtownshire and Kirkcudbrightshire. It is administered as part of the council area of Dumfries and Galloway. A native or ...
, Scotland. It has been given a calibrated radiocarbon date of 4040 BC to 3640 BC and is on display in the
National Museum of Scotland The National Museum of Scotland in Edinburgh, Scotland, was formed in 2006 with the merger of the new Museum of Scotland, with collections relating to Scottish antiquities, culture and history, and the adjacent Royal Scottish Museum (opened in ...
. Yew is the wood of choice for longbow making; the
heartwood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin t ...
is always on the inside of the bow with the sapwood on the outside. This makes most efficient use of their properties as heartwood is best in compression whilst sapwood is superior in tension. However, much yew is knotty and twisted, and therefore unsuitable for bowmaking; most trunks do not give good staves and even in a good trunk much wood has to be discarded. There was a tradition of planting yew trees in churchyards throughout Britain and Ireland, among other reasons, as a resource for bows, such as at " Ardchattan Priory whose yew trees, according to other accounts, were inspected by
Robert the Bruce Robert I (11 July 1274 – 7 June 1329), popularly known as Robert the Bruce (Scottish Gaelic: ''Raibeart an Bruis''), was King of Scots from 1306 to his death in 1329. One of the most renowned warriors of his generation, Robert eventual ...
and cut to make at least some of the
longbow A longbow (known as warbow in its time, in contrast to a hunting bow) is a type of tall bow that makes a fairly long draw possible. A longbow is not significantly recurved. Its limbs are relatively narrow and are circular or D-shaped in cross ...
s used at the Battle of Bannockburn." The trade of yew wood to England for longbows was so robust that it depleted the stocks of good-quality, mature yew over a vast area. The first documented import of yew bowstaves to England was in 1294. In 1423 the Polish king commanded protection of yews in order to cut exports, facing nearly complete destruction of local yew stock. In 1470 compulsory archery practice was renewed, and hazel, ash, and
laburnum ''Laburnum'', sometimes called golden chain or golden rain, is a genus of two species of small trees in the subfamily Faboideae of the pea family Fabaceae. The species are '' Laburnum anagyroides''—common laburnum and '' Laburnum alpinum'' ...
were specifically allowed for practice bows. Supplies still proved insufficient, until by the Statute of Westminster in 1472, every ship coming to an English port had to bring four bowstaves for every
tun TUN or tun may refer to: Biology * Tun shells, large sea snails of the family ''Tonnidae'' * Tun, a tardigrade in its cryptobiotic state * Tun or Toon, common name for trees of the genus '' Toona'' Places * Tun, Sweden, a locality in Västra G ...
. Richard III of England increased this to ten for every tun. This stimulated a vast network of extraction and supply, which formed part of royal monopolies in southern Germany and Austria. In 1483, the price of bowstaves rose from two to eight pounds per hundred, and in 1510 the Venetians would only sell a hundred for sixteen pounds. In 1507 the Holy Roman Emperor asked the
Duke of Bavaria The following is a list of rulers during the history of Bavaria. Bavaria was ruled by several dukes and kings, partitioned and reunited, under several dynasties. Since 1949, Bavaria has been a democratic state in the Federal Republic of Germ ...
to stop cutting yew, but the trade was profitable, and in 1532 the royal
monopoly A monopoly (from Greek el, μόνος, mónos, single, alone, label=none and el, πωλεῖν, pōleîn, to sell, label=none), as described by Irving Fisher, is a market with the "absence of competition", creating a situation where a speci ...
was granted for the usual quantity "if there are that many." In 1562, the Bavarian government sent a long plea to the Holy Roman Emperor asking him to stop the cutting of yew, and outlining the damage done to the forests by its selective extraction, which broke the canopy and allowed wind to destroy neighbouring trees. In 1568, despite a request from
Saxony Saxony (german: Sachsen ; Upper Saxon: ''Saggsn''; hsb, Sakska), officially the Free State of Saxony (german: Freistaat Sachsen, links=no ; Upper Saxon: ''Freischdaad Saggsn''; hsb, Swobodny stat Sakska, links=no), is a landlocked state of ...
, no royal monopoly was granted because there was no yew to cut, and the next year
Bavaria Bavaria ( ; ), officially the Free State of Bavaria (german: Freistaat Bayern, link=no ), is a state in the south-east of Germany. With an area of , Bavaria is the largest German state by land area, comprising roughly a fifth of the total lan ...
and
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
similarly failed to produce enough yew to justify a royal monopoly. Forestry records in this area in the 17th century do not mention yew, and it seems that no mature trees were to be had. The English tried to obtain supplies from the Baltic, but at this period bows were being replaced by guns in any case.


