HOME

TheInfoList




A seed is an
embryonic ''Embryonic'' is the twelfth studio album by experimental rock band the Flaming Lips released on October 13, 2009, on Warner Bros. Records, Warner Bros. The band's first double album, it was released to generally positive reviews. Production News ...
plant Plants are predominantly photosynthetic Photosynthesis is a process used by plants and other organisms to Energy transformation, convert light energy into chemical energy that, through cellular respiration, can later be released to fuel ...

plant
enclosed in a protective outer covering. The formation of the seed is part of the process of
reproduction Reproduction (or procreation or breeding) is the biological process Biological processes are those processes that are vital for an organism In biology, an organism (from Ancient Greek, Greek: ὀργανισμός, ''organismos'') is ...

reproduction
in seed plants, the
spermatophytes The spermatophytes (; ), also known as phanerogams (taxon Phanerogamae) or phaenogams (taxon Phaenogamae), comprise those plant Plants are predominantly photosynthetic eukaryotes of the Kingdom (biology), kingdom Plantae. Historically, the p ...
, including the
gymnosperm The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also ...
and
angiosperm Flowering plants include multiple members of the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek#REDIRECT Greek Greek may refer to: Greece Anything of, from, or related to Greece Greece ...

angiosperm
plants. Seeds are the product of the ripened
ovule In seed plant The spermatophytes, also known as phanerogams (taxon Phanerogamae) or phaenogams (taxon Phaenogamae), comprise those plants that produce seeds, hence the alternative name seed plants. They are a subset of the embryophytes or l ...

ovule
, after
fertilization Fertilisation or fertilization (see American and British English spelling differences#-ise.2C -ize .28-isation.2C -ization.29, spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes ...

fertilization
by
pollen Pollen is a powdery substance consisting of pollen grains which are Sporophyte, microsporophytes of spermatophyta, seed plants, which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the ga ...

pollen
and some growth within the mother plant. The
embryo An embryo is the early stage of development of a multicellular organism A multicellular organism is an organism that consists of more than one cell (biology), cell, in contrast to a unicellular organism. All species of animals, Embryophyte, la ...

embryo
develops from the
zygote A zygote (, ) is a eukaryotic Eukaryotes () are organism In biology, an organism () is any organic, life, living system that functions as an individual entity. All organisms are composed of cells (cell theory). Organisms are c ...

zygote
, and the seed coat from the
integuments
integuments
of the ovule. Seeds have been an important development in the reproduction and success of gymnosperm and angiosperm plants, relative to more primitive plants such as
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plant Vascular plants (from Latin ''vasculum'': duct), also known as Tracheophyta (the tracheophytes , from Greek τραχεῖα ἀρτηρία ''trācheia art ...

fern
s,
moss Mosses are small, non-vascular plant, non-vascular flowerless plants in the taxonomic phylum, division Bryophyta (, ) ''sensu stricto''. Bryophyta (''sensu lato'', Wilhelm Philippe Schimper, Schimp. 1879) may also refer to the parent group bryo ...

moss
es and
liverworts The Marchantiophyta () are a division of non-vascular land plant The Embryophyta (), or land plants, are the most familiar group of green plant Plants are predominantly photosynthetic eukaryotes of the Kingdom (biology), kingdom Plantae ...
, which do not have seeds and use water-dependent means to propagate themselves. Seed plants now dominate biological
niches
niches
on land, from
forest A forest is an area of land dominated by tree In botany, a tree is a perennial plant with an elongated Plant stem, stem, or trunk (botany), trunk, supporting branches and leaves in most species. In some usages, the definition of a ...

forest
s to
grassland Grasslands are areas where the vegetation Vegetation is an assemblage of plant Plants are predominantly photosynthetic eukaryotes of the Kingdom (biology), kingdom Plantae. Historically, the plant kingdom encompassed all living things t ...

grassland
s both in hot and cold
climates Climate is the long-term average of weather, typically averaged over a period of 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologic ...

climates
. The term "seed" also has a general meaning that antedates the above – anything that can be
sown Sowing is the process of planting. An area or object that has had seeds planted in it will be described as a sowed area. Plants which are usually sown Among the major field crops, oat The oat (''Avena sativa''), sometimes called the ...

sown
, e.g. "seed"
potato The potato is a starch Starch or amylum is a polymeric A polymer (; Greek '' poly-'', "many" + '' -mer'', "part") is a substance Substance may refer to: * Substance (Jainism), a term in Jain ontology to denote the base or owner of attr ...

potato
es, "seeds" of
corn Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American North America is a continent entirely within the Northern Hemisphere and almost all within the Western Hemisphere. It can ...

corn
or sunflower "seeds". In the case of
sunflower ''Helianthus'' () is a genus Genus /ˈdʒiː.nəs/ (plural genera /ˈdʒen.ər.ə/) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, vi ...

sunflower
and corn "seeds", what is sown is the seed enclosed in a shell or
husk Husk (or hull) in botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" come ...
, whereas the potato is a
tuber Tubers are enlarged structures used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing season, and ...
. In the angiosperms (flowering plants), the ovary ripens to a
fruit In botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the ...

fruit
which contains the seed and serves to disseminate it. Many structures commonly referred to as "seeds" are actually dry fruits. Sunflower seeds are sometimes sold commercially while still enclosed within the hard wall of the fruit, which must be split open to reach the seed. Different groups of plants have other modifications, the so-called
stone fruits ), showing both fruit and seed A seed is an embryonic plant enclosed in a protective outer covering. The formation of the seed is part of the process of reproduction Reproduction (or procreation or breeding) is the biological process ...
(such as the
peach The peach (''Prunus persica'') is a deciduous In the fields of horticulture Horticulture is the art of cultivating plants in gardens to produce food and medicinal ingredients, or for comfort and ornamental purposes. Horticulturists grow ...

peach
) have a hardened fruit layer (the endocarp) fused to and surrounding the actual seed.
Nuts Nut often refers to: * Nut (fruit) A nut is a fruit composed of an inedible hard shell and a seed, which is generally edible. In general usage and in a Culinary nut, culinary sense, a wide variety of dried seeds are called nuts, but in a botan ...
are the one-seeded, hard-shelled fruit of some plants with an
indehiscent 200px, Dehiscence of the follicular fruit of milkweed ('' Asclepias syriaca'') revealing seeds within Dehiscence is the splitting of a mature plant structure along a built-in line of weakness in order to release its contents. This is common among ...
seed, such as an
acorn The acorn, or oaknut, is the nut Nut often refers to: * Nut (fruit), a fruit composed of a hard shell and a seed * Nut (food), collective noun for dry and edible fruits or seeds * Nut (hardware), a fastener used with a bolt Nut or Nuts may al ...

acorn
or
hazelnut The hazelnut is the nut (fruit), fruit of the hazel, hazel tree and therefore includes any of the nuts deriving from species of the genus ''Corylus'', especially the nuts of the species ''Corylus avellana''. They are also known as cobnuts or ...

hazelnut
.


History

Around 450 million years ago, it is believed that plants evolved from the oceans to dry land. According to fossil records, hair-like algae that formed on the edges of water evolved to survive
drought A drought is an event of prolonged shortages in the water supply, whether atmospheric (below-average precipitation In meteorology Meteorology is a branch of the atmospheric sciences Atmospheric science is the study of the Earth's atm ...

drought
. By 360 million years ago this evolution led to the development of a zygote that protected from the environment but nurished by the mother plant.


