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Two Great Andamanese men, circa 1875

The Andamanese are the various indigenous peoples of the Andaman Islands, part of India's Andaman and Nicobar Islands union territory in the southeastern part of the Bay of Bengal in Southeast Asia. The Andamanese peoples are among the various groups considered Negrito, owing to their dark skin and diminutive stature. All Andamanese traditionally lived a hunter-gatherer lifestyle, and appear to have lived in substantial isolation for thousands of years.[1] It is suggested that the Andamanese settled in the Andaman Islands around the latest glacial maximum, around 26,000 years ago.[2][3]

The Andamanese peoples included the Great Andamanese and Jarawas of the Great Andaman archipelago, the Jangil of Rutland Island, the Onge of Little Andaman, and the Sentinelese of North Sentinel Island.[4] At the end of the 18th century, when they first came into sustained contact with outsiders, an estimated 7,000 Andamanese remained. In the next century, they were largely wiped out by diseases, violence, and loss of territory. Today, only roughly 400–450 Andamanese remain, with the Jangil being extinct. Only the Jarawa and the Sentinelese maintain a steadfast independence, refusing most attempts at contact by outsiders.

The Andamanese are a designated Scheduled Tribe in India's constitution.[5]

History

Until the late 18th century, the Andamanese culture, language, and genetics were preserved from outside influences by their fierce reaction to visitors, which included killing any shipwrecked foreigners, and by the remoteness of the islands. The various tribes and their mutually unintelligible languages thus are believed to have evolved on their own over millennia.

Origins

The Andamanese are the various indigenous peoples of the Andaman Islands, part of India's Andaman and Nicobar Islands union territory in the southeastern part of the Bay of Bengal in Southeast Asia. The Andamanese peoples are among the various groups considered Negrito, owing to their dark skin and diminutive stature. All Andamanese traditionally lived a hunter-gatherer lifestyle, and appear to have lived in substantial isolation for thousands of years.[1] It is suggested that the Andamanese settled in the Andaman Islands around the latest glacial maximum, around 26,000 years ago.[2][3]

The Andamanese peoples included the Great Andamanese and Jarawas of the Great Andaman archipelago, the Jangil of Rutland Island, the Onge of Little Andaman, and the Sentinelese of North Sentinel Island.[4] At the end of the 18th century, when they first came into sustained contact

The Andamanese peoples included the Great Andamanese and Jarawas of the Great Andaman archipelago, the Jangil of Rutland Island, the Onge of Little Andaman, and the Sentinelese of North Sentinel Island.[4] At the end of the 18th century, when they first came into sustained contact with outsiders, an estimated 7,000 Andamanese remained. In the next century, they were largely wiped out by diseases, violence, and loss of territory. Today, only roughly 400–450 Andamanese remain, with the Jangil being extinct. Only the Jarawa and the Sentinelese maintain a steadfast independence, refusing most attempts at contact by outsiders.

The Andamanese are a designated Scheduled Tribe in India's constitution.[5]

Until the late 18th century, the Andamanese culture, language, and genetics were preserved from outside influences by their fierce reaction to visitors, which included killing any shipwrecked foreigners, and by the remoteness of the islands. The various tribes and their mutually unintelligible languages thus are believed to have evolved on their own over millennia.

Origins

Map of human haplotype migration based on mitochondrial DNA, with Key (coloured) indicating periods in numbered thousands of years before the present. Note the route of the mtDNA haplogroup M through the Indian mainland and the Andaman Islands, possibly on to Southeast Asia

According to Chaubey and Endicott (2013), the Andaman Islands were settled less than 26,000 years ago, by people who were not direct descendants of the first migrants out of Africa.[6][note 1] According to Wang et al. (2011),

...the Andaman archipelago was likely settled by modern humans from northeast India via the land-bridge which connected the Andaman archipelago and Myanmar around the Last Glacial Maximum (LGM), a scenario in well agreement with the evidence from linguistic and palaeoclimate studies.[7]

It was previously assumed that the Andaman ancestors were part of the initial Great Coastal Migration that was the first expansion of humanity out of Africa, via the Arabian peninsula, along the coastal regions of the Indian mainland and toward Southeast Asia, China, and Oceania.[8][9] The Andamanese were considered to be a pristine example of a hypothesized Negrito population, which showed similar physical characteristics, and was supposed to have existed throughout southeast Asia. The existence of a specific Negrito-population is nowadays

