Wildlife Of North Macedonia
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Over 22,500 species of wildlife have been recorded in
North Macedonia North Macedonia, ; sq, Maqedonia e Veriut, (Macedonia before February 2019), officially the Republic of North Macedonia,, is a country in Southeast Europe. It gained independence in 1991 as one of the successor states of Socialist Feder ...
. Over 10,000 of these are insects, which include 3,000 beetle species and large numbers of
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
, flies, and
Hymenoptera Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Par ...
. Aside from insects, other large arthropod groups include
Chelicerata The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including Opiliones, harvestmen, scorpions, spiders, Solifugae, solif ...
(mostly spiders) and crustaceans. Among vertebrates, more than 300 species of birds recorded, although not all nest in the country. There are over 80 species of both fish and mammals, 32 reptiles, and 14 amphibians. Over 4,200 plants have been identified, of which more than 3,700 are
vascular plant Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They al ...
s. The majority of existing forest is
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
, and the amount of forest has expanded slightly in recent years. Over 2,000 species of algae have been found, most of them within lakes. There are also 2,000 species of identified fungi, with 90% of these being
Basidiomycota Basidiomycota () is one of two large divisions that, together with the Ascomycota, constitute the subkingdom Dikarya (often referred to as the "higher fungi") within the kingdom Fungi. Members are known as basidiomycetes. More specifically, Basi ...
, and at least 450
lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.topography Topography is the study of the forms and features of land surfaces. The topography of an area may refer to the land forms and features themselves, or a description or depiction in maps. Topography is a field of geoscience and planetary sci ...
has contributed to large variation within local climates, which together with the presence of ice age refugia has resulted in significant diversity and
endemism Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
. Although North Macedonia is landlocked, it has numerous rivers and lakes supporting aquatic wildlife. The forest and caves in the west of the country support a number of unique and endangered animals. The three largest lakes, Lake Ohrid, Lake Prespa, and Lake Dojran, are further areas of relatively high diversity. There is substantial interaction between people and wildlife, especially in rural areas. Various plant and fungi species are foraged for local use and for export, and many animal species are hunted. Other threats to wildlife include land use change, pollution, and climate change. A number of animal and plant species have become locally extinct. A variety of laws protect some species and habitats and regulate their use, and action plans have been developed by the
government A government is the system or group of people governing an organized community, generally a state. In the case of its broad associative definition, government normally consists of legislature, executive, and judiciary. Government is a ...
, covering topics including the environment, biodiversity, and water. Four national parks have been established, and North Macedonia is party to a number of European and International conventions that relate to wildlife and the environment.