Seed production

Seeds are produced in several related groups of plants, and their manner of production distinguishes the
angiosperm Flowering plants include multiple members of the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek#REDIRECT Greek Greek may refer to: Greece Anything of, from, or related to Greece Greece ...

angiosperm
s ("enclosed seeds") from the
gymnosperm The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also ...
s ("naked seeds"). Angiosperm seeds are produced in a hard or fleshy structure called a
fruit In botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the ...

fruit
that encloses the seeds for protection in order to secure healthy growth. Some fruits have layers of both hard and fleshy material. In gymnosperms, no special structure develops to enclose the seeds, which begin their development "naked" on the bracts of cones. However, the seeds do become covered by the
cone A cone is a three-dimensional Three-dimensional space (also: 3-space or, rarely, tri-dimensional space) is a geometric setting in which three values (called parameter A parameter (from the Ancient Greek language, Ancient Greek wikt:πα ...
scales as they develop in some species of
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ex ...
. Seed production in natural plant populations varies widely from year to year in response to weather variables, insects and diseases, and internal cycles within the plants themselves. Over a 20-year period, for example, forests composed of
loblolly pine ''Pinus taeda'', commonly known as loblolly pine, is one of several pine A pine is any Pinophyta, conifer in the genus ''Pinus'' () of the family (biology), family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The Plant List ...

loblolly pine
and
shortleaf pine ''Pinus echinata'', the shortleaf pine, is a species of pine A pine is any Pinophyta, conifer in the genus ''Pinus'' () of the family (biology), family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The Plant List compiled by ...

shortleaf pine
produced from 0 to nearly 5 million sound pine seeds per hectare. Over this period, there were six bumper, five poor, and nine good seed crops, when evaluated for production of adequate
seedling A seedling is a young sporophyte '' Tortula muralis''. In mosses, the gametophyte is the dominant generation, while the sporophytes consist of sporangium-bearing stalks growing from the tips of the gametophytes A sporophyte () is the dip ...

seedling
s for natural forest reproduction.


Development

Angiosperm Flowering plants include multiple members of the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek#REDIRECT Greek Greek may refer to: Greece Anything of, from, or related to Greece Greece ...

Angiosperm
(flowering plants) seeds consist of three genetically distinct constituents: (1) the embryo formed from the zygote, (2) the endosperm, which is normally triploid, (3) the seed coat from tissue derived from the maternal tissue of the ovule. In angiosperms, the process of seed development begins with
double fertilization Double fertilization is a complex fertilization Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual orga ...
, which involves the fusion of two male gametes with the egg cell and the central cell to form the primary
endosperm 350px, right The endosperm is a tissue produced inside the seed A seed is an embryonic ''Embryonic'' is the twelfth studio album by experimental rock band the Flaming Lips released on October 13, 2009, on Warner Bros. Records, Warner Bros. The ...
and the zygote. Right after fertilization, the zygote is mostly inactive, but the primary endosperm divides rapidly to form the endosperm tissue. This tissue becomes the food the young plant will consume until the roots have developed after
germination seedlings, three days after germination Germination is the process by which an organism In biology, an organism () is any organic, life, living system that functions as an individual entity. All organisms are composed of cells (cell t ...

germination
.


Ovule

After fertilization, the
ovules In seed plant The spermatophytes, also known as phanerogams (taxon Phanerogamae) or phaenogams (taxon Phaenogamae), comprise those plants that produce seeds, hence the alternative name seed plants. They are a subset of the embryophytes or l ...
develop into the seeds. The ovule consists of a number of components: * The funicle (''funiculus, funiculi'') or seed stalk which attaches the ovule to the
placenta The placenta is a temporary fetal that begins developing from the shortly after . It plays critical roles in facilitating nutrient, gas and waste exchange between the physically separate maternal and fetal circulations, and is an important pro ...

placenta
and hence
ovary The ovary is an organ found in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the b ...
or fruit wall, at the pericarp. * The nucellus, the remnant of the
megasporangium '' mold asexual sporophyte generation) leaf Image:Equisetum arvense sporangia.jpg, 200px, ''Equisetum arvense'' Greek ''σπόρος ()'' ‘spore’ + ''ἀγγεῖον ()'' ‘vessel’) is an enclosure in which spores )'', growing on ...
and main region of the ovule where the megagametophyte develops. * The micropyle, a small pore or opening in the apex of the integument of the ovule where the pollen tube usually enters during the process of fertilization. * The
chalaza The chalaza (; from Greek "hailstone"; plural ''chalazas'' or ''chalazae'', ) is a structure inside bird and reptile eggs and plant ovules. It attaches or suspends the yolk or nucellus within the larger structure. In animals In the eggs of ...
, the base of the ovule opposite the micropyle, where integument and nucellus are joined together. The shape of the ovules as they develop often affects the final shape of the seeds. Plants generally produce ovules of four shapes: the most common shape is called anatropous, with a curved shape. Orthotropous ovules are straight with all the parts of the ovule lined up in a long row producing an uncurved seed. Campylotropous ovules have a curved megagametophyte often giving the seed a tight "C" shape. The last ovule shape is called amphitropous, where the ovule is partly inverted and turned back 90 degrees on its stalk (the funicle or funiculus). In the majority of flowering plants, the zygote's first division is transversely oriented in regards to the long axis, and this establishes the polarity of the embryo. The upper or chalazal pole becomes the main area of growth of the embryo, while the lower or micropylar pole produces the stalk-like suspensor that attaches to the micropyle. The suspensor absorbs and manufactures nutrients from the endosperm that are used during the embryo's growth.


Embryo

The main components of the embryo are: * The cotyledons, the seed leaves, attached to the embryonic axis. There may be one (
Monocotyledons Monocotyledons (), commonly referred to as monocots, ( Lilianae '' sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds A seed is an Plant embryogenesis, embryonic plant enclosed in a testa (botany), pr ...
), or two (
Dicotyledons 200px, Young castor oil plant showing its prominent two embryonic leaves (cotyledons), that differ from the adult leaves. The dicotyledons, also known as dicots (or more rarely dicotyls), are one of the two groups into which all the flowering ...
). The cotyledons are also the source of nutrients in the non-endospermic dicotyledons, in which case they replace the endosperm, and are thick and leathery. In endospermic seeds, the cotyledons are thin and papery. Dicotyledons have the point of attachment opposite one another on the axis. * The
epicotylAn epicotyl is important for the beginning stages of a plant's life. It is the region of a seedling stem above the stalks of the seed leaves of an embryo plant. It grows rapidly, showing hypogeal germination, and extends the stem above the soil surfa ...
, the embryonic axis above the point of attachment of the cotyledon(s). * The plumule, the tip of the epicotyl, and has a feathery appearance due to the presence of young leaf primordia at the apex, and will become the shoot upon germination. * The
hypocotyl Diagram of Scouler's willow (''Salix scouleriana'') seed, indicating position of hypocotyl. The hypocotyl (short for "hypocotyledonous stem", meaning "below seed leaf") is the stem of a germinating seedling A seedling is a young sporophyte ...
, the embryonic axis below the point of attachment of the cotyledon(s), connecting the epicotyl and the radicle, being the stem-root transition zone. * The
radicle Image:Salix scouleriana.seed.jpg, 250px, seed of Scouler's willow (''Salix scouleriana'') In botany, the radicle is the first part of a seedling (a growing plant embryo) to emerge from the seed during the process of germination. The radicle is the ...
, the basal tip of the hypocotyl, grows into the primary root. Monocotyledonous plants have two additional structures in the form of sheaths. The plumule is covered with a
coleoptile Schematic image of wheat coleoptile (above) and flag leaf (below). Coleoptile is the pointed protective sheath covering the emerging shoot in monocotyledons such as grasses in which few leaf primordia and shoot apex of monocot embryo remain enclo ...

coleoptile
that forms the first leaf while the radicle is covered with a
coleorhiza
coleorhiza
that connects to the primary root and
adventitiousImportant structures in plant development are buds, shoot In botany, shoots consist of plant stem, stems including their appendages, the leaves and lateral buds, flowering stems and flower buds. The new growth from seed germination that grows upw ...
roots form the sides. Here the hypocotyl is a rudimentary axis between radicle and plumule. The seeds of corn are constructed with these structures; pericarp, scutellum (single large cotyledon) that absorbs nutrients from the endosperm, plumule, radicle, coleoptile, and coleorhiza – these last two structures are sheath-like and enclose the plumule and radicle, acting as a protective covering.