Musical instruments

The late Robert Lundberg, a noted luthier who performed extensive research on historical lute-making methodology, states in his 2002 book ''Historical Lute Construction'' that yew was historically a prized wood for lute construction. European legislation establishing use limits and requirements for yew limited supplies available to luthiers, but it was apparently as prized among medieval, renaissance, and baroque lute builders as
Brazilian rosewood ''Dalbergia nigra'', commonly known as the Bahia rosewood, jacarandá-da-Bahia, Brazilian rosewood, Rio rosewood, jacarandá-do-brasil, pianowood, caviúna, graúna, jacarandá-una or obuina is a species of legume in the family Fabaceae. Descri ...
is among contemporary guitar-makers for its quality of sound and beauty.


Horticulture

Today European yew is widely used in landscaping and ornamental horticulture. Due to its dense, dark green, mature foliage, and its tolerance of even very severe pruning, it is used especially for formal
hedge A hedge or hedgerow is a line of closely spaced shrubs and sometimes trees, planted and trained to form a barrier or to mark the boundary of an area, such as between neighbouring properties. Hedges that are used to separate a road from adjoin ...
s and
topiary Topiary is the horticultural practice of training perennial plants by clipping the foliage and twigs of trees, shrubs and subshrubs to develop and maintain clearly defined shapes, whether geometric or fanciful. The term also refers to plants w ...
. Its relatively slow growth rate means that in such situations it needs to be clipped only once per year (in late summer). European yew will tolerate a wide range of soils and situations, including shallow chalk soils and shade, although in deep shade its foliage may be less dense. However it cannot tolerate waterlogging, and in poorly-draining situations is liable to succumb to the root-rotting pathogen ''
Phytophthora cinnamomi ''Phytophthora cinnamomi'' is a soil-borne water mould that produces an infection which causes a condition in plants variously called " root rot", "dieback", or (in certain ''Castanea'' species), "ink disease". The plant pathogen is one of the wo ...
''. ''T. baccata'' is tolerant of urban pollution, cold, and heat, though soil compaction e.g. by roads can harm it. It is slow-growing, taking about 20 years to grow tall, and vertical growth effectively stops after 100 years. With its soft bark, the tree can be killed over time by rubbing such as by climbing children. In Europe, ''Taxus baccata'' grows naturally north to Molde in southern Norway, but it is used in gardens further north. It is also popular as a bonsai in many parts of Europe and makes a handsome small- to large-sized bonsai. Well over 200
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s of ''T. baccata'' have been named. The most popular of these are the
Irish yew ''Taxus baccata'' is a species of evergreen tree in the family (botany), family Taxaceae, native to western, central and southern Europe (including Britain and Ireland), northwest Africa, northern Iran, and southwest Asia.Rushforth, K. (1999). '' ...
(''T. baccata'' 'Fastigiata'), a fastigiate cultivar of the European yew selected from two trees found growing in Ireland, and the several cultivars with yellow leaves, collectively known as "golden yew". In some locations, e.g. when hemmed in by buildings or other trees, an Irish yew can reach 20 feet in height without exceeding 2 feet in diameter at its thickest point, although with age many Irish yews assume a fat cigar shape rather than being truly columnar.