Environment

North Macedonia spans in the middle of the
Balkan Peninsula The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
. The landlocked country is centred around the Vardar river valley, with the national borders being marked by mountain ranges. Its diverse landscape creates a variety of local climates, which can be divided into eight
biome A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
s. The varied
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s of the country, including both natural and man-made environments, can be divided into 28 broad types, which are further subdivided into 120 at the third level of EUNIS classification. These are spread across 38 classes of landscapes. The overall climate is
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
, with the area undergoing four seasons throughout the year. However, the varied geography means that there is significant local climatic variety.
Precipitation In meteorology, precipitation is any product of the condensation of atmospheric water vapor that falls under gravitational pull from clouds. The main forms of precipitation include drizzle, rain, sleet, snow, ice pellets, graupel and hail. ...
also varies significantly. Up to of snow can fall on some mountains. North Macedonia contains of
surface water Surface water is water located on top of land forming terrestrial (inland) waterbodies, and may also be referred to as ''blue water'', opposed to the seawater and waterbodies like the ocean. The vast majority of surface water is produced by prec ...
. Its 35 rivers fall into three
river basins A drainage basin is an area of land where all flowing surface water converges to a single point, such as a river mouth, or flows into another body of water, such as a lake or ocean. A basin is separated from adjacent basins by a perimeter, the ...
, with one basin each flowing into the
Aegean Sea The Aegean Sea ; tr, Ege Denizi (Greek language, Greek: Αιγαίο Πέλαγος: "Egéo Pélagos", Turkish language, Turkish: "Ege Denizi" or "Adalar Denizi") is an elongated embayment of the Mediterranean Sea between Europe and Asia. It ...
, the
Adriatic Sea The Adriatic Sea () is a body of water separating the Italian Peninsula from the Balkan Peninsula. The Adriatic is the northernmost arm of the Mediterranean Sea, extending from the Strait of Otranto (where it connects to the Ionian Sea) to t ...
, and the
Black Sea The Black Sea is a marginal mediterranean sea of the Atlantic Ocean lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bounded by Bulgaria, Georgia, Roma ...
. The majority of the country, , falls within the Agean Basin, which contains Lake Dojran. of this basin is covered by the Vardar river and its tributaries. The remaining of the Aegean drainage basin is covered by the
Strumica Strumica ( mk, Струмица, ) is the largest city2002 census results
in English and Macedon ...
river and its tributaries. The main river of the Adriatic basin is Black Drim, which begins at Lake Ohrid. Lake Prespa also falls within this basin. Under some accounting there are also three minor drainage basins, which combined cover just over 1% of North Macedonia. Altogether, lakes cover 2% of the country, about . The three large lakes mentioned above are
tectonic lakes A lake is an area filled with water, localized in a Depression (geology), basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the World Ocean, oce ...
. Lake Ohrid is , and has an average depth of . It is thought to be 2–3 million years old. Lake Prespa is , with an average depth of . Lake Dojran is , with a maximum depth of just . These three lakes are split between North Macedonia and neighbouring countries. Around 43
glacial lakes A glacial lake is a body of water with origins from glacier activity. They are formed when a glacier erodes the land and then melts, filling the depression created by the glacier. Formation Near the end of the last glacial period, roughly 10,0 ...
exist in the mountains. There are also 111 are artificial lakes, including large
reservoir A reservoir (; from French ''réservoir'' ) is an enlarged lake behind a dam. Such a dam may be either artificial, built to store fresh water or it may be a natural formation. Reservoirs can be created in a number of ways, including contro ...
s. A complex geological history means the country's surface rocks originate from many periods of Earth's history. There are 13 mountains whose heights exceed , with the highest being above sea level. Around 50% of the country is considered mountainous, containing many gorges and valleys. Caves, especially in limestone mountains found in the west, support a variety of cave fauna, and contain their own lakes. Forests cover of the country, 90% of which is within state-owned land. Agricultural land covers , including both actively used land and land that is able to be quickly cultivated.
Pasture Pasture (from the Latin ''pastus'', past participle of ''pascere'', "to feed") is land used for grazing. Pasture lands in the narrow sense are enclosed tracts of farmland, grazed by domesticated livestock, such as horses, cattle, sheep, or swine ...
s make up 757,000 of this agricultural land.