According to Chaubey and Endicott (2013), the Andaman Islands were settled less than 26,000 years ago, by people who were not direct descendants of the first migrants out of Africa.[6][note 1] According to Wang et al. (2011),

...the Andaman archipelago was likely settled by modern humans from northeast India via the land-bridge which connected the Andaman archipelago and Myanmar around the Last Glacial Maximum (LGM), a scenario in well agreement with the evidence from linguistic and palaeoclimate studies.[7]

It was previously assumed that the Andaman ancestors were part of the initial Great Coastal Migration that was the first expansion of humanity out of Africa, via the Arabian peninsula, along the coastal regions of the Indian mainland and toward Southeast Asia, China, and Oceania.[8][9] The Andamanese were considered to be a pristine example of a hypothesized Negrito popul

...the Andaman archipelago was likely settled by modern humans from northeast India via the land-bridge which connected the Andaman archipelago and Myanmar around the Last Glacial Maximum (LGM), a scenario in well agreement with the evidence from linguistic and palaeoclimate studies.[7]

It was previously assumed that the Andaman ancestors were part of the initial Great Coastal Migration that was the first expansion of humanity out of Africa, via the Arabian penins

It was previously assumed that the Andaman ancestors were part of the initial Great Coastal Migration that was the first expansion of humanity out of Africa, via the Arabian peninsula, along the coastal regions of the Indian mainland and toward Southeast Asia, China, and Oceania.[8][9] The Andamanese were considered to be a pristine example of a hypothesized Negrito population, which showed similar physical characteristics, and was supposed to have existed throughout southeast Asia. The existence of a specific Negrito-population is nowadays doubted. Their commonalities could be the result of evolutionary convergence and/or a shared history.[10][11]

Bulbeck (20

Bulbeck (2013) shows the Andamanese maternal mtDNA is entirely mitochondrial Haplogroup M.[12] Their Y-DNA belong to the D haplogroup which has not been seen outside of the Andamans, a fact that underscores the insularity of these tribes. Analysis of mtDNA, which is inherited exclusively by maternal descent, confirms the above results. All Onge belong to mtDNA haplogroup M, which is unique to Onge people. This haplogroup, which is descended from Africa, represents the entire lineage of the Onge and Adamanese, and is the predominant mtDNA among the Semang in Malaysia. Moreover, haplogroup M is the single most common mtDNA haplogroup in Asia, where it represents 60% of all maternal lineages.[13][14]

A 2017 study by Mondal et al. finds that the Riang lineage of D1a3 (a Tibeto-Burmese population) and the Andamanese D1a3 have their nearest related lineages in East Asia. The Jarawa and Onge shared this D1a3 lineage with each other within the last ~7000 years, but had diverged from the Japanese D1a2 lineage ~53000 years ago". They further suggest that: “This strongly suggests that haplogroup D does not indicate a separate ancestry for Andamanese populations. Rather, haplogroup D was part of the standing variation carried by the OOA expansion, and later lost from most of the populations except in Andaman and partially in Japan and Tibet”.[15]

The Andamanese's protective isolation changed with the first British colonial presence and subsequent settlements, which proved disastrous for them. Lacking immunity against common infectious diseases of the Eurasian mainland, the large Jarawa habitats on the southeastern regions of South Andaman Island likely were depopulated by disease within four years (1789–1793) of the initial British colonial settlement in 1789.[16] Epidemics of pneumonia, measles and influenza spread rapidly and exacted heavy tolls, as did alcoholism.[16] In the 19th century, the measles killed 50% of the Andamanese population.[17] By 1875, the Andamanese were already "perilously close to extinction," yet attempts to contact, subdue and co-opt them continued unrelentingly. In 1888, the British government set in place a policy of "organized gift giving" that continued in varying forms until well into the 20th century.[18]

Great Andamanese men, women and children, 1876

There is evidence that some sections of the British Indian administration were working deliberately to annihilate the tribes.[19] After the mid-19th century, British established penal colonies on the islands and an increasing number of mainland Indian and Karen settlers arrived, encroaching on former territories of the Andamanese. This accelerated the d

There is evidence that some sections of the British Indian administration were working deliberately to annihilate the tribes.[19] After the mid-19th century, British established penal colonies on the islands and an increasing number of mainland Indian and Karen settlers arrived, encroaching on former territories of the Andamanese. This accelerated the decline of the tribes.