Seed coat

The maturing
ovule In seed plant The spermatophytes, also known as phanerogams (taxon Phanerogamae) or phaenogams (taxon Phaenogamae), comprise those plants that produce seeds, hence the alternative name seed plants. They are a subset of the embryophytes or l ...

ovule
undergoes marked changes in the integuments, generally a reduction and disorganization but occasionally a thickening. The seed coat forms from the two integuments or outer layers of cells of the ovule, which derive from tissue from the mother plant, the inner integument forms the tegmen and the outer forms the testa. (The seed coats of some monocotyledon plants, such as the grasses, are not distinct structures, but are fused with the fruit wall to form a
pericarp Fruit anatomy is the plant anatomy Plant anatomy or phytotomy is the general term for the study of the internal structure A structure is an arrangement and organization of interrelated elements in a material object or system A system is ...
.) The testae of both monocots and dicots are often marked with patterns and textured markings, or have wings or tufts of hair. When the seed coat forms from only one layer, it is also called the testa, though not all such testae are homologous from one species to the next. The funiculus abscisses (detaches at fixed point – abscission zone), the scar forming an oval depression, the hilum. Anatropous ovules have a portion of the funiculus that is adnate (fused to the seed coat), and which forms a longitudinal ridge, or raphe, just above the hilum. In bitegmic ovules (e.g. ''Gossypium'' described here) both inner and outer integuments contribute to the seed coat formation. With continuing maturation the cells enlarge in the outer integument. While the inner epidermis may remain a single layer, it may also divide to produce two to three layers and accumulates starch, and is referred to as the colourless layer. By contrast, the outer epidermis becomes tanniferous. The inner integument may consist of eight to fifteen layers. (Kozlowski 1972) As the cells enlarge, and starch is deposited in the outer layers of the pigmented zone below the outer epidermis, this zone begins to lignify, while the cells of the outer epidermis enlarge radially and their walls thicken, with nucleus and cytoplasm compressed into the outer layer. these cells which are broader on their inner surface are called palisade cells. In the inner epidermis, the cells also enlarge radially with plate like thickening of the walls. The mature inner integument has a palisade layer, a pigmented zone with 15–20 layers, while the innermost layer is known as the fringe layer. (Kozlowski 1972)


Gymnosperms

In gymnosperms, which do not form ovaries, the ovules and hence the seeds are exposed. This is the basis for their nomenclature – naked seeded plants. Two sperm cells transferred from the pollen do not develop the seed by double fertilization, but one sperm nucleus unites with the egg nucleus and the other sperm is not used. Sometimes each sperm fertilizes an egg cell and one zygote is then aborted or absorbed during early development. The seed is composed of the embryo (the result of fertilization) and tissue from the mother plant, which also form a cone around the seed in coniferous plants such as
pine A pine is any conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The divi ...

pine
and
spruce A spruce is a tree In botany, a tree is a perennial plant with an elongated Plant stem, stem, or trunk (botany), trunk, supporting branches and leaves in most species. In some usages, the definition of a tree may be narrower, including ...

spruce
.


Shape and appearance

A large number of terms are used to describe seed shapes, many of which are largely self-explanatory such as ''Bean-shaped'' (reniform) – resembling a kidney, with lobed ends on either side of the hilum, ''Square'' or ''Oblong'' – angular with all sides more or less equal or longer than wide, ''Triangular'' – three sided, broadest below middle, ''Elliptic'' or ''Ovate'' or ''Obovate'' – rounded at both ends, or egg shaped (ovate or obovate, broader at one end), being rounded but either symmetrical about the middle or broader below the middle or broader above the middle. Other less obvious terms include discoid (resembling a disc or plate, having both thickness and parallel faces and with a rounded margin), ellipsoid, globose (
spherical of a sphere A sphere (from Greek language, Greek —, "globe, ball") is a geometrical object in three-dimensional space Three-dimensional space (also: 3-space or, rarely, tri-dimensional space) is a geometric setting in which three values ...
), or subglobose (Inflated, but less than spherical), lenticular, oblong, ovoid, reniform and sectoroid. Striate seeds are striped with parallel, longitudinal lines or ridges. The commonest colours are brown and black, other colours are infrequent. The surface varies from highly polished to considerably roughened. The surface may have a variety of appendages (see Seed coat). A seed coat with the consistency of
cork Cork or CORK may refer to: Materials * Cork (material), an impermeable buoyant plant product ** Cork (plug), a cylindrical or conical object used to seal a container ***Wine cork Places Ireland * Cork (city) ** Metropolitan Cork, also known as G ...
is referred to as suberose. Other terms include crustaceous (hard, thin or brittle).


Structure

A typical seed includes two basic parts: # an
embryo An embryo is the early stage of development of a multicellular organism A multicellular organism is an organism that consists of more than one cell (biology), cell, in contrast to a unicellular organism. All species of animals, Embryophyte, la ...

embryo
; # a seed coat. In addition, the
endosperm 350px, right The endosperm is a tissue produced inside the seed A seed is an embryonic ''Embryonic'' is the twelfth studio album by experimental rock band the Flaming Lips released on October 13, 2009, on Warner Bros. Records, Warner Bros. The ...
forms a supply of nutrients for the embryo in most monocotyledons and the endospermic dicotyledons.


Seed types

Seeds have been considered to occur in many structurally different types (Martin 1946). These are based on a number of criteria, of which the dominant one is the embryo-to-seed size ratio. This reflects the degree to which the developing cotyledons absorb the nutrients of the endosperm, and thus obliterate it. Six types occur amongst the monocotyledons, ten in the dicotyledons, and two in the gymnosperms (linear and spatulate). This classification is based on three characteristics: embryo morphology, amount of endosperm and the position of the embryo relative to the endosperm.


Embryo

In endospermic seeds, there are two distinct regions inside the seed coat, an upper and larger endosperm and a lower smaller embryo. The
embryo An embryo is the early stage of development of a multicellular organism A multicellular organism is an organism that consists of more than one cell (biology), cell, in contrast to a unicellular organism. All species of animals, Embryophyte, la ...

embryo
is the fertilised ovule, an immature
plant Plants are predominantly photosynthetic Photosynthesis is a process used by plants and other organisms to Energy transformation, convert light energy into chemical energy that, through cellular respiration, can later be released to fuel ...

plant
from which a new plant will grow under proper conditions. The embryo has one
cotyledon A cotyledon (; ; ; , gen. (), ) is a significant part of the embryo within the seed of a plant, and is defined as "the embryonic leaf in seed-bearing plants, one or more of which are the first to appear from a germination, germinating see ...
or seed leaf in
monocotyledon Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one Embryo#Plant embryos, embryonic leaf, or cotyledon. Th ...
s, two cotyledons in almost all
dicotyledon The dicotyledons, also known as dicots (or more rarely dicotyls), are one of the two groups into which all the flowering plant The flowering plants, also known as Angiospermae (), or Magnoliophyta (), are the most diverse group of Embryophyte ...
s and two or more in gymnosperms. In the fruit of
grains A grain is a small, hard, dry – with or without an attached or layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are s and . After being harvested, dry ...

grains
(caryopses) the single monocotyledon is shield shaped and hence called a scutellum. The scutellum is pressed closely against the endosperm from which it absorbs food and passes it to the growing parts. Embryo descriptors include small, straight, bent, curved, and curled.