Plants

At least 3700 species of
vascular plant Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They al ...
s have been identified in North Macedonia, of which 116 are
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
or near-endemic. 3200 of these are
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s. Additionally, there are 573 non-vascular plant species, none of which are endemic. Over 400 of these species are
moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...
es, while 100 are
liverworts The Marchantiophyta () are a division of non-vascular land plants commonly referred to as hepatics or liverworts. Like mosses and hornworts, they have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of ge ...
. The estimated wood mass in the country was in 2008. Forested land expanded by 3.5% in the decade leading up to 2013, reflecting a decrease in annual logging, deliberate efforts to expand forested area, and a reduction in livestock. This recovery is not shared by all forest types however, with
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extant ...
and mixed forest coverage reducing in size. As of 2013, 58% of the forest was
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
, 30% was mixed, 7% was coniferous, and 5% was considered degraded. Some deliberate forestation has been undertaken with the
invasive Invasive may refer to: *Invasive (medical) procedure *Invasive species *Invasive observation, especially in reference to surveillance *Invasively progressive spread of disease from one organ in the body to another, especially in reference to cancer ...
''
Robinia pseudoacacia ''Robinia pseudoacacia'', commonly known in its native territory as black locust, is a medium-sized hardwood deciduous tree, belonging to the tribe Robinieae of the legume family Fabaceae. It is endemic to a few small areas of the United States, ...
'' and '' Ailanthus altissima''. Broadleaf tree woodlands, dominated by alder,
birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech-oak family Fagaceae. The genus ''Betula'' contains 30 ...
, poplar, and
willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist s ...
species, are some of the country's most endangered habitats. Swamp woodlands are dominated by ''
Alnus glutinosa ''Alnus glutinosa'', the common alder, black alder, European alder, European black alder, or just alder, is a species of tree in the family Betulaceae, native to most of Europe, southwest Asia and northern Africa. It thrives in wet locations whe ...
''. Beech forests are dominated by
oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...
s and
beech Beech (''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to temperate Europe, Asia, and North America. Recent classifications recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engle ...
species. Oaks,
chestnut The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. They are native to temperate regions of the Northern Hemisphere. The name also refers to the edible nuts they produce. The unrelat ...
s, hophornbeams, and ''
Carpinus orientalis ''Carpinus orientalis'', known as the Oriental hornbeam, is a hornbeam native to Hungary, the Balkans, Italy, Crimea, Turkey, Iran, and the Caucasus. and occurs usually on hot dry sites at lower altitudes in comparison to the ''Carpinus betulus' ...
'' dominate deciduous woodlands. Other habitats contain many conifer species. 314 species of flowering plants (11% of all flowering plants in the country) are protected. 14 species of vascular plants assessed by the
United Nations Environment Programme The United Nations Environment Programme (UNEP) is responsible for coordinating responses to environmental issues within the United Nations system. It was established by Maurice Strong, its first director, after the United Nations Conference on th ...
are considered at a risk of extinction: four are critically endangered, five are endangered, and five are vulnerable. Eight moss species are protected. Only 11 Macedonian species appear in the Bern Convention, whereas five are listed in
European Union The European Union (EU) is a supranational political and economic union of member states that are located primarily in Europe. The union has a total area of and an estimated total population of about 447million. The EU has often been des ...
Habitats Directive appendices. Plant species that have become locally extinct in North Macedonia include '' Acorus calamus'', ''
Sagittaria sagittifolia ''Sagittaria sagittifolia'' (also called arrowhead due to the shape of its leaves) is a flowering plant in the family Alismataceae, native to wetlands most of Europe from Ireland and Portugal to Finland and Bulgaria, and in Russia, Ukraine, Siber ...
'', ''
Lysimachia thyrsiflora ''Lysimachia thyrsiflora'', the tufted loosestrife, is a plant in the genus ''Lysimachia''. It is native to large sections of the northern Northern Hemisphere, including Eurasia and North America. It often grows in marshes, shorelines of lakes an ...
'', ''
Aldrovanda vesiculosa ''Aldrovanda vesiculosa'', commonly known as the waterwheel plant, is the sole extant species in the flowering plant genus ''Aldrovanda'' of the family Droseraceae. The plant captures small aquatic invertebrates using traps similar to those of ...
'', and ''
Nymphaea alba ''Nymphaea alba'', the white waterlily, European white water lily or white nenuphar , is an aquatic flowering plant in the family Nymphaeaceae. It is native to North Africa, temperate Asia, Europe and tropical Asia (Jammu and Kashmir). Descript ...
''. Others, such as '' Jacobaea paludosa'', '' Ranunculus lingua'', and ''
Gentiana pneumonanthe ''Gentiana pneumonanthe'', the marsh gentian, is a species of the genus ''Gentiana ''Gentiana'' is a genus of flowering plants belonging to the gentian family (Gentianaceae), the tribe Gentianeae, and the monophyletic subtribe Gentianinae. ...
'', are near locally extinct if not extinct already. '' Carex elata'' is also on the brink of extinction.


Fungi

Of the 2000 identified
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
taxa in North Macedonia, none are endemic. Around 1800 belong to
Basidiomycota Basidiomycota () is one of two large divisions that, together with the Ascomycota, constitute the subkingdom Dikarya (often referred to as the "higher fungi") within the kingdom Fungi. Members are known as basidiomycetes. More specifically, Basi ...
(including 550 within
Agaricales The fungal order Agaricales, also known as gilled mushrooms (for their distinctive gills) or euagarics, contains some of the most familiar types of mushrooms. The order has 33 extant families, 413 genera, and over 13,000 described species, alo ...
and 450 within
Aphyllophorales The Aphyllophorales is an obsolete order of fungi in the Basidiomycota. The order is entirely artificial, bringing together a miscellany of species now grouped among the clavarioid fungi, corticioid fungi, cyphelloid fungi, hydnoid fungi, and p ...
), while 200 species are
Ascomycota Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The def ...
. In addition, at least 450 species of
lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. ''
Entomophaga maimaiga ''Entomophaga maimaiga'' is a Japanese fungus which has shown striking success in managing spongy moth populations in North America. Etymology ''Maimaiga'' is the Japanese name for the spongy moth. History In 1908, shortly after classical e ...
'', an entomopathogenic fungus, is found on gypsy moths, following its introduction in neighbouring Bulgaria as a way to control the moth. So far, 122 species have been identified as threatened, and 213 species from the ascomycetes and basidiomycetes have been preliminarily assessed for
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
status. Of these, 21 were critically endangered, 30 endangered, 71 vulnerable, 40 near threatened, 9 least concern, and 42 data deficient. Ten of the critically endangered species have been found in just a single location.