Many Andamanese succumbed to British expeditions to avenge the killing of shipwrecked sailors. In the 1867 Andaman Islands Expedition, dozens of Onge were killed by British naval personnel following the death of shipwrecked sailors, which resulted in four Victoria Crosses for the British soldiers.[20][21][22]

In 1923, the British ornithologist and anthropologist Many Andamanese succumbed to British expeditions to avenge the killing of shipwrecked sailors. In the 1867 Andaman Islands Expedition, dozens of Onge were killed by British naval personnel following the death of shipwrecked sailors, which resulted in four Victoria Crosses for the British soldiers.[20][21][22]

In 1923, the British ornithologist and anthropologist Frank Finn, who visited the islands in the 1890s, while working for the Indian Museum, described the Andamanese as "The World's Most Primitive People", writing:[23]

I used to envy the pigmies their simple costume, which in the case of the ladies was a wisp and a waistband, and in that of the men, nothing at all. Their interests are looked after by an English Civil Servant, who has to see that no one sells them drink, or interferes with them in any way; but even this officer-in-charge, as he is styled, dares not go among them where he is not known, and considerable tact is required in getting an introduction to the local chief.

In the 1940s, the Jarawa were bombed by imperial Japanese forces for their hostility. This attack of the Japanese was criticized as war crime by many observers.[24]

In 1974, a film crew and anthropologist Triloknath Pandit attempted friendly contact by leaving a tethered pig, some pots and pans, some fruit, and toys on the beach at North Sentinel Island. One of the islanders shot the film director in the thigh with an arrow. The following year, European visitors were repulsed with arrows.[25][26][27]

On 2 August 1981, the Hong Kong freighter ship Primrose grounded on the North Sentinel Island reef. A few days later, crewmen on the immobile vessel observed that small black men were carrying spears and arrows and building boats on the beach. The captain of the Primrose radioed for an urgent airdrop of fire

On 2 August 1981, the Hong Kong freighter ship Primrose grounded on the North Sentinel Island reef. A few days later, crewmen on the immobile vessel observed that small black men were carrying spears and arrows and building boats on the beach. The captain of the Primrose radioed for an urgent airdrop of firearms so the crew could defend themselves, but did not receive them. Heavy seas kept the islanders away from the ship. After a week, the crew were rescued by an Indian navy helicopter.[28]

On 4 January 1991, Triloknath Pandit made the first known friendly contact with the Sentinelese.[27]

Until 1996, the Jarawa met most visitors with flying arrows. From time to time, they attacked and killed poachers on the lands reserved to them by the Indian government. They also killed some workers building the Andaman Trunk Road (ATR), which traverses Jarawa lands. One of the earliest peaceful contacts with the Jarawa occurred in 1996. Settlers found a teenaged Jarawa boy named Enmei near Kadamtala town. The boy was immobilized with a broken foot. They took Enmei to a hospital, where he received good care. Over several weeks, Enmei learned a few words of Hindi before returning to his jungle home. The following year, Jarawa individuals and small groups began appearing along roadsides and occasionally venturing into settlements to steal food. The ATR may have interfered with traditional Jarawa food sources.[29][30][31]

On 17 November 2018, a United States missionary, John Allen Chau, was killed when he tried to introduce Christianity to the Sentinelese tribe. The Sentinelese have been protected from contact with the outside world. Trips to the Island are prohibited by Indian law.[32] Chau was brought near the island by local fishermen, who were later arrested during the investigation into his death.[33] Indian authorities attempted to retrieve Chau's remains without success.[34]

The five major groups of Andamanese are:

  • Great Andamanese, traditionally of the Great Andaman archipelago but now living on Strait Island. Their population was 52 as of 2010.[35]
  • Jarawa traditionally of the southern part of South Andaman Island in the Great Andaman archipelago but now living in the ex-Great Andamanese homeland in the West Coast and central parts of South and Middle Andaman Islands. Their population was 380 as of 2011.[36]
  • By the end of the eighteenth century, there were an estimated 5,000 Great Andamanese living on Great Andaman. Altogether they comprised ten distinct tribes with different languages. The population quickly dwindled to 600 in 1901 and to 19 by 1961.[37] It has increased slowly after that, following their move to a reservation on Strait Island. As of 2010, the population was 52, representing a mix of the former tribes.[35]

    The Jarawa originally inhabited southeastern Jarawa Island and have migrated to the west coast of Great Andaman in the wake of the Great Andamanese. The Onge once lived throughout Little Andaman and now are confined to two reservations on the island. The Jangil, who originally inhabited Rutland Island, were extinct by 1931; the last individual was sighted in 1907.[24] Only the Sentinelese are still living in their original homeland on North Sentinel Island, largely undisturbed, and have fiercely resisted all attempts at contact.