Nutrient storage

Within the seed, there usually is a store of
nutrient A nutrient is a substance Substance may refer to: * Substance (Jainism), a term in Jain ontology to denote the base or owner of attributes * Chemical substance, a material with a definite chemical composition * Matter, anything that has mass and t ...
s for the
seedling A seedling is a young sporophyte '' Tortula muralis''. In mosses, the gametophyte is the dominant generation, while the sporophytes consist of sporangium-bearing stalks growing from the tips of the gametophytes A sporophyte () is the dip ...

seedling
that will grow from the embryo. The form of the stored nutrition varies depending on the kind of plant. In angiosperms, the stored food begins as a tissue called the
endosperm 350px, right The endosperm is a tissue produced inside the seed A seed is an embryonic ''Embryonic'' is the twelfth studio album by experimental rock band the Flaming Lips released on October 13, 2009, on Warner Bros. Records, Warner Bros. The ...
, which is derived from the mother plant and the pollen via
double fertilization Double fertilization is a complex fertilization Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual orga ...
. It is usually
triploid Polyploidy is a condition in which the biological cell, cells of an organism have more than two paired (Homologous chromosome, homologous) sets of chromosomes. Most species whose cells have Cell nucleus, nuclei (eukaryotes) are diploid, meaning ...
, and is rich in
oil An oil is any nonpolar In chemistry Chemistry is the science, scientific study of the properties and behavior of matter. It is a natural science that covers the Chemical element, elements that make up matter to the chemical compound ...
or
starch Starch or amylum is a polymeric A polymer (; Greek '' poly-'', "many" + '' -mer'', "part") is a substance Substance may refer to: * Substance (Jainism), a term in Jain ontology to denote the base or owner of attributes * Chemical substance ...
, and
protein Proteins are large biomolecule , showing alpha helices, represented by ribbons. This poten was the first to have its suckture solved by X-ray crystallography by Max Perutz and Sir John Cowdery Kendrew in 1958, for which they received a No ...

protein
. In gymnosperms, such as
conifers Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All exta ...

conifers
, the food storage tissue (also called endosperm) is part of the female gametophyte, a
haploid Ploidy () is the number of complete sets of chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. Most eukaryotic chromosomes include packaging proteins called histones which, aided by ...
tissue. The endosperm is surrounded by the
aleurone Aleurone (from Greek#REDIRECT Greek Greek may refer to: Greece Anything of, from, or related to Greece Greece ( el, Ελλάδα, , ), officially the Hellenic Republic, is a country located in Southeast Europe. Its population is approximatel ...
layer (peripheral endosperm), filled with proteinaceous aleurone grains. Originally, by analogy with the animal
ovum The egg cell, or ovum (plural ova), is the female reproductive Reproduction (or procreation or breeding) is the biological process by which new individual organisms – "offspring" – are produced from their "parent" or parents. Reproduct ...

ovum
, the outer nucellus layer ( perisperm) was referred to as
albumen Egg white is the clear liquid (also called the albumen or the glair/glaire) contained within an egg Diagram of a chicken egg in its 9th day. Membranes: allantois, chorion, amnion, and vitellus/ yolk. An egg is the organic vessel containing ...
, and the inner endosperm layer as vitellus. Although misleading, the term began to be applied to all the nutrient matter. This terminology persists in referring to endospermic seeds as "albuminous". The nature of this material is used in both describing and classifying seeds, in addition to the embryo to endosperm size ratio. The endosperm may be considered to be farinaceous (or mealy) in which the cells are filled with
starch Starch or amylum is a polymeric A polymer (; Greek '' poly-'', "many" + '' -mer'', "part") is a substance Substance may refer to: * Substance (Jainism), a term in Jain ontology to denote the base or owner of attributes * Chemical substance ...
, as for instance cereal grains, or not (non-farinaceous). The endosperm may also be referred to as "fleshy" or "cartilaginous" with thicker soft cells such as
coconut The coconut tree (''Cocos nucifera'') is a member of the palm tree family (biology), family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, t ...

coconut
, but may also be oily as in ''
Ricinus ''Ricinus communis'', the castor bean or castor oil plant, is a species of perennial flowering plant in the Euphorbia, spurge family, Euphorbiaceae. It is the sole species in the monotypic genus, ''Ricinus'', and subtribe, Ricininae. The evolu ...

Ricinus
'' (castor oil), '' Croton'' and
Poppy in New Delhi, India A poppy is a flowering plant in the subfamily Papaveroideae of the family Papaveraceae. Poppies are herbaceous plants, often grown for their colourful flowers. One species of poppy, ''Papaver somniferum'', is the source of ...

Poppy
. The endosperm is called "horny" when the cell walls are thicker such as and
coffee Coffee is a drink prepared from roasted s, the seeds of from certain s in the ' genus. From the coffee fruit, the seeds are separated to produce a stable, raw product: unroasted ''green coffee''. The seeds are then , a process which transfo ...

coffee
, or "ruminated" if mottled, as in
nutmeg Nutmeg is the seed A seed is an embryonic ''Embryonic'' is the twelfth studio album by experimental rock band the Flaming Lips released on October 13, 2009, on Warner Bros. Records, Warner Bros. The band's first double album, it was relea ...

nutmeg
, palms and
Annonaceae The Annonaceae are a family In , family (from la, familia) is a of people related either by (by recognized birth) or (by marriage or other relationship). The purpose of families is to maintain the well-being of its members and of socie ...

Annonaceae
. In most monocotyledons (such as
grasses Poaceae () or Gramineae () is a large and nearly ubiquitous Family (biology), family of monocotyledonous flowering plants known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in ...
and ) and some (endospermic or albuminous) dicotyledons (such as
castor bean ''Ricinus communis'', the castor bean or castor oil plant, is a species of perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used ...
s) the embryo is embedded in the endosperm (and nucellus), which the seedling will use upon
germination seedlings, three days after germination Germination is the process by which an organism In biology, an organism () is any organic, life, living system that functions as an individual entity. All organisms are composed of cells (cell t ...

germination
. In the non-endospermic dicotyledons the endosperm is absorbed by the embryo as the latter grows within the developing seed, and the cotyledons of the embryo become filled with stored food. At maturity, seeds of these species have no endosperm and are also referred to as exalbuminous seeds. The exalbuminous seeds include the
legumes A legume () is a in the family (or Leguminosae), or the or of such a plant. When used as a dry , the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for and , and as soil-enhancing . Well-kno ...
(such as
bean A bean is the seed of one of several of the , which are used as vegetables for human or animal food. They can be cooked in many different ways, including boiling, frying, and baking, and are used in many traditional dishes throughout th ...

bean
s and
pea The pea is most commonly the small spherical seed A seed is an embryonic plant enclosed in a protective outer covering. The formation of the seed is part of the process of reproduction Reproduction (or procreation or breeding) is ...

pea
s), trees such as the
oak An oak is a tree In botany, a tree is a perennial plant with an elongated Plant stem, stem, or trunk (botany), trunk, supporting branches and leaves in most species. In some usages, the definition of a tree may be narrower, including on ...

oak
and
walnut A walnut is the Nut (fruit), nut of any tree of the genus ''Juglans'' (family Juglandaceae), particularly the Persian or English walnut, ''Juglans regia''. A walnut is the edible seed of a drupe, and thus not a true botanical nut. It is com ...

walnut
, vegetables such as
squash Squash may refer to: Sports * Squash (sport), the high-speed racquet sport also known as squash racquets * Squash (professional wrestling), an extremely one-sided match in professional wrestling * Squash tennis, a game similar to squash racquets ...
and
radish The radish (''Raphanus raphanistrum ''Raphanus raphanistrum'', the sea radish, wild radish, white charlock or jointed charlock, is a flowering plant in the family Brassicaceae. One of its subspecies, ''Raphanus raphanistrum'' subsp. ''sativ ...

radish
, and
sunflower ''Helianthus'' () is a genus Genus /ˈdʒiː.nəs/ (plural genera /ˈdʒen.ər.ə/) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, vi ...

sunflower
s. According to Bewley and Black (1978), Brazil nut storage is in hypocotyl, this place of storage is uncommon among seeds. All gymnosperm seeds are albuminous.