Animals


Invertebrates

There are 13,493 recorded
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
species in North Macedonia. The vast majority of these (90%) are
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s. 6 of the 10
sponge Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through t ...
s found in the three major lakes are endemic. '' Eunapius carteri dojranensis'' is endemic to Lake Doyran. ''
Ochridaspongia rotunda ''Ochridaspongia rotunda'' is freshwater sponge endemic to Lake Ohrid Lake Ohrid ( mk, Охридско Езеро , al, Liqeni i Ohrit , also referred as ''Liqeni i Pogradecit'';) is a lake which straddles the mountainous border between the ...
'', '' Ochridospongia interlithonis'', '' Ochridospongilla stankovici'', and ''
Spongilla stankovici Overview ''Spongilla'' is a genus of freshwater sponges with over 200 different species. Spongilla was first publicly recognized in 1696 by Leonard Plukenet and can be found in lakes, ponds and slow streams.''Spongilla'' have a leuconoid body f ...
'' are endemic to Lake Ohrid. ''
Spongilla Overview ''Spongilla'' is a genus of freshwater sponges with over 200 different species. Spongilla was first publicly recognized in 1696 by Leonard Plukenet and can be found in lakes, ponds and slow streams.''Spongilla'' have a leuconoid body f ...
prespensis'' is endemic to Lake Prespa. Of the three
Cnidaria Cnidaria () is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in freshwater and marine environments, predominantly the latter. Their distinguishing feature is cnidocytes, specialized cells that th ...
species, one is invasive. The 229
flatworm The flatworms, flat worms, Platyhelminthes, or platyhelminths (from the Greek πλατύ, ''platy'', meaning "flat" and ἕλμινς (root: ἑλμινθ-), ''helminth-'', meaning "worm") are a phylum of relatively simple bilaterian, unsegment ...
s are made up of 158
Neodermata Neodermata is a clade of rhabditophoran flatworms containing the parasitic groups Trematoda, Monogenea and Cestoda. Description All neodermatans are parasites, in many groups having a free-swimming larval stage. The most striking feature uniti ...
and 71
Turbellaria The Turbellaria are one of the traditional sub-divisions of the phylum Platyhelminthes (flatworms), and include all the sub-groups that are not exclusively parasitic. There are about 4,500 species, which range from to large freshwater forms more ...
. The two known Nematomorpha species known are '' Gordius nonmaculatus'' and '' Gordius aquaticus''. The only known Nemertea species is '' Prostoma graecense'', which was found in Lake Ohrid.
Annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...
s consist of 140 Oligochaeta, 30
leech Leeches are segmented parasitic or predatory worms that comprise the subclass Hirudinea within the phylum Annelida. They are closely related to the oligochaetes, which include the earthworm, and like them have soft, muscular segmented bodie ...
es, and 5 Branchiobdellida. 53 of the 180 identified annelids species are endemic, all in Lake Ohrid. 38 of these endemic species are oligochaetes, and 11 are Hirudinea. There are at least 870
nematode The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-Parasitism, parasitic nematodes also known as eelworms. They are a diverse animal phylum inhab ...
species, including 450 forest-dwelling species, 80 plant
parasite Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has ...
s, and many living in lakes. The
Mollusca Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
species represented are mostly
Gastropoda The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. ...
, which have 301 species. At least 195 of these are land snails, and 107 are aquatic snails. The remaining 19 mollusc species are
Bivalvia Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of w ...
. Of 92 molluscs considered endemic, 88 of them are gastropods and four of them are bivalves. One gastropod species in Lake Prespa is invasive.