    Languages

    The Andamanese languages are considered to be the fifth language family of India, following the Indo-European, Dravidian, Austroasiatic, and Sino-Tibetan.[38]

    While some connections have been tentatively proposed with other language families,[39] the consensus view is currently t

    The Jarawa originally inhabited southeastern Jarawa Island and have migrated to the west coast of Great Andaman in the wake of the Great Andamanese. The Onge once lived throughout Little Andaman and now are confined to two reservations on the island. The Jangil, who originally inhabited Rutland Island, were extinct by 1931; the last individual was sighted in 1907.[24] Only the Sentinelese are still living in their original homeland on North Sentinel Island, largely undisturbed, and have fiercely resisted all attempts at contact.

    The Andamanese languages are considered to be the fifth language family of India, following the Indo-European, Dravidian, Austroasiatic, and Sino-Tibetan.[38]

    While some connections have been tentatively proposed with other language families,[39] the consensus view is currently that Andamanese languages form a separate language family – or rather, two unrelated linguistic families: Greater Andamanese[40] and Ongan.

    [39] the consensus view is currently that Andamanese languages form a separate language family – or rather, two unrelated linguistic families: Greater Andamanese[40] and Ongan.

    Until contact, the Andamanese were strict hunter-gatherers. They did not practice cultivation, and lived off hunting indigenous pigs, fishing, and gathering. Their only weapons were the bow, adzes, and wooden harpoons. With the Aboriginal Tasmanians, the Andamanese were one of only two peoples who in the nineteenth century knew of no method for making fire.[41]:229 They instead carefully preserved embers[41]:229 in hollowed-out trees from fires caused by lightning strikes.

    The men wore girdles made of hibiscus fiber which carried useful tools and weapons for when they went hunting. The women on the other hand wore a tribal dress containing leaves that were held by a belt. A majority of them had painted bodies as well. They usually slept on leaves or mats and had either permanent or temporary habitation among the tribes. All habitations were man made.[42]

    Some of the tribe members were credited to having supernatural powers. They were called oko-pai-ad, which meant dreamer. They were thought to have an influence on the members of the tribe and would bring misfortune to those who did not believe in their abilities. Traditional knowledge practitioners were the ones who helped with healthcare. The medicine that was used to cure illnesses were herbal most of the time. Various types of medicinal plants were used by the islanders. 77 total traditional knowledge practitioners were identified and 132 medicinal plants were used. The members of the tribes found various ways to use leaves in their everyday lives including clothing, medicine, and to sleep on.[43]

    Anthropologist A.R. Radcliffe Brown argued that the Andamanese had no government and made decisions by group consensus.[44]

    Religion

    The native Andamanese religion and belief system is a form of Animism. [42]

    Some of the tribe members were credited to having supernatural powers. They were called oko-pai-ad, which meant dreamer. They were thought to have an influence on the members of the tribe and would bring misfortune to those who did not believe in their abilities. Traditional knowledge practitioners were the ones who helped with healthcare. The medicine that was used to cure illnesses were herbal most of the time. Various types of medicinal plants were used by the islanders. 77 total traditional knowledge practitioners were identified and 132 medicinal plants were used. The members of the tribes found various ways to use leaves in their everyday lives including clothing, medicine, and to sleep on.[43]

    Anthropologist A.R. Radcliffe Brown argued that the Andamanese had no government and made decisions by group consensus.[44]

    The native Andamanese religion and belief system is a form of Animism. Ancestor worship is an important element in the religious traditions of the Andaman islands.[45]

    Physical appearance

    Dental characteristics also group the Andamanese between Negrito and East-Asian samples.[46]