Seed coat

The seed coat develops from the maternal tissue, the
integument In biology, integument is the natural covering of an organism or an organ, such as its skin Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, an ...
s, originally surrounding the ovule. The seed coat in the mature seed can be a paper-thin layer (e.g.
peanut The peanut, also known as the groundnut, goober (US), pindar (US) or monkey nut (UK), and taxonomically classified as ''Arachis hypogaea'', is a legume A legume () is a plant Plants are predominantly photosynthetic Photosynth ...

peanut
) or something more substantial (e.g. thick and hard in
honey locust The honey locust (''Gleditsia triacanthos''), also known as the thorny locust or thorny honeylocust, is a deciduous In the fields of horticulture and botany, the term ''deciduous'' (; ) means "falling off at maturity" and "tending to fall of ...

honey locust
and
coconut The coconut tree (''Cocos nucifera'') is a member of the palm tree family (biology), family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, t ...

coconut
), or fleshy as in the
sarcotesta The sarcotesta is a fleshy seedcoat, a type of testa. Examples of seeds with a sarcotesta are pomegranate and some cycad seeds. The sarcotesta of pomegranate seeds consists of epidermal cells derived from the integument, and there are no arils ...
of
pomegranate The pomegranate (''Punica granatum'') is a fruit In botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who speci ...

pomegranate
. The seed coat helps protect the embryo from mechanical injury, predators, and drying out. Depending on its development, the seed coat is either bitegmic or unitegmic. Bitegmic seeds form a testa from the outer integument and a tegmen from the inner integument while unitegmic seeds have only one integument. Usually, parts of the testa or tegmen form a hard protective mechanical layer. The mechanical layer may prevent water penetration and germination. Amongst the barriers may be the presence of lignified sclereids. The outer integument has a number of layers, generally between four and eight organised into three layers: (a) outer epidermis, (b) outer pigmented zone of two to five layers containing
tannin Tannins (or tannoids) are a class of , ic s that bind to and s and various other organic compounds including s and s. The term ''tannin'' (from ''tanner'', from ''tannāre'', from ''tannum'', ) refers to the use of oak and other bark ...
and starch, and (c) inner epidermis. The endotegmen is derived from the inner epidermis of the inner integument, the exotegmen from the outer surface of the inner integument. The endotesta is derived from the inner epidermis of the outer integument, and the outer layer of the testa from the outer surface of the outer integument is referred to as the exotesta. If the exotesta is also the mechanical layer, this is called an exotestal seed, but if the mechanical layer is the endotegmen, then the seed is endotestal. The exotesta may consist of one or more rows of cells that are elongated and pallisade like (e.g.
Fabaceae The Fabaceae or Leguminosae,International Code of Nomenc ...

Fabaceae
), hence 'palisade exotesta'. In addition to the three basic seed parts, some seeds have an appendage, an
aril An aril (pronounced ), also called an arillus, is a specialized outgrowth from a seed that partly or completely covers the seed. An arillode or false aril is sometimes distinguished: whereas an aril grows from the attachment point of the seed ...

aril
, a fleshy outgrowth of the funicle (), (as in
yew Yew is a common name given to various species of trees. The name is most prominently given to any of various coniferous Conifers are a group of cone-bearing seed plants The spermatophytes (; ), also known as phanerogams (taxon Phanerogamae ...

yew
and
nutmeg Nutmeg is the seed A seed is an embryonic ''Embryonic'' is the twelfth studio album by experimental rock band the Flaming Lips released on October 13, 2009, on Warner Bros. Records, Warner Bros. The band's first double album, it was relea ...

nutmeg
) or an oily appendage, an
elaiosome '' seeds bearing orange elaiosomes Image:TrilliumRecurvatumSeeds.jpg">''Trillium recurvatum'' seeds Elaiosomes ( grc, ἔλαιον ''élaion'' "oil" + ''sóma'' "body") are fleshy structures that are attached to the seeds of many plant species. ...
(as in ''Corydalis''), or hairs (trichomes). In the latter example these hairs are the source of the textile crop cotton. Other seed appendages include the raphe (a ridge), wings, caruncles (a soft spongy outgrowth from the outer integument in the vicinity of the micropyle), spines, or tubercles. A scar also may remain on the seed coat, called the hilum, where the seed was attached to the ovary wall by the funicle. Just below it is a small pore, representing the micropyle of the ovule.


Size and seed set

Seeds are very diverse in size. The dust-like orchid seeds are the smallest, with about one million seeds per gram; they are often embryonic seeds with immature embryos and no significant energy reserves. Orchids and a few other groups of plants are myco-heterotrophy, mycoheterotrophs which depend on mycorrhizal fungi for nutrition during germination and the early growth of the seedling. Some terrestrial orchid seedlings, in fact, spend the first few years of their lives deriving energy from the fungi and do not produce green leaves. At over 20 kg, the largest seed is the ''coco de mer''. Plants that produce smaller seeds can generate many more seeds per flower, while plants with larger seeds invest more resources into those seeds and normally produce fewer seeds. Small seeds are quicker to ripen and can be dispersed sooner, so autumn all blooming plants often have small seeds. Many annual plants produce great quantities of smaller seeds; this helps to ensure at least a few will end in a favorable place for growth. Herbaceous perennials and woody plants often have larger seeds; they can produce seeds over many years, and larger seeds have more energy reserves for germination and seedling growth and produce larger, more established seedlings after germination.


Functions

Seeds serve several functions for the plants that produce them. Key among these functions are nourishment of the
embryo An embryo is the early stage of development of a multicellular organism A multicellular organism is an organism that consists of more than one cell (biology), cell, in contrast to a unicellular organism. All species of animals, Embryophyte, la ...

embryo
, biological dispersal, dispersal to a new location, and dormancy during unfavorable conditions. Seeds fundamentally are means of reproduction, and most seeds are the product of sexual reproduction which produces a remixing of genetic material and phenotype variability on which natural selection acts. Plant seeds hold endophyte, endophytic microorganisms that can perform various functions, the most important of which is protection against disease.


Embryo nourishment

Seeds protect and nourish the embryo or young plant. They usually give a seedling a faster start than a sporeling from a spore, because of the larger food reserves in the seed and the multicellularity of the enclosed embryo.


Dispersal

Unlike animals, plants are limited in their ability to seek out favorable conditions for life and growth. As a result, plants have evolved many ways to Biological dispersal, disperse their offspring by dispersing their seeds (see also vegetative reproduction). A seed must somehow "arrive" at a location and be there at a time favorable for germination and growth. When the fruits open and release their seeds in a regular way, it is called dehiscence (botany), dehiscent, which is often distinctive for related groups of plants; these fruits include Capsule (fruit), capsules, Follicle (fruit), follicles, Legume, legumes, Silicle, silicles and Silique, siliques. When fruits do not open and release their seeds in a regular fashion, they are called indehiscent, which include the fruits Achene, achenes, Caryopsis, caryopses, Nut (fruit), nuts, samara (fruit), samaras, and utricle (fruit), utricles.


By wind (anemochory)

* Some seeds (e.g.,
pine A pine is any conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The divi ...

pine
) have a wing that aids in wind dispersal. * The dustlike seeds of orchids are carried efficiently by the wind. * Some seeds (e.g. milkweed, Populus, poplar) have hairs that aid in wind dispersal. Other seeds are enclosed in fruit structures that aid wind dispersal in similar ways: * Dandelion achenes have hairs. * Maple samaras have two wings.


By water (hydrochory)

* Some plants, such as ''Mucuna'' and ''Dioclea (plant), Dioclea'', produce buoyant seeds termed sea-beans or drift seeds because they float in rivers to the oceans and wash up on beaches.