Arthropods

At least 11,800 arthropod species have been identified.Red List Of Butterflies (Lepidoptera: Hesperioidea & Papilionoidea) For Republic Of Macedonia Of the 1126
Chelicerata The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including Opiliones, harvestmen, scorpions, spiders, Solifugae, solif ...
species present, at least 60 are considered endemic. The most numerous order within Chelicerata is that of
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s, of which there are 767 species (nine of which are endemic). The next largest taxa is the
Acari Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but genetic analysis does not show clear evid ...
, whose 250 species include 164
Hydrachnidia Hydrachnidia, also known as "water mites", Hydrachnidiae, Hydracarina or Hydrachnellae, are among the most abundant and diverse groups of benthic arthropods, composed of 6,000 described species from 57 families. As water mites of Africa, Asia, ...
and four
Halacaridae Halacaridae is a family of meiobenthic mites found in marine, brackish, and freshwater habitats around the world. It includes more than 1100 described species belonging to 64 genera It is the largest marine radiation of arachnids. Description ...
. Within Halacroidea, the
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
'' Stygohalacarus'' genus, represented by ''Stygohalacarus scupiensis'', is endemic to North Macedonia, as is '' Copidognathus profundus''. The
fen raft spider The great raft spider or fen raft spider (''Dolomedes plantarius'') is a European species of spider in the family Pisauridae. Like other ''Dolomedes'' spiders, it is semiaquatic, hunting its prey on the surface of water. It occurs mainly in neutr ...
, living in marshes near Mount Belasica, is known to eat fish. The remaining identified Cherlicerata species are 54
pseudoscorpion Pseudoscorpions, also known as false scorpions or book scorpions, are small, scorpion-like arachnids belonging to the order Pseudoscorpiones, also known as Pseudoscorpionida or Chelonethida. Pseudoscorpions are generally beneficial to humans sin ...
s, 50 opiliones, three
scorpion Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always end ...
s, and two
Solifugae Solifugae is an order of animals in the class Arachnida known variously as camel spiders, wind scorpions, sun spiders, or solifuges. The order includes more than 1,000 described species in about 147 genera. Despite the common names, they are ne ...
(''
Galeodes ''Galeodes'' is a genus of solifuge Solifugae is an Order (biology), order of animals in the Class (biology), class Arachnida known variously as camel spiders, wind scorpions, sun spiders, or solifuges. The order includes more than 1,000 de ...
elegans'' and ''Galeodes graceus''). 19 Opiliones are endemic, along with 16 pseudoscorpions.