    When comparing dental morphology the focus is on overall size and tooth shape. To measure the size and shape, Penrose's size and shape statistic is used. To calculate tooth size, the sum of the tooth area is taken. Factor analysis is applied to tooth size to achieve tooth shape. Results have shown that the dental morphology of Andaman Islanders resembles that of South Asians, whereas Philippine Negrito groups more closely resemble southeast Asians in their dental morphology. Therefore, the dental morphology of the Andamanese indicates a retention of dental morphology from early South Asians in the early-mid Holocene. Overall the dental morphology of the Andamanese is found to be distant from that of modern East Asians. However it is noted that the tooth size of the Andamanese is similar to that of the Han Chinese and Japanese.[12]

    Genetics

    [46]

    When comparing dental morphology the focus is on overall size and tooth shape. To measure the size and shape, Penrose's size and shape statistic is used. To calculate tooth size, the sum of the tooth area is taken. Factor analysis is applied to tooth size to achieve tooth shape. Results have shown that the dental morphology

    When comparing dental morphology the focus is on overall size and tooth shape. To measure the size and shape, Penrose's size and shape statistic is used. To calculate tooth size, the sum of the tooth area is taken. Factor analysis is applied to tooth size to achieve tooth shape. Results have shown that the dental morphology of Andaman Islanders resembles that of South Asians, whereas Philippine Negrito groups more closely resemble southeast Asians in their dental morphology. Therefore, the dental morphology of the Andamanese indicates a retention of dental morphology from early South Asians in the early-mid Holocene. Overall the dental morphology of the Andamanese is found to be distant from that of modern East Asians. However it is noted that the tooth size of the Andamanese is similar to that of the Han Chinese and Japanese.[12]

    Genetic analysis, both of nuclear DNA[8][47] and mitochondrial DNA[48] provide information about the origins of the Andamanese. The Andamanese are most genetically similar to the Malaysian Negrito tribes.[49]

    Nuclear DNA

    The Andamanese show a very small genetic variation, which is indicative of populations that have experienced a population bottleneck and then developed in isolation for a long period.

    An allele has been discovered among the Jarawas that is found nowhere else in the world. Blood samples of 116 Jarawas were collected and tested for Duffy blood group and malarial parasite infectivity. Results showed a total absence of both Fya and Fyb antigens in two areas (Kadamtala and R.K Nallah) and low prevalence of both Fya antigen in another two areas (Jirkatang and Tirur). There was an absence of malarial parasite population bottleneck and then developed in isolation for a long period.

    An allele has been discovered among the Jarawas that is found nowhere else in the world. Blood samples of 116 Jarawas were collected and tested for Duffy blood group and malarial parasite infectivity. Results showed a total absence of both allele has been discovered among the Jarawas that is found nowhere else in the world. Blood samples of 116 Jarawas were collected and tested for Duffy blood group and malarial parasite infectivity. Results showed a total absence of both Fya and Fyb antigens in two areas (Kadamtala and R.K Nallah) and low prevalence of both Fya antigen in another two areas (Jirkatang and Tirur). There was an absence of malarial parasite Plasmodium vivax infection though Plasmodium falciparum infection was present in 27·59% of cases. A very high frequency of Fy (a–b–) in the Jarawa tribe from all the four jungle areas of Andaman Islands along with total absence of P. vivax infections suggests the selective advantage offered to Fy (a–b–) individuals against P. vivax infection.[50]

    Overall, the Andamanese showed closest relations with other Oceanic populations. The Nicobarese in contrast were observed to share close genetic relations with adjacent Indo-Mongoloid populations of Northeast India and Myanmar.[47] Bulbeck (2013) likewise noted that the Andamanese's nuclear DNA clusters with that of other Andamanese Islanders, as they carry a unique branch of Haplogroup D (D1a3) and maternal M (mtDNA).[12]

    The use of single-nucleotide polymorphism (SNP) shows that the genome of Andamanese people is closest to those of other Oceanic Negrito groups and distinct from South Asians and East Asians.[12] Analysis of mtDNA, which is inherited exclusively by maternal descent, confirms the above results. All Onge belong to M32 mtDNA, a subgroup of M which is unique to Onge people.[48]

    According to Reich et al. (2009). South Asian populations of the Indian subcontinent are composed, to significantly varying degrees, of mixtures of ancestry