By animals (zoochory)

* Seeds (burr (fruit), burrs) with barbs or hooks (e.g. acaena, burdock, Rumex, dock) which attach to animal fur or feathers, and then drop off later. * Seeds with a fleshy covering (e.g. apple, cherry, juniper) are eaten by animals (birds, mammals, reptiles, fish) which then disperse these seeds in their faeces, droppings. * Seeds (nut (fruit), nuts) are attractive long-term storable food resources for animals (e.g.
acorn The acorn, or oaknut, is the nut Nut often refers to: * Nut (fruit), a fruit composed of a hard shell and a seed * Nut (food), collective noun for dry and edible fruits or seeds * Nut (hardware), a fastener used with a bolt Nut or Nuts may al ...

acorn
s, hazelnut,
walnut A walnut is the Nut (fruit), nut of any tree of the genus ''Juglans'' (family Juglandaceae), particularly the Persian or English walnut, ''Juglans regia''. A walnut is the edible seed of a drupe, and thus not a true botanical nut. It is com ...

walnut
); the seeds are stored some distance from the parent plant, and some escape being eaten if the animal forgets them. Myrmecochory is the dispersal of seeds by ants. Foraging ants disperse seeds which have appendages called
elaiosome '' seeds bearing orange elaiosomes Image:TrilliumRecurvatumSeeds.jpg">''Trillium recurvatum'' seeds Elaiosomes ( grc, ἔλαιον ''élaion'' "oil" + ''sóma'' "body") are fleshy structures that are attached to the seeds of many plant species. ...
s (e.g. bloodroot, trilliums, acacias, and many species of Proteaceae). Elaiosomes are soft, fleshy structures that contain nutrients for animals that eat them. The ants carry such seeds back to their nest, where the elaiosomes are eaten. The remainder of the seed, which is hard and inedible to the ants, then germinates either within the nest or at a removal site where the seed has been discarded by the ants.Ricklefs, Robert E. (1993) ''The Economy of Nature'', 3rd ed., p. 396. (New York: W.H. Freeman). . This dispersal relationship is an example of Mutualism (biology), mutualism, since the plants depend upon the ants to disperse seeds, while the ants depend upon the plants seeds for food. As a result, a drop in numbers of one partner can reduce success of the other. In South Africa, the Argentine ant (''Linepithema humile'') has invasive species, invaded and displaced native species of ants. Unlike the native ant species, Argentine ants do not collect the seeds of ''Mimetes cucullatus'' or eat the elaiosomes. In areas where these ants have invaded, the numbers of ''Mimetes'' seedlings have dropped.


Dormancy

Seed dormancy has two main functions: the first is synchronizing germination with the optimal conditions for survival of the resulting seedling; the second is spreading germination of a batch of seeds over time so a catastrophe (e.g. late frosts, drought, herbivory) does not result in the death of all offspring of a plant (Bet-hedging (biology), bet-hedging). Seed dormancy is defined as a seed failing to germinate under environmental conditions optimal for germination, normally when the environment is at a suitable temperature with proper soil moisture. This true dormancy or innate dormancy is therefore caused by conditions within the seed that prevent germination. Thus dormancy is a state of the seed, not of the environment. Induced dormancy, enforced dormancy or seed quiescence occurs when a seed fails to germinate because the external environmental conditions are inappropriate for germination, mostly in response to conditions being too dark or light, too cold or hot, or too dry. Seed dormancy is not the same as seed persistence in the soil or on the plant, though even in scientific publications dormancy and persistence are often confused or used as synonyms. Often, seed dormancy is divided into four major categories: exogenous; endogenous; combinational; and secondary. A more recent system distinguishes five classes: morphological, physiological, morphophysiological, physical, and combinational dormancy. Exogenous dormancy is caused by conditions outside the embryo, including: * Physical dormancy or hard seed coats occurs when seeds are Permeability (fluid), impermeable to water. At dormancy break, a specialized structure, the ‘water gap’, is disrupted in response to environmental cues, especially temperature, so water can enter the seed and germination can occur. Plant families where physical dormancy occurs include Anacardiaceae, Cannaceae, Convulvulaceae,
Fabaceae The Fabaceae or Leguminosae,International Code of Nomenc ...

Fabaceae
and Malvaceae. * Chemical dormancy considers species that lack physiological dormancy, but where a chemical prevents germination. This chemical can be leached out of the seed by rainwater or snow melt or be deactivated somehow. Leaching of chemical inhibitors from the seed by rain water is often cited as an important cause of dormancy release in seeds of desert plants, but little evidence exists to support this claim. Endogenous dormancy is caused by conditions within the embryo itself, including: * In morphological dormancy, germination is prevented due to morphological characteristics of the embryo. In some species, the embryo is just a mass of cells when seeds are dispersed; it is not differentiated. Before germination can take place, both differentiation and growth of the embryo have to occur. In other species, the embryo is differentiated but not fully grown (underdeveloped) at dispersal, and embryo growth up to a species specific length is required before germination can occur. Examples of plant families where morphological dormancy occurs are Apiaceae, Cycadaceae, Liliaceae, Magnoliaceae and Ranunculaceae.Baskin, C.C. and Baskin, J.M. (1998) Seeds: Ecology, biogeography, and evolution of dormancy and germination.San Diego, Academic Press.Baskin, J.M. and Baskin, C.C. (2004) A classification system for seed dormancy. Seed Science Research 14:1–16. * Morphophysiological dormancy includes seeds with underdeveloped embryos, and also have physiological components to dormancy. These seeds, therefore, require a dormancy-breaking treatments, as well as a period of time to develop fully grown embryos. Plant families where morphophysiological dormancy occurs include Apiaceae, Aquifoliaceae, Liliaceae, Magnoliaceae, Papaveraceae and Ranunculaceae. Some plants with morphophysiological dormancy, such as ''Asarum'' or ''Trillium'' species, have multiple types of dormancy, one affects radicle (root) growth, while the other affects plumule (shoot) growth. The terms "double dormancy" and "two-year seeds" are used for species whose seeds need two years to complete germination or at least two winters and one summer. Dormancy of the radicle (seedling root) is broken during the first winter after dispersal while dormancy of the shoot bud is broken during the second winter. * Physiological dormancy means the embryo, due to physiological causes, cannot generate enough power to break through the seed coat, endosperm or other covering structures. Dormancy is typically broken at cool wet, warm wet, or warm dry conditions. Abscisic acid is usually the growth inhibitor in seeds, and its production can be affected by light. **Drying, in some plants, including a number of grasses and those from seasonally arid regions, is needed before they will germinate. The seeds are released, but need to have a lower moisture content before germination can begin. If the seeds remain moist after dispersal, germination can be delayed for many months or even years. Many herbaceous plants from temperate climate zones have physiological dormancy that disappears with drying of the seeds. Other species will germinate after dispersal only under very narrow temperature ranges, but as the seeds dry, they are able to germinate over a wider temperature range. * In seeds with combinational dormancy, the seed or fruit coat is impermeable to water and the embryo has physiological dormancy. Depending on the species, physical dormancy can be broken before or after physiological dormancy is broken. * Secondary dormancy* is caused by conditions after the seed has been dispersed and occurs in some seeds when nondormant seed is exposed to conditions that are not favorable to germination, very often high temperatures. The mechanisms of secondary dormancy are not yet fully understood, but might involve the loss of sensitivity in receptors in the plasma membrane. The following types of seed dormancy do not involve seed dormancy, strictly speaking, as lack of germination is prevented by the environment, not by characteristics of the seed itself (see Germination): * Photodormancy or light sensitivity affects germination of some seeds. These photoblastic seeds need a period of darkness or light to germinate. In species with thin seed coats, light may be able to penetrate into the dormant embryo. The presence of light or the absence of light may trigger the germination process, inhibiting germination in some seeds buried too deeply or in others not buried in the soil. * Thermodormancy is seed sensitivity to heat or cold. Some seeds, including cocklebur and amaranth, germinate only at high temperatures (30 °C or 86 °F); many plants that have seeds that germinate in early to midsummer have thermodormancy, so germinate only when the soil temperature is warm. Other seeds need cool soils to germinate, while others, such as celery, are inhibited when soil temperatures are too warm. Often, thermodormancy requirements disappear as the seed ages or dries. Not all seeds undergo a period of dormancy. Seeds of some mangroves are viviparous; they begin to germinate while still attached to the parent. The large, heavy root allows the seed to penetrate into the ground when it falls. Many garden plant seeds will germinate readily as soon as they have water and are warm enough; though their wild ancestors may have had dormancy, these cultivated plants lack it. After many generations of selective pressure by plant breeders and gardeners, dormancy has been selected out. For annual plant, annuals, seeds are a way for the species to survive dry or cold seasons. Ephemeral plants are usually annuals that can go from seed to seed in as few as six weeks.