=Insects

= The at least 10,081 species of insects make up around 85% of the country's arthropod species. At least 3,145
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
species have been identified, split between 84
families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideall ...
. The
ground beetle Ground beetles are a large, cosmopolitan distribution, cosmopolitan family (biology), family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it i ...
s make up a large number of these, represented by 573 species. Within the
Staphylinoidea Staphylinoidea is a superfamily of beetles. It is a very large and diverse group with worldwide distribution. Description Adult staphylinoids are generally small beetles no more than a few millimetres long, though Staphylinidae can reach 50 m ...
superfamily there are 516 species, including representatives of the
Staphylinidae The rove beetles are a family (Staphylinidae) of beetles, primarily distinguished by their short elytra (wing covers) that typically leave more than half of their abdominal segments exposed. With roughly 63,000 species in thousands of genera, the ...
(383 species),
Cholevinae Cholevinae is a subfamily of small carrion beetles in the family Leiodidae. There are more than 260 genera and 1,100 described species in Cholevinae. See also For a list of genera in Cholevinae, see List of Leiodidae genera These 379 genera b ...
(57),
Hydraenidae Hydraenidae is a family of very small aquatic beetles, sometimes called "Minute moss beetles", with a worldwide distribution. They are around 0.8 to 3.3 mm in length. The adults store air on the underside of the body as well as beneath the el ...
(42),
Silphidae Silphidae is a family of beetles that are known commonly as large carrion beetles, carrion beetles or burying beetles. There are two subfamilies: Silphinae and Nicrophorinae. Nicrophorines are sometimes known as sexton beetles. The number of sp ...
(14),
Ptiliidae Ptiliidae is a family of very tiny beetles with a cosmopolitan distribution. This family contains the smallest of all beetles, with a length when fully grown of . The weight is approximately 0.4 milligrams. They are colloquially called feat ...
(4), and
Agyrtidae Agyrtidae, or primitive carrion beetles, are a small family of beetles belonging to Staphylinoidea. They are found in mostly temperate areas of the Northern Hemisphere and in New Zealand. Characteristics Agyrtidae are small or middle-sized bee ...
(1) families. Water beetles have been identified from the Dytiscidae (62), Haliplidae (11), Gyrinidae (6), and
Noteridae Noteridae is a family of adephagan water beetles closely related to the Dytiscidae, and formerly classified with them. They are mainly distinguished by the presence of a distinctive "noterid platform" underneath, in the form of a plate between t ...
(2) families. Within
Hydrophiloidea Hydrophiloidea, known as water scavenger beetles, is a superfamily of beetles. Until recently it included only a single family, the Hydrophilidae (water scavenger beetles), but several of the subfamilies have been removed and raised to family ra ...
, there are 68 Histeridae species and 33
Hydrophilidae Hydrophilidae, also known colloquially as water scavenger beetles, is a family of beetles. Aquatic hydrophilids are notable for their long maxillary palps, which are longer than their antennae. Several of the former subfamilies of Hydrophilidae ...
species. There are 14 families of Scarabaeoidea represented, which altogether contain 172 species. Within
Cucujoidea Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark ...
, there are perhaps over 200 species, of which 116 are
Nitidulidae The sap beetles, also known as Nitidulidae, are a family (biology), family of beetles. They are small (2–6 mm) ovoid, usually dull-coloured beetles, with knobbed antenna (biology), antennae. Some have red or yellow spots or bands. They fe ...
and 33 are
Coccinellidae Coccinellidae () is a widespread family of small beetles ranging in size from . They are commonly known as ladybugs in North America and ladybirds in Great Britain. Some entomologists prefer the names ladybird beetles or lady beetles as they ...
. Within
Chrysomeloidea The Chrysomeloidea are an enormous superfamily of beetles, with tens of thousands of species. The largest families are Cerambycidae, long-horned beetles, with more than 35,000 species, and Chrysomelidae, leaf beetles, with more than 13,000 specie ...
, there are 340 leaf beetles and 176
longhorn beetle The longhorn beetles (Cerambycidae), also known as long-horned or longicorns, are a large family of beetles, with over 35,000 species described. Most species are characterized by extremely long antennae, which are often as long as or longer than ...
s. The 88 species of Bostrichiformia are split between four families: Bostrichidae, Dermestidae, Ptinidae, and
Nosodendridae Nosodendridae is a family of beetles, with only 67 species in three extant genera, which are found worldwide. '' Nosodendron'', the largest genus, is found in forests and attracted to yeast generated slime on the wounds of trees, and likely consu ...
. There are 100 species of Elateroidea. Most are click beetles (74), with the rest being soldier beetles (15), fireflies (7), and net-winged beetles (4). There are 44 identified
Cleroidea Cleroidea is a small superfamily of beetles containing over 10,000 species. Most of the members of the group are somewhat slender, often with fairly soft, flexible elytra, and typically hairy or scaly. Description Cleroidea is defined by the fo ...
species. 17 each are from
Malachiinae Malachiinae is a subfamily of beetles of the family Melyridae and having a global distribution. Description Malachiinae have peculiar orange structures along the sides of the abdomen, which may be everted and saclike or withdrawn into the body ...
and