    According to Reich et al. (2009). South Asian populations of the Indian subcontinent are composed, to significantly varying degrees, of mixtures of ancestry from: a group (known as "ASI" or "Ancestral South Indian") closest to but distinct from Andaman islanders, and populations from Western Eurasia (comprising a component termed "ANI" or "Ancestral North Indian").[51] Reich et al. speculate that the Andamanese split from mainland Asia 1,700 generations ago.[52] Andamanese are the only South Asian population in the study that lacked any Ancestral North Indian admixture.[52] According to Basu et al. (2016), the populations of the Andaman Islands archipelago form a distinct ancestry, which is "coancestral to Oceanic populations and not close to South Asians (India)." They conclude that the Andamanese, though they may be the closest existing group to the ancient "ASI" ancestral component in modern South Asians, have a distinct ancestry and are not closely related to other South Asians but are closer to Southeast Asian Negritos.[53]

    Moorjani et al. 2013 state that the "ASI" component in South Asians, though not closely related to any living group, is "related (distantly) to indigenous Andaman Islanders." Moorjani et al. also suggest possible gene flow into the Andamanese from a population related to the ASI. The study concluded that “almost all groups speaking Indo-European or Dravidian languages lie along a gradient of varying relatedness to West Eurasians in PCA (referred to as “Indian cline”)”.[46]

    A genetic analysis from Chaubey et al. 2015 found evidence of East Asian (Han Chinese-related) ancestry in Andamanese people. They estimated 32% East Asian ancestry in the Onge and 31% in the Great Andamanese, but suggest that this finding may in fact reflect the genetic affinity of the Andamanese to Melanesian and Southeast Asian Negrito populations rather than true East Asian admixture (stating that "The Han ancestry measured in Andaman Negrito is probably partially capturing both Melanesian and Malaysian Negrito ancestry"),[3] as a previous study by Chaubey et al. suggested "a deep common ancestry" between Andamanese, Melanesians and other Negrito groups (as well as South Asians), and an affinity between Southeast Asian Negritos and Melanesians with East Asians.[54]

    McColl et al. (2018) Analysed 26 ancient samples from Southeast Asia and Japan spanning from the late Neolithic to the Iron Age, along with an ancient Ikawazu Jōmon sample from southeast Honshu.The Jomon female skeleton which was analyzed shows typical Jōmon morphology.[55] However this Jōmon individual partially shares some ancestry with prehistoric Hoabinhians, which in turn also share some ancestry with the Onge, Jehai (Peninsular Malaysia) in mainland Southeast Asia along with Indian groups and Papua New Guineans, which represents possible gene flow from that group into the Jōmon population.[56] The Jōmon individual is best modeled as a mix of Hòabìnhian (La368) and East Asian ancestry while present-day East Asians can be modeled as a mixture of an Önge-like population and a population related to the Tiányuán individual.[57][58] However, there is still lack of ancient genome data to understand the peopling history of East Eurasians. It is required to analyze more ancient genome data, if there found appropriate skeletons, in order to fill the gap and to prove the speculation.[59]

    A study by Narasimhan et al. (2018) concludes that ANI and ASI were formed in the 2nd millennium BCE, and were preceded by a mixture of AASI (Ancient Ancestral South Indian, i.e. hunter-gatherers sharing a common root with the Andamanese); and Iranian agriculturalists who arrived in India ca. 4700–3000 BCE, and "must have reached the Indus Valley by the 4th millennium BCE".[60] Narasimhan et al. observe that samples from the Indus periphery population are always mixes of the same two proximal sources of AASI and Iranian agriculturalist-related ancestry; with "one of the Indus Periphery individuals having ~42% AASI ancestry and the other two individuals having ~14-18% AASI ancestry" (with the remainder of their ancestry being from the Iranian agriculturalist-related population). According Narasimhan the genetic makeup of the ASI population consisted of about 73% AASI and about 27% from Iran-related peoples.[60]

    Another genetic study by Yelmen et al. (2019) shows that the native South Asian genetic component is distinct from the Andamanese and thus that the Andamanese (Onge) are an imperfect and imprecise proxy for "ASI" ancestry in South Asians (there is difficulty detecting ASI ancestry in the North Indian Gujarati when the Andamanese Onge are used). Yemen et al. suggest that the South Indian tribal Paniya people would serve as a better proxy than the Andamanese (Onge) for the "native South Asian" component in modern South Asians.[61]

