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Chlorobi
The green sulfur bacteria are a phylum of obligately anaerobic photoautotrophic bacteria that metabolize sulfur. Green sulfur bacteria are nonmotile (except ''Chloroherpeton thalassium'', which may glide) and capable of anoxygenic photosynthesis. They live in anaerobic aquatic environments. In contrast to plants, green sulfur bacteria mainly use sulfide ions as electron donors. They are autotrophs that utilize the reverse tricarboxylic acid cycle to perform carbon fixation. They are also mixotrophs and reduce nitrogen. Characteristics Green sulfur bacteria are gram-negative rod or spherical shaped bacteria. Some types of green sulfur bacteria have gas vacuoles that allow for movement. They are photolithoautotrophs, and use light energy and reduced sulfur compounds as the electron source. Electron donors include H2, H2S, S. The major photosynthetic pigment in these bacteria is Bacteriochlorophylls ''c'' or ''d'' in green species and ''e'' in brown species, and is located in t ...
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Chlorobaculum Tepidum
''Chlorobaculum tepidum'', previously known as ''Chlorobium tepidum'', is an anaerobic, thermophilic green sulfur bacteria first isolated from New Zealand. Cells are gram-negative and non-motile rods of variable length. They contain bacteriochlorophyll c and chlorosomes. Genome structure ''Chlorobaculum tepidum'' contains a genome that contains 2.15 Mbp. There are a total of 2,337 genes (of these genes, there are 2,245 protein coding genes and 56 tRNA and rRNA coding genes). It synthesizes chlorophyll ''a'' and bacteriochlorophylls (BChls) ''a'' and ''c'' and is a model organism used to elucidate the biosynthesis of BChl ''c''. Several of its carotenoid metabolic pathways (including a novel lycopene cyclase) have similar counterparts in cyanobacteria. See also * List of bacterial orders * List of bacteria genera This article lists the genera of the bacteria. The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN) and Natio ...
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Chlorosome
A chlorosome is a photosynthetic antenna complex found in green sulfur bacteria (GSB) and some green filamentous anoxygenic phototrophs (FAP) ( Chloroflexaceae, Oscillochloridaceae; both members of Chloroflexia). They differ from other antenna complexes by their large size and lack of protein matrix supporting the photosynthetic pigments. Green sulfur bacteria are a group of organisms that generally live in extremely low-light environments, such as at depths of 100 metres in the Black Sea. The ability to capture light energy and rapidly deliver it to where it needs to go is essential to these bacteria, some of which see only a few photons of light per chlorophyll per day. To achieve this, the bacteria contain chlorosome structures, which contain up to 250,000 chlorophyll molecules. Chlorosomes are ellipsoidal bodies, in GSB their length varies from 100 to 200 nm, width of 50-100 nm and height of 15 – 30 nm, in FAP the chlorosomes are somewhat smaller. Structu ...
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Winogradsky Column
The Winogradsky column is a simple device for culturing a large diversity of microorganisms. Invented in the 1880s by Sergei Winogradsky, the device is a column of pond mud and water mixed with a carbon source such as newspaper (containing cellulose), blackened marshmallows or egg-shells (containing calcium carbonate), and a sulfur source such as gypsum (calcium sulfate) or egg yolk. Incubating the column in sunlight for months results in an Wikt:aerobic, aerobic/Hypoxia (environmental), anaerobic gradient as well as a sulfide gradient. These two gradients promote the growth of different microorganisms such as ''Clostridium'', ''Desulfovibrio'', ''Chlorobium'', ''Chromatium'', ''Rhodomicrobium'', and ''Beggiatoa'', as well as many other species of bacteria, cyanobacteria, and algae. The column provides numerous gradients, depending on additive nutrients, from which the variety of aforementioned organisms can grow. The aerobic water phase and anaerobic mud or soil phase are one ...
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Carbon Fixation
Biological carbon fixation or сarbon assimilation is the process by which inorganic carbon (particularly in the form of carbon dioxide) is converted to organic compounds by living organisms. The compounds are then used to store energy and as structure for other biomolecules. Carbon is primarily fixed through photosynthesis, but some organisms use a process called chemosynthesis in the absence of sunlight. Organisms that grow by fixing carbon are called autotrophs, which include photoautotrophs (which use sunlight), and lithoautotrophs (which use inorganic oxidation). Heterotrophs are not themselves capable of carbon fixation but are able to grow by consuming the carbon fixed by autotrophs or other heterotrophs. "Fixed carbon", "reduced carbon", and "organic carbon" may all be used interchangeably to refer to various organic compounds. Chemosynthesis is carbon fixation driven by chemical energy, rather than from sunlight. Sulfur- and hydrogen-oxidizing bacteria often use the ...