Persistence and seed banks


Germination

Seed germination is a process by which a seed embryo develops into a seedling. It involves the reactivation of the metabolic pathways that lead to growth and the emergence of the radicle or seed root and plumule or shoot. The emergence of the seedling above the soil surface is the next phase of the plant's growth and is called seedling establishment. Three fundamental conditions must exist before germination can occur. (1) The embryo must be alive, called seed viability. (2) Any dormancy requirements that prevent germination must be overcome. (3) The proper environmental conditions must exist for germination. Far red light can prevent germination. Seed viability is the ability of the embryo to germinate and is affected by a number of different conditions. Some plants do not produce seeds that have functional complete embryos, or the seed may have no embryo at all, often called empty seeds. Predators and pathogens can damage or kill the seed while it is still in the fruit or after it is dispersed. Environmental conditions like flooding or heat can kill the seed before or during germination. The age of the seed affects its health and germination ability: since the seed has a living embryo, over time cells die and cannot be replaced. Some seeds can live for a long time before germination, while others can only survive for a short period after dispersal before they die. Seed vigor is a measure of the quality of seed, and involves the viability of the seed, the germination percentage, germination rate, and the strength of the seedlings produced. The germination percentage is simply the proportion of seeds that germinate from all seeds subject to the right conditions for growth. The germination rate is the length of time it takes for the seeds to germinate. Germination percentages and rates are affected by seed viability, dormancy and environmental effects that impact on the seed and seedling. In agriculture and horticulture quality seeds have high viability, measured by germination percentage plus the rate of germination. This is given as a percent of germination over a certain amount of time, 90% germination in 20 days, for example. 'Dormancy' is covered above; many plants produce seeds with varying degrees of dormancy, and different seeds from the same fruit can have different degrees of dormancy. It's possible to have seeds with no dormancy if they are dispersed right away and do not dry (if the seeds dry they go into physiological dormancy). There is great variation amongst plants and a dormant seed is still a viable seed even though the germination rate might be very low. Environmental conditions affecting seed germination include; water, oxygen, temperature and light. Three distinct phases of seed germination occur: water imbibition; lag phase; and
radicle Image:Salix scouleriana.seed.jpg, 250px, seed of Scouler's willow (''Salix scouleriana'') In botany, the radicle is the first part of a seedling (a growing plant embryo) to emerge from the seed during the process of germination. The radicle is the ...
emergence. In order for the seed coat to split, the embryo must imbibe (soak up water), which causes it to swell, splitting the seed coat. However, the nature of the seed coat determines how rapidly water can penetrate and subsequently initiate
germination seedlings, three days after germination Germination is the process by which an organism In biology, an organism () is any organic, life, living system that functions as an individual entity. All organisms are composed of cells (cell t ...

germination
. The rate of imbibition is dependent on the permeability of the seed coat, amount of water in the environment and the area of contact the seed has to the source of water. For some seeds, imbibing too much water too quickly can kill the seed. For some seeds, once water is imbibed the germination process cannot be stopped, and drying then becomes fatal. Other seeds can imbibe and lose water a few times without causing ill effects, but drying can cause secondary dormancy.


Repair of DNA damage

During seed dormancy, often associated with unpredictable and stressful environments, DNA repair#DNA damage, DNA damage accumulates as the seeds age. In rye seeds, the reduction of DNA integrity due to damage is associated with loss of seed viability during storage. Upon germination, seeds of ''Vicia faba'' undergo DNA repair. A plant DNA ligase that is involved in repair of single- and double-strand breaks during seed germination is an important determinant of seed longevity. Also, in Arabidopsis seeds, the activities of the DNA repair enzymes Poly ADP ribose polymerases (PARP) are likely needed for successful germination. Thus DNA damages that accumulate during dormancy appear to be a problem for seed survival, and the enzymatic repair of DNA damages during germination appears to be important for seed viability.


Inducing germination

A number of different strategies are used by gardeners and horticulturists to break seed dormancy. Scarification allows water and gases to penetrate into the seed; it includes methods to physically break the hard seed coats or soften them by chemicals, such as soaking in hot water or poking holes in the seed with a pin or rubbing them on sandpaper or cracking with a press or hammer. Sometimes fruits are harvested while the seeds are still immature and the seed coat is not fully developed and sown right away before the seed coat become impermeable. Under natural conditions, seed coats are worn down by rodents chewing on the seed, the seeds rubbing against rocks (seeds are moved by the wind or water currents), by undergoing freezing and thawing of surface water, or passing through an animal's digestive tract. In the latter case, the seed coat protects the seed from digestion, while often weakening the seed coat such that the embryo is ready to sprout when it is deposited, along with a bit of fecal matter that acts as fertilizer, far from the parent plant. Microorganisms are often effective in breaking down hard seed coats and are sometimes used by people as a treatment; the seeds are stored in a moist warm sandy medium for several months under nonsterile conditions. Stratification (botany), Stratification, also called moist-chilling, breaks down physiological dormancy, and involves the addition of moisture to the seeds so they absorb water, and they are then subjected to a period of moist chilling to after-ripen the embryo. Sowing in late summer and fall and allowing to overwinter under cool conditions is an effective way to stratify seeds; some seeds respond more favorably to periods of oscillating temperatures which are a part of the natural environment. Leaching or the soaking in water removes chemical inhibitors in some seeds that prevent germination. Rain and melting snow naturally accomplish this task. For seeds planted in gardens, running water is best – if soaked in a container, 12 to 24 hours of soaking is sufficient. Soaking longer, especially in stagnant water, can result in oxygen starvation and seed death. Seeds with hard seed coats can be soaked in hot water to break open the impermeable cell layers that prevent water intake. Other methods used to assist in the germination of seeds that have dormancy include prechilling, predrying, daily alternation of temperature, light exposure, potassium nitrate, the use of plant growth regulators, such as gibberellins, cytokinins, ethylene, thiourea, sodium hypochlorite, and others. Some seeds germinate best after a fire. For some seeds, fire cracks hard seed coats, while in others, chemical dormancy is broken in reaction to the presence of smoke. Liquid smoke is often used by gardeners to assist in the germination of these species.


Sterile seeds

Seeds may be sterile for few reasons: they may have been irradiated, unpollinated, cells lived past expectancy, or bred for the purpose.


Evolution and origin of seeds

The issue of the origin of seed plants remains unsolved. However, more and more data tends to place this origin in the middle Devonian. The description in 2004 of the proto-seed ''Runcaria heinzelinii'' in the Givetian of Belgium is an indication of that ancient origin of seed-plants. As with modern ferns, most land plants before this time reproduced by sending into the air spores that would land and become whole new plants. Taxonomists have described early "true" seeds from the upper Devonian, which probably became the theater of their true first evolutionary radiation. With this radiation came an evolution of seed size, shape, dispersal and eventually the radiation of gymnosperms and angiosperms and
monocotyledon Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one Embryo#Plant embryos, embryonic leaf, or cotyledon. Th ...
s and
dicotyledon The dicotyledons, also known as dicots (or more rarely dicotyls), are one of the two groups into which all the flowering plant The flowering plants, also known as Angiospermae (), or Magnoliophyta (), are the most diverse group of Embryophyte ...
s. Seed plants progressively became one of the major elements of nearly all ecosystems.


True to the seed

Also called growing true, refers to plants whose seed will yield the same type of plant as the original plant. Open pollinated plants, which include heirlooms, will almost always grow true to seed if another variety does not cross-pollinate them.


Seed microbiome

Seeds harbor a diverse microbial community. Most of these microorganisms are transmitted from the seed to the developing seedlings.


Economic importance


Seed market

In the United States farmers spent $22 billion on seeds in 2018, a 35 percent increase since 2010. DowDuPont and Monsanto account for 72 percent of corn and soybean seed sales in the U.S. with the average price of a bag of GMO Sweet corn, corn seed is priced at $270.