    Two more recent genetic studies (Shinde et al. 2019 and Narasimhan et al. 2019) on remains from the Indus Valley civilisation (of parts of Bronze Age northwest India and east Pakistan) found them to have a mixed ancestry: The samples analyzed by Shinde et al. had about 50-98% of their genome from Iranian hunter-gatherer peoples, which were also ancestral to Iranian farmers, and from 2-50% from South-east Asian hunter-gatherers sharing a common ancestry with the Andamanese, with the Iranian ancestry being on average predominant. The samples analyzed by Narasimhan et al. had 45–82% Iranian-related ancestry and 11–50% AASI (or Andamanese-related hunter-gatherer ancestry). The analysed samples of both studies have little to none of the "Steppe ancestry" component associated with later Indo-European migrations into India. The authors found that the respective amounts of those ancestries varied significantly between individuals, and concluded that more samples are needed to get the full picture of Indian population history. Further Narasimhan et al. 2019 notes that the high correlation between Dravidian and ASI ancestry may suggest that the Dravidians originated from the AASI component and are native to peninsular South Asia (south and east India).[62][63]

    The male Y-chromosome in humans is inherited exclusively through paternal descent. All sampled males of Onges (23/23) and Jarawas (4/4) belong to Paragroup D-M174*(D1a3).[64][65][66][67] However, male Great Andamanese do not appear to carry these clades. A low resolution study suggests that they belong to haplogroups K, L, O and P1 (P-M45).[68]

    A 2017 study by Mondal et al. finds that Andamanese D lineages diverged from Japanese D lineages lineage ~53000 years ago" and that Andamanese D lineages and those of the Riang (a Tibeto-Burmese population) have their nearest related lineages in East Asia, also finding that the D lineages of the Jarawa and Onge (both Andamanese groups) share a common ancestry more recently, within the last ~7000 years. They further suggest that: “This strongly suggests that haplogroup D does not indicate a separate ancestry for Andamanese populations. Rather, haplogroup D was part of the standing variation carried by the OOA expansion, and later lost from most of the populations except in Andaman and partially in Japan and Tibet”.[15]

    Mitochondrial DNA

    Analysis of Riang (a Tibeto-Burmese population) have their nearest related lineages in East Asia, also finding that the D lineages of the Jarawa and Onge (both Andamanese groups) share a common ancestry more recently, within the last ~7000 years. They further suggest that: “This strongly suggests that haplogroup D does not indicate a separate ancestry for Andamanese populations. Rather, haplogroup D was part of the standing variation carried by the OOA expansion, and later lost from most of the populations except in Andaman and partially in Japan and Tibet”.[15]

    Analysis of mtDNA, which is inherited exclusively by maternal descent, confirms the above results. The Onge and all the Adamanan Islanders belong strictly to the mitochondrial Haplogroup M. All Andamanese belong to M31 and M32 mtDNA, a subgroup of M which is unique to Andamanese people.[12][48] The analysis of 20 coding regions in 20 samples of ancient Andamanese people and 12 samples of modern Indian populations changed the topology of the two lineages in South Asians. The data received suggests an M31a lineage in South Asians and in East Asians.[69] Other mainland specific subgroups of M are distributed in Asia, where they represent 60% of all maternal lineages.[68][70][71] According to Endicott et al. (2002), this haplogroup originated with the earliest settlers of India during the coastal migration that may have brought the ancestors of the Andamanese to the Indian mainland, the Andaman Islands, and farther afield to Southeast Asia.[72]

    Archaic Admixture

    Unlike some Negrito populations of Southeast Asia, Andaman Islanders have not been found to have Denisovan ancestry.[73] However, they are estimated, like all other non-African populations, to possess approximately 1-2% Neanderthal ancestry.[74] A study suggests that all Asian and Australo-Papuan populations, including Andaman Islanders, also share between 2.6 and 3.4% of the genetic profile of a previously unknown hominin that was genetically roughly equidistant to Denisovans and Neanderthals.[75][74] However, a 2018 study by Skoglund et al. lead by the Harvard Reich lab team did not replicate that result and did not find evidence for admixture from an unknown hominin in the Andamanese or other Asians.[76] Wall et al. 2019 also did not find evidence of unknown archaic hominin admixture in Andamanese.[77]

    See also