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The All-Species Living Tree Project
The All-Species Living Tree' Project is a collaboration between various academic groups/institutes, such as ARB Project, ARB, SILVA rRNA database project, and List of Prokaryotic names with Standing in Nomenclature, LPSN, with the aim of assembling a database of 16S ribosomal RNA, 16S rRNA sequences of all validly published species of ''Bacteria'' and ''Archaea''. At one stage, 23S ribosomal RNA, 23S sequences were also collected, but this has since stopped. Currently there are over 10,950 species in the aligned dataset and several more are being added either as new species are discovered or species that are not represented in the database are sequenced. Initially the latter group consisted of 7% of species. Similar (and more recent) projects include the Genomic Encyclopedia of Bacteria and Archaea (GEBA), which focused on whole genome sequencing of bacteria and archaea. Tree The tree was created by maximum likelihood analysis without bootstrap: consequently accuracy is traded ...
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Gram-negative Bacteria
Gram-negative bacteria are bacteria that do not retain the crystal violet stain used in the Gram staining method of bacterial differentiation. They are characterized by their cell envelopes, which are composed of a thin peptidoglycan cell wall sandwiched between an inner cytoplasmic cell membrane and a bacterial outer membrane. Gram-negative bacteria are found in virtually all environments on Earth that support life. The gram-negative bacteria include the model organism ''Escherichia coli'', as well as many pathogenic bacteria, such as ''Pseudomonas aeruginosa'', '' Chlamydia trachomatis'', and ''Yersinia pestis''. They are a significant medical challenge as their outer membrane protects them from many antibiotics (including penicillin), detergents that would normally damage the inner cell membrane, and lysozyme, an antimicrobial enzyme produced by animals that forms part of the innate immune system. Additionally, the outer leaflet of this membrane comprises a complex lipopol ...
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Black Smoker
A hydrothermal vent is a fissure on the seabed from which geothermally heated water discharges. They are commonly found near volcanically active places, areas where tectonic plates are moving apart at mid-ocean ridges, ocean basins, and hotspots. Hydrothermal deposits are rocks and mineral ore deposits formed by the action of hydrothermal vents. Hydrothermal vents exist because the earth is both geologically active and has large amounts of water on its surface and within its crust. Under the sea, they may form features called black smokers or white smokers. Relative to the majority of the deep sea, the areas around hydrothermal vents are biologically more productive, often hosting complex communities fueled by the chemicals dissolved in the vent fluids. Chemosynthetic bacteria and Archaea form the base of the food chain, supporting diverse organisms, including giant tube worms, clams, limpets and shrimp. Active hydrothermal vents are thought to exist on Jupiter's moon Europa an ...
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Bacteria
Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among the first life forms to appear on Earth, and are present in most of its habitats. Bacteria inhabit soil, water, acidic hot springs, radioactive waste, and the deep biosphere of Earth's crust. Bacteria are vital in many stages of the nutrient cycle by recycling nutrients such as the fixation of nitrogen from the atmosphere. The nutrient cycle includes the decomposition of dead bodies; bacteria are responsible for the putrefaction stage in this process. In the biological communities surrounding hydrothermal vents and cold seeps, extremophile bacteria provide the nutrients needed to sustain life by converting dissolved compounds, such as hydrogen sulphide and methane, to energy. Bacteria also live in symbiotic and parasitic relationsh ...
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Photoautotrophic
Photoautotrophs are organisms that use light energy and inorganic carbon to produce organic materials. Eukaryotic photoautotrophs absorb energy through the chlorophyll molecules in their chloroplasts while prokaryotic photoautotrophs use chlorophylls and bacteriochlorophylls present in their cytoplasm. All known photoautotrophs perform photosynthesis. Examples include plants, algae, and cyanobacteria. Origin and the Great Oxidation Event Chemical and geological evidence indicate that photosynthetic cyanobacteria existed about 2.6 billion years ago and anoxygenic photosynthesis had been taking place since a billion years before that. Oxygenic photosynthesis was the primary source of oxygenation and led to the Great Oxidation Event (the Oxygen Catastrophe) roughly 2.4 to 2.1 billion years ago. Although the end of the Great Oxidation Event was marked by a significant decrease in gross primary productivity that eclipsed extinction events, the development of aerobic respiration enabled ...
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Phylum
In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about 8 phyla. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguished them as a group ("a self-contained unity" ...
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