Edible seeds

Many seeds are edible and the majority of human calories comes from seeds, especially from cereals, legumes and Nut (fruit), nuts. Seeds also provide most cooking oils, many beverages and spices and some important food additives. In different seeds the Embryo#Embryos of plants and animals, seed embryo or the
endosperm 350px, right The endosperm is a tissue produced inside the seed A seed is an embryonic ''Embryonic'' is the twelfth studio album by experimental rock band the Flaming Lips released on October 13, 2009, on Warner Bros. Records, Warner Bros. The ...
dominates and provides most of the
nutrient A nutrient is a substance Substance may refer to: * Substance (Jainism), a term in Jain ontology to denote the base or owner of attributes * Chemical substance, a material with a definite chemical composition * Matter, anything that has mass and t ...
s. The storage
protein Proteins are large biomolecule , showing alpha helices, represented by ribbons. This poten was the first to have its suckture solved by X-ray crystallography by Max Perutz and Sir John Cowdery Kendrew in 1958, for which they received a No ...

protein
s of the embryo and endosperm differ in their amino acid content and physical properties. For example, the gluten of wheat, important in providing the Elasticity (physics), elastic property to bread dough is strictly an endosperm protein. Seeds are used to propagate many crops such as cereals, legumes, forestry, forest trees, turfgrasses, and pasture grasses. Particularly in developing countries, a major constraint faced is the inadequacy of the marketing channels to get the seed to poor farmers. Thus the use of farmer-retained seed remains quite common. Seeds are also eaten by animals (seed predation), and are also fed to livestock or provided as birdseed.


Poison and food safety

While some seeds are edible, others are harmful, poisonous or deadly. Plants and seeds often contain chemical compounds to discourage herbivores and seed predation, seed predators. In some cases, these compounds simply taste bad (such as in Mustard (condiment), mustard), but other compounds are toxic or break down into toxic compounds within the digestive system. Children, being smaller than adults, are more susceptible to poisoning by plants and seeds. A deadly poison, ricin, comes from seeds of the
castor bean ''Ricinus communis'', the castor bean or castor oil plant, is a species of perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used ...
. Reported lethal doses are anywhere from two to eight seeds, though only a few deaths have been reported when castor beans have been ingested by animals. In addition, seeds containing amygdalin – apple, apricot, bitter almond,
peach The peach (''Prunus persica'') is a deciduous In the fields of horticulture Horticulture is the art of cultivating plants in gardens to produce food and medicinal ingredients, or for comfort and ornamental purposes. Horticulturists grow ...

peach
, plum, cherry, quince, and others – when consumed in sufficient amounts, may cause cyanide poisoning. Other seeds that contain poisons include annona, cotton, custard apple, datura, uncooked durian, golden chain, Horse-chestnut (tree), horse-chestnut, Delphinium, larkspur, locoweed, lychee, nectarine, rambutan, rosary pea, sour sop, sugar apple, wisteria, and
yew Yew is a common name given to various species of trees. The name is most prominently given to any of various coniferous Conifers are a group of cone-bearing seed plants The spermatophytes (; ), also known as phanerogams (taxon Phanerogamae ...

yew
. The seeds of the strychnine tree are also poisonous, containing the poison strychnine. The seeds of many legumes, including the common bean (''Phaseolus vulgaris''), contain proteins called lectins which can cause gastric distress if the beans are eaten without cooking. The common bean and many others, including the soybean, also contain trypsin inhibitors which interfere with the action of the digestive enzyme trypsin. Normal cooking processes degrade lectins and trypsin inhibitors to harmless forms.


Other uses

Cotton fiber grows attached to Gossypium, cotton plant seeds. Other seed fibers are from kapok tree, kapok and milkweed. Many important nonfood oils are extracted from seeds. Linseed oil is used in paints. Oil from jojoba and crambe are similar to whale oil. Seeds are the source of some medicines including castor oil, tea tree oil and the quack cancer drug Amygdalin, Laetrile. Many seeds have been used as beads in necklaces and rosaries including Job's tears, Chinaberry, Abrus precatorius, rosary pea, and
castor bean ''Ricinus communis'', the castor bean or castor oil plant, is a species of perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used ...
. However, the latter three are also poisonous. Other seed uses include: * Seeds once used as weights for Weighing scale, balances. * Seeds used as toys by children, such as for the game Conkers. * Resin from ''Clusia rosea'' seeds used to caulk boats. * Nematicide from milkweed seeds. * Cottonseed meal used as animal feed and fertilizer.


Seed records

* The Oldest viable seed, oldest viable carbon-14-dated seed that has grown into a plant was a Judean date palm seed about 2,000 years old, recovered from excavations at Herod the Great's palace on Masada in Israel. It was germinated in 2005. (A reported regeneration of ''Silene stenophylla'' (narrow-leafed campion) from material preserved for 31,800 years in the Siberian permafrost was achieved using fruit tissue, not seed.) * The largest seed is produced by the coco de mer, or "double coconut palm", ''Lodoicea maldivica''. The entire
fruit In botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the ...

fruit
may weigh up to 23 kilograms (50 pounds) and usually contains a single seed. * The smallest seeds are produced by epiphytic orchids. They are only 85 micrometers long, and weigh 0.81 micrograms. They have no
endosperm 350px, right The endosperm is a tissue produced inside the seed A seed is an embryonic ''Embryonic'' is the twelfth studio album by experimental rock band the Flaming Lips released on October 13, 2009, on Warner Bros. Records, Warner Bros. The ...
and contain underdeveloped embryos. * The earliest fossil seeds are around 365 million years old from the Famennian, Late Devonian of West Virginia. The seeds are preserved immature
ovule In seed plant The spermatophytes, also known as phanerogams (taxon Phanerogamae) or phaenogams (taxon Phaenogamae), comprise those plants that produce seeds, hence the alternative name seed plants. They are a subset of the embryophytes or l ...

ovule
s of the
plant Plants are predominantly photosynthetic Photosynthesis is a process used by plants and other organisms to Energy transformation, convert light energy into chemical energy that, through cellular respiration, can later be released to fuel ...

plant
''Elkinsia polymorpha''.


In religion

The Book of Genesis in the Old Testament begins with an explanation of how all plant forms began:
And God said, Let the earth bring forth grass, the herb yielding seed, and the fruit tree yielding fruit after his kind, whose seed is in itself, upon the earth: and it was so. And the earth brought forth grass, and herb yielding seed after its kind, and the tree yielding fruit, whose seed was in itself, after its kind: and God saw that it was good. And the evening and the morning were the third day.
The Quran speaks of seed germination thus:
It is Allah Who causeth the seed-grain and the date-stone to split and sprout. He causeth the living to issue from the dead, and He is the one to cause the dead to issue from the living. That is Allah: then how are ye deluded away from the truth?Quran, Translation: Abdullah Yusuf Ali, Al-An'aam 95:6


See also

* Biological dispersal * Carpology * Genetically modified crops * List of world's largest seeds * Recalcitrant seed * Seed company * Seed enhancement * Seed library * Seed orchard * Seed paper * Seed saving * Seed testing * Seed trap * Seedbed * Soil seed bank * Selective embryo abortion


References


Bibliography

*
A.C. Martin. The Comparative Internal Morphology of Seeds. American Midland Naturalist Vol. 36, No. 3 (Nov., 1946), pp. 513–660

M.B. McDonald, Francis Y. Kwong (eds.). Flower Seeds: Biology and Technology. CABI, 2005.
*
also available on lineVolume I

Edred John Henry Corner. The Seeds of Dicotyledons. Cambridge University Press, 1976.

United States Forest Service. Woody Plant Seed Manual. 1948

Stuppy, W. Glossary of Seed and Fruit Morphological Terms. Royal botanical gardens, Kew 2004


External links



*[https://web.archive.org/web/20070528090503/http://www.kew.org/msbp/ The Millennium Seed Bank Project] Kew Garden's ambitious preservation project
The Svalbard Global Seed Vault
– a backup facility for the world's seed banks
Plant Physiology online: Types of Seed Dormancy and the Roles of Environmental Factors
*[https://web.archive.org/web/20141218084446/http://theseedsite.co.uk/ The Seed Site]: collecting, storing, sowing, germinating, and exchanging seeds, with pictures of seeds, seedpods and seedlings.
Plant Fix: Check out various plant seeds and learn more information about them
{{Authority control Seeds, Botany Plant reproduction Plant sexuality