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Apoikozoa
Choanozoa is a clade of opisthokont eukaryotes consisting of the choanoflagellates (Choanoflagellatea) and the animals (Animalia, Metazoa). The sister-group relationship between the choanoflagellates and animals has important implications for the origin of the animals. The clade was identified in 2015 by Graham Budd and Sören Jensen, who used the name Apoikozoa. The 2018 revision of the classification first proposed by the International Society of Protistologists in 2012 recommends the use of the name Choanozoa. Introduction A close relationship between choanoflagellates and animals has long been recognised, dating back at least to the 1840s. A particularly striking and famous similarity between the single-celled choanoflagellates and multicellular animals is provided by the collar cells of sponges and the overall morphology of the choanoflagellate cell. The relationship has since been confirmed by multiple molecular analyses. This proposed homology was however thrown into ...
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Codosiga
''Codonosiga'' is a genus of choanoflagellate in the family Codonosigidae Codonosigidae were a Choanoflagellate The choanoflagellates are a group of free-living unicellular and colonial flagellate eukaryotes considered to be the closest living relatives of the animals. Choanoflagellates are collared flagellates, hav .... It is currently considered as a synonym of Codosiga, with 29 species in the family. References Footnotes * James-Clark, H. (1868). On the Spongia Ciliatae as Infusoria Flagellata; or, observations on the structure, animality, and relationship of Leucosolenia botryoides, Bowerbank. Memoirs of the Boston Society of Natural History, 1 (3): 305–340. Also published in the Proceedings of this Society on June 20, 1866 (vol. 11, p. 15), in the ''American Journal of Science in November'' 1866, in the Annals and Magazine of Natural History in January 1867, and in the Annals and Magazine of Natural History in 1868 (4th ser., vol. 1: 133–142, 188–215, 25 ...
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Heterotrophs
A heterotroph (; ) is an organism that cannot produce its own food, instead taking nutrition from other sources of organic carbon, mainly plant or animal matter. In the food chain, heterotrophs are primary, secondary and tertiary consumers, but not producers. Living organisms that are heterotrophic include all animals and fungi, some bacteria and protists, and many parasitic plants. The term heterotroph arose in microbiology in 1946 as part of a classification of microorganisms based on their type of nutrition. The term is now used in many fields, such as ecology in describing the food chain. Heterotrophs may be subdivided according to their energy source. If the heterotroph uses chemical energy, it is a chemoheterotroph (e.g., humans and mushrooms). If it uses light for energy, then it is a photoheterotroph (e.g., green non-sulfur bacteria). Heterotrophs represent one of the two mechanisms of nutrition (trophic levels), the other being autotrophs (''auto'' = self, ''troph'' ...
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Opisthosporidia
Opisthosporidia is a superphylum of intracellular parasites with amoeboid vegetative stage, defined as a common group of eukaryotic groups Microsporidia, Cryptomycota (also known as Rozellida, Rozellomycota, or Rozellosporidia) and Aphelidea. They have been considered to represent a monophyletic lineage with shared ecological and structural features, being a sister clade of the Fungi. Together with the Fungi they represent a sister clade of the Cristidiscoidea, together forming the Holomycota. Several other basal groups of the freshwater, marine and soil-inhabiting Holomycota were identified in recent studies, as the 'basal clone group 1' (BCG1=NCLC1), 'basal clone group 2' (BCG2), 'basal marine group' (NAMAKO-37), 'basal group GS01', the inner relationships of Opisthosporidia were clarified and their monophyly questioned: Cryptomycota and Microsporidia were proposed to join the phylum Rozellomycota, while Aphelidea were considered as a separate, though related phylum and all t ...
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Fungi
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota'' (''t ...
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Nuclearia Sp Nikko (cropped)
''Nuclearia'' is a nucleariid Nucleariida is a group of amoebae with filose pseudopods, known mostly from soils and freshwater. They are distinguished from the superficially similar vampyrellids mainly by having mitochondria with discoid cristae, in the absence of superfici ... genus. Species include: * Species ?'' Astrodisculus affinis'' Schouteden 1905 * Species ?'' Astrodisculus araneiformis'' Schewiakoff 1893 * Species ?'' Astrodisculus laciniatus'' Penard 1904 'Chlamydaster lacinatus'' (Penard 1904) Rainer 1968* Species ?'' Astrodisculus marinus'' Kufferath 1952 * Species ?'' Astrodisculus minutus'' Greeff 1869 * Species ?'' Heliophrys variabilis'' * Species ?'' Nuclearina similis'' * Species ?'' N. amphizonellae'' Penard 1917 * Species ?'' N. conspicua'' West 1903 * Species ?'' N. delicatula'' Cienkowsky 1865 * Species ?'' N. lohmanni'' Kufferath 1952 * Species ?'' N. pseudotenelloides'' * Species '' N. flavescens'' (Greef 1869) Patterson 1984 'Astrodisculus flavescen ...
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Nucleariid
Nucleariida is a group of amoebae with filose pseudopods, known mostly from soils and freshwater. They are distinguished from the superficially similar vampyrellids mainly by having mitochondria with discoid cristae, in the absence of superficial granules, and in the way they consume food. Classification Molecular studies indicate that nucleariids are closely related to fungi. and more distantly to the lineage that gave rise to choanoflagellates and metazoa opisthokonts, the group which includes animals, fungi. Some use a broad definition of Opisthokonta to include all of these organisms with flattened mitochondrial cristae. The genera ''Rabdiophrys'', ''Pinaciophora'', and ''Pompholyxophrys'', freshwater forms with hollow siliceous scales or spines, were included in Nucleariida by some. This was disputed by Smith and Chao who placed them in the Rhizaria. Their affinity with the nucleariids has been confirmed. Historically, nucleariids were included among the heliozoa as the ...
