Zoarcoidei is a suborder of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

es belonging to the order

Scorpaeniformes The Scorpaeniformes are a diverse order of ray-finned fish, including the lionfishes and sculpins, but have also been called the Scleroparei. It is one of the five largest orders of bony fishes by number of species, with over 1,320. They are ...

. The suborder includes the

wolffish Anarhichadidae, the wolffishes, sea wolves or wolf eels, is a family of marine ray finned fishes belonging to the order Scorpaeniformes. These are predatory, eel shaped fishes which are native to the cold waters of the Arctic, North Pacific and ...

es, gunnels and

eelpout The eelpouts are the ray-finned fish family Zoarcidae. As the common name suggests, they are somewhat eel-like in appearance. All of the roughly 300 species are marine and mostly bottom-dwelling, some at great depths. Eelpouts are predominantly ...

s. The suborder includes about 400 species. These fishes predominantly found in the boreal seas of the northern hemisphere but they have colonised the southern hemisphere.

Taxonomy

Zoarcoidei was first proposed as a taxonomic grouping by the American zoologist Theodore Gill in 1893 as the

superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...

Zoarceoidea. The 5th edition of ''

Fishes of the World ''Fishes of the World'' by the American ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classification of the ...

'' classifies the Zoarcoidei as a suborder within the order Scorpaeniformes. Other authorities classify this taxon as the

infraorder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...

Zoarcales wihin the suborder Cottoidei of the

Perciformes Perciformes (), also called the Percomorpha or Acanthopteri, is an order or superorder of ray-finned fish. If considered a single order, they are the most numerous order of vertebrates, containing about 41% of all bony fish. Perciformes means ...

because removing the Scorpaeniformes from the Perciformes renders that taxon non

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...

. The monophyly of this grouping has still not been fully ascertained but it is generally accepted that the most basal

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

is Bathymasteridae.

Timeline

ImageSize = width:1000px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:10px Period = from:-65.5 till:10 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:5 start:-65.5 ScaleMinor = unit:year increment:1 start:-65.5 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:NAM2 bar:NAM3 bar:NAM4 bar:NAM5 bar:NAM6 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -65.5 till: -55.8 color:paleocene text:

Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...

from: -55.8 till: -33.9 color:eocene text:

Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...

from: -33.9 till: -23.03 color:oligocene text:

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...

from: -23.03 till: -5.332 color:miocene text:

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text:

Pleist. The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...

from: -0.0117 till: 0 color:holocene text: H. bar:eratop from: -65.5 till: -23.03 color:paleogene text:

Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...

from: -23.03 till: -2.588 color:neogene text:

Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...

from: -2.588 till: 0 color:quaternary text: Q. PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left color:miocene bar:NAM1 from: -11.608 till: 0 text: Araeosteus color:pliocene bar:NAM2 from: -5.332 till: 0 text:

Anarhichas ''Anarhichas'' is a genus of marine ray-finned fishes belonging to the family Anarhichadidae, the wolffishes or sea wolves. These fishes are found in the northern North Atlantic and North Pacific oceans. Taxonomy ''Anarhichas'' was first propos ...

color:pliocene bar:NAM3 from: -5.332 till: 0 text: Lycodopsis color:pliocene bar:NAM4 from: -5.332 till: 0 text: Lyconectes color:pleistocene bar:NAM5 from: -2.588 till: 0 text:

Anarrhichthys The wolf eel (''Anarrhichthys ocellatus'') is a species of marine ray-finned fish belonging to the family Anarhichadidae, the wolf fishes. It is found in the North Pacific Ocean. Despite its common name and resemblance, it is not a true eel. It ...

color:pleistocene bar:NAM6 from: -2.588 till: 0 text:

Zoarces ''Zoarces'' is a genus of marine ray-finned fishes belonging to the family (biology), family Zoarcidae, the eelpouts. It is the only genus in the subfamily Zoarcinae. These eelpouts are found in the northern Atlantic and northern Pacific Oceans. ...

PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: -65.5 till: -55.8 color:paleocene text:

from: -55.8 till: -33.9 color:eocene text:

from: -33.9 till: -23.03 color:oligocene text:

from: -23.03 till: -5.332 color:miocene text:

from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text:

from: -0.0117 till: 0 color:holocene text: H. bar:era from: -65.5 till: -23.03 color:paleogene text:

from: -23.03 till: -2.588 color:neogene text:

from: -2.588 till: 0 color:quaternary text: Q.

Families

Zoarcoidei has the following families classified under it: *

Anarhichadidae Anarhichadidae, the wolffishes, sea wolves or wolf eels, is a family of marine ray finned fishes belonging to the order Scorpaeniformes. These are predatory, eel shaped fishes which are native to the cold waters of the Arctic, North Pacific and ...

Bonaparte, 1835 (

es) * Bathymasteridae

Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...

Gilbert Gilbert may refer to: People and fictional characters * Gilbert (given name), including a list of people and fictional characters *Gilbert (surname), including a list of people Places Australia * Gilbert River (Queensland) * Gilbert River (South ...

, 1883 ( ronquils) * Cryptacanthodidae

Gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...

, 1861 *

Pholidae ''Pholidae'' is a family of marine ray-finned fishes, known as gunnels, in the scorpaeniform suborder Zoarcoidei. These are fishes of the littoral zone and are mainly found in North Pacific Ocean, with two species found in the North Atlantic Ocea ...

Gill, 1893 (gunnels) * Ptilichthyidae Jordan & Gilbert, 1883 (

Quillfish The quillfish, (''Ptilichthys goodei''), is a species of marine ray-finned fish, it is the only species in the genus ''Ptilichthys'' and family Ptilichthyidae. This fish occurs in the northern North Pacific Ocean. Taxonomy The quillfish was fir ...

) * Scytalinidae Jordan & Starks, 1895 * Stichaeidae Gill, 1864 (

prickleback Stichaeidae, the pricklebacks or shannies, are a family of marine ray-finned fishes in the suborder Zoarcoidei of the order Scorpaeniformes. Most species are found in the North Pacific Ocean with a few in the North Atlantic Ocean. Taxonomy St ...

s) * Eulophiidae H. M. Smith, 1902 (Spinous eelpouts) *

Zaproridae The prowfish (''Zaprora silenus'') is a species of Scorpaeniformes, scorpaeniform marine fish found in the northern Pacific Ocean. It is the only Extant taxon, extant member of the family, Zaproridae. Another species, ''Araeosteus rothi'', is k ...

Jordan, 1896 ( prowfish) *

Zoarcidae The eelpouts are the ray-finned fish family (biology), family Zoarcidae. As the common name suggests, they are somewhat eel-like in appearance. All of the roughly 300 species are ocean, marine and mostly bottom-dwelling, some at great depths. Eel ...

Swainson, 1839 (

Etymology

Zoarcoidei is based on the genus name ''

'' which was coined by

Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuvier ...

in 1829 and which means "live bearing", as in the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

Zoarces viviparus The viviparous eelpout (''Zoarces viviparus''), also known as the, viviparous blenny and European eelpout is species of marine ray-finned fish belonging to the family Zoarcidae, the eelpouts. It is notable for being ovoviviparous and gives birth ...

'', the viviparous blenny.

Characteristics

The Zoarcoidei families all share a single feature, the possession of a single nostril, and there is no other features or group of features which mark out the Zoarcoids as a taxonomic grouping.

Distribution

The Zoarcoidei is thought to have originated in the northern hemisphere, particularly the northwestern Pacific Ocean and one of the families, the Zoarcidae, has colonised the southern hemisphere on a number of occasions.

References

* {{Taxonbar, from=Q300930 Scorpaeniformes Ray-finned fish suborders Taxa named by Theodore Gill