AGM cultivars

The following cultivars have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's Award of Garden Merit:- *''T. baccata'' *''T. baccata'' 'Fastigiata' (Irish yew) *''T. baccata'' 'Fastigiata Aureomarginata' (golden Irish yew) *''T. baccata'' 'Icicle' *''T. baccata'' 'Repandens' *''T. baccata'' 'Repens Aurea' *''T. baccata'' 'Semperaurea' *''T. baccata'' 'Standishii'


Privies

In England, yew has historically been sometimes associated with privies (outside toilets), possibly because the smell of the plant keeps insects away.


Conservation

Historically, ''T. baccata'' populations were gravely threatened by felling for longbows and destruction to protect livestock from poisoning. It is now endangered in parts of its range due to intensive land use. The species is also harvested to meet pharmaceutical demand for taxanes. Trees are often damaged by
browsing Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...
and bark stripping. Yew's thin bark makes it vulnerable to fire. Its toxicity protects against many insects, but the yew mite causes significant bud mortality, and seedlings can be killed by fungi. Clippings from ancient specimens in the UK, including the Fortingall Yew, were taken to the Royal Botanic Gardens in Edinburgh to form a mile-long hedge. The purpose of this "Yew Conservation Hedge Project" is to maintain the DNA of ''Taxus baccata''. Another conservation programme was run in
Catalonia Catalonia (; ca, Catalunya ; Aranese Occitan: ''Catalonha'' ; es, Cataluña ) is an autonomous community of Spain, designated as a '' nationality'' by its Statute of Autonomy. Most of the territory (except the Val d'Aran) lies on the nort ...
in the early 2010s by the
Forest Sciences Centre of Catalonia A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
(CTFC) in order to protect genetically endemic yew populations and preserve them from overgrazing and forest fires. In the framework of this programme, the 4th International Yew Conference was organised in the Poblet Monastery in 2014. There has also been a conservation programme in northern Portugal and
Northern Spain Spain is a country located in southwestern Europe occupying most (about 82 percent) of the Iberian Peninsula. It also includes a small exclave inside France called Llívia, as well as the Balearic Islands in the Mediterranean, the Canary Isla ...
(Cantabrian Range).


See also

*
List of poisonous plants Plants that produce toxins are referred to as poisonous plants. Plants that cause irritation on contact are also described as "poisonous". The toxins in poisonous plants affect herbivores, and deter them from consuming the plants. Plants cannot ...
*
List of plants poisonous to equines Many plants are poisonous to equines; the species vary depending on location, climate, and grazing conditions. In many cases, entire genera are poisonous to equines and include many species spread over several continents. Plants can cause reactio ...


References


Further reading

* Chetan, A. and Brueton, D. (1994) ''The Sacred Yew'', London: Arkana, * Hartzell, H. (1991) ''The yew tree: a thousand whispers: biography of a species'', Eugene: Hulogosi, * Simón, F. M. (2005) Religion and Religious Practices of the Ancient Celts of the Iberian Peninsula,
e-Keltoi
', v. 6, p. 287-345, online *


External links


Monumentaltrees.com: Images and location details of ancient yews

Life+ TAXUS.cat: Taxus conservation programme in Catalonia
*
Forest Sciences Centre of Catalonia A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
(CTFC)
Biology Conservation Department

''Taxus baccata''
distribution map, genetic conservation units, and related resources from the
European Forest Genetic Resources Programme European Forest Genetic Resources Programme (EUFORGEN) is an international network that supports the conservation and sustainable use of forest genetic resources in Europe. The programme’s tasks include to coordinate and promote '' in situ'' an ...
(EUFORGEN) {{Authority control baccata Flora of Europe Flora of Iran Flora of North Africa Flora of the Azores Flora of the Caucasus Flora of Turkey Garden plants of Asia Garden plants of Europe Medicinal plants of Asia Medicinal plants of Europe Ornamental trees Poisonous plants Plants described in 1753 Plants used in bonsai Taxa named by Carl Linnaeus Trees of humid continental climate Trees of mild maritime climate