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Fonticulida
''Fonticula'' is a genus of cellular slime mold which forms a fruiting body in a volcano shape. As long ago as 1979 it has been known to not have a close relationship with either the Dictyosteliida or the Acrasidae, the two well-established groups of cellular slime molds. In 1979, ''Fonticula'' was made a new genus of its own due to the unique characteristics of its fruiting body, with only one species: ''Fonticula alba''. The life cycle of ''Fonticula alba'' alternates between an amoeboid vegetative stage and aggregative fruiting stage. The fruiting body of the genus has a unique shape, as its sorocarp resembles a volcano and sorus looks like a ball of hot lava emerging from that volcano. Molecular phylogenies have found alignments in genes of ''Fonticula alba'' to subgroups in Opisthokonta. A 2009 study has found that ''Fonticula'' is the sister taxa to ''Nuclearia'', thus making it related to the kingdom Fungi. ''Fonticula'', ''Nuclearia'', and Fungi have been united into th ...
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Cristidiscoidea
Cristidiscoidea or Nucleariae is a proposed basal holomycota clade in which Fonticula and Nucleariida emerged, as sister of the fungi. Since it is close to the divergence between the main lineages of fungi and animals, the study of Cristidiscoidea can provide crucial information on the divergent lifestyles of these groups and the evolution of opisthokonts and slime mold multicellularity. The holomycota tree is following Tedersoo et al. Classification * Class Cristidiscoidea Cavalier-Smith 1998 ** Order Fonticulida Cavalier-Smith 1993 *** Family Fonticulidae Worley, Raper & Hohl 1979 **** Genus ''Fonticula'' Worley, Raper & Hohl 1979 **** Genus '' Parvularia'' López-Escardó 2017 ***** Species '' P. atlantis'' López-Escardó 2017 ** Order Nucleariida Cavalier-Smith 1993 *** Family Nucleariidae Cann & Page 1979 **** Genus ''Nuclearia ''Nuclearia'' is a nucleariid Nucleariida is a group of amoebae with filose pseudopods, known mostly from soils and freshwater. They ar ...
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Holomycota
Holomycota or Nucletmycea are a basal Opisthokont clade as sister of the Holozoa. It consists of the Cristidiscoidea and the kingdom Fungi. The position of nucleariids, unicellular free-living phagotrophic amoebae, as the earliest lineage of Holomycota suggests that animals and fungi independently acquired complex multicellularity from a common unicellular ancestor and that the osmotrophic lifestyle (one of the fungal hallmarks) was originated later in the divergence of this eukaryotic lineage. Opisthosporidians is a recently proposed taxonomic group that includes aphelids, Microsporidia and Cryptomycota, three groups of endoparasites. ''Rozella'' ( Cryptomycota) is the earliest diverging fungal genus in which chitin has been observed at least in some stages of their life cycle, although the chitinous cell wall (another fungal hallmark) and osmotrophy originated in a common ancestor of Blastocladiomycota and Chytridiomycota, which still contain some ancestral characteristics suc ...
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Kimberella
''Kimberella'' is an extinct genus of bilaterian known only from rocks of the Ediacaran period. The slug-like organism fed by scratching the microbial surface on which it dwelt in a manner similar to the gastropods, although its affinity with this group is contentious. Specimens were first found in Australia's Ediacara Hills, but recent research has concentrated on the numerous finds near the White Sea in Russia, which cover an interval of time from . As with many fossils from this time, its evolutionary relationships to other organisms are hotly debated. Paleontologists initially classified ''Kimberella'' as a type of Cubozoan, but, since 1997, features of its anatomy and its association with scratch marks resembling those made by a radula have been interpreted as signs that it may have been a mollusc. Although some paleontologists dispute its classification as a mollusc, it is generally accepted as being at least a bilaterian. The classification of ''Kimberella'' is imp ...
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Ediacaran Biota
The Ediacaran (; formerly Vendian) biota is a Taxonomy (biology), taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period (). These were composed of enigmatic tubular and frond-shaped, mostly sessility (zoology), sessile, organisms. Trace fossils of these organisms have been found worldwide, and represent the earliest known complex multicellular organisms. The Ediacaran biota may have undergone evolutionary radiation in a proposed event called the Avalon explosion, . This was after the Earth had thawed from the Cryogenian period's Snowball Earth, extensive glaciation. This biota largely disappeared with the rapid increase in biodiversity known as the Cambrian explosion. Most of the currently existing body plans of animals first appeared in the fossil, fossil record of the Cambrian rather than the Ediacaran. For macroorganisms, the Cambrian biota appears to have almost completely replaced the organisms that dominate ...
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Ediacaran Period
The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and the beginning of the Phanerozoic Eon. It is named after the Ediacara Hills of South Australia. The Ediacaran Period's status as an official geological period was ratified in 2004 by the International Union of Geological Sciences (IUGS), making it the first new geological period declared in 120 years. Although the period takes its name from the Ediacara Hills where geologist Reg Sprigg first discovered fossils of the eponymous Ediacaran biota in 1946, the type section is located in the bed of the Enorama Creek within Brachina Gorge in the Flinders Ranges of South Australia, at . The Ediacaran marks the first appearance of widespread multicellular fauna following the end of Snowball Earth glaciation events, the so-called Ediacaran biota, wh ...
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