Dasytinae Dasytinae is a subfamily of soft-wing flower beetles, beetles of the family Melyridae, historically sometimes treated as a separate family, "Dasytidae". Description and distribution Dasytinae are typically small (<8 mm) and parallel-sided ...
. One Dasytidae species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the Šar Mountains. The remaining 10 are from the
Cleridae Cleridae are a family of beetles of the superfamily Cleroidea. They are commonly known as checkered beetles. The family Cleridae has a worldwide distribution, and a variety of habitats and feeding preferences. Cleridae have many niches and fe ...
family. Other beetles include 290
Curculionidae The Curculionidae are a family of weevils, commonly called snout beetles or true weevils. They are one of the largest animal families, with 6,800 genera and 83,000 species described worldwide. They are the sister group to the family Brentidae. T ...
and 45
darkling beetle Darkling beetle is the common name for members of the beetle family Tenebrionidae. The number of species in the Tenebrionidae is estimated at more than 20,000 and the family is cosmopolitan in distribution. Taxonomy ''Tenebrio'' is the Latin gen ...
s. A total of 2,638 species of
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
have been identified. These are split between 68 families, with family size ranging from a single species to 528 species (the
Noctuidae The Noctuidae, commonly known as owlet moths, cutworms or armyworms, are a family of moths. They are considered the most controversial family in the superfamily Noctuoidea because many of the clades are constantly changing, along with the other f ...
). 67 species have been provisionally assessed for IUCN Red List status within North Macedonia. One has been identified as endangered, with 15 being vulnerable and 24 being near threatened. There are over 1,500 species of flies, representing 54 families. The largest family is that of the hoverflies, represented by 262 species. Some flies are considered potential
disease vectors In epidemiology, a disease vector is any living agent that carries and transmits an infectious pathogen to another living organism; agents regarded as vectors are organisms, such as parasites or microbes. The first major discovery of a disease vec ...
, and one, '' Obolodiplosis robiniae'', is invasive.
Hymenoptera Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Par ...
is another diverse order, represented by about 1077 species. One relatively well-studied group are
parasitoid wasp Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causin ...
s. 150 species of
Braconidae The Braconidae are a family of parasitoid wasps. After the closely related Ichneumonidae, braconids make up the second-largest family in the order Hymenoptera, with about 17,000 recognized species and many thousands more undescribed. One analysis ...
have been found. The 129 Ichneumonidae species (21 of which fall within Cryptinae) present include three endemics: '' Hadrodactylus tiphae balcanicus'', ''
Mesochorus ''Mesochorus'' is a genus of ichneumon wasps in the family Ichneumonidae The Ichneumonidae, also known as the ichneumon wasps, Darwin wasps, or ichneumonids, are a family (biology), family of parasitoid wasps of the insect order Hymenoptera. T ...
venerandus'', and '' Gelis balcanicus''.
Eulophidae The Eulophidae are a large family of hymenopteran insects, with over 4,300 described species in some 300 genera. The family includes the genus ''Elasmus'', which used to be treated as a separate family, "Elasmidae", and is now treated as a subf ...
has 40 species. Non-parasitic
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...
s include 75 members of the family Vespidae and 37 members of the family Mutillidae. In addition to wasps, there are at least 113 Apidae bee species and 99 ant species. Within
Hemiptera Hemiptera (; ) is an order (biology), order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, Reduviidae, assassin bugs, Cimex, bed bugs, and shield bugs. ...
, 776 species of
Heteroptera The Heteroptera are a group of about 40,000 species of insects in the order Hemiptera. They are sometimes called "true bugs", though that name more commonly refers to the Hemiptera as a whole. "Typical bugs" might be used as a more unequivocal al ...
(15 endemic) have been identified. However, local expertise in this taxon is lacking. It has been suggested that the discovered number makes up only 62% of the total species present, taking into account the rate of new discoveries, the diversity of habitats, and studies from neighbouring countries. Knowledge about Homoptera is also very limited, with most studies covering a small number of species from specific locations. The known Homoptera consist of 134 Sternorrhyncha, 15 Cicadomorpha, and 13 Fulgoromorpha. There are 175 identified
orthoptera Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grassho ...
species. One, ''
Bradyporus ''Bradyporus'' is a genus of bush crickets, found in south-eastern Europe and western and central Asia. It is the type genus of the subfamily Bradyporinae The Bradyporinae are a subfamily in the family Tettigoniidae (bush crickets or katydid ...
macrogaster macrogaste'', is critically endangered. Four are considered endangered, while another eight are considered vulnerable (another 10 are data deficient). Some species are known only from a single location, or even a single record. There are perhaps 106 caddisfly species, although estimates vary. The 68 species of identified
Neuropterida The Neuropterida are a clade, sometimes placed at superorder level, of holometabolous insects with over 5,700 described species, containing the orders Neuroptera (lacewings, antlions), Megaloptera (alderflies, dobsonflies), and Raphidioptera (sna ...
consist of 59
Neuroptera The insect order Neuroptera, or net-winged insects, includes the lacewings, mantidflies, antlions, and their relatives. The order consists of some 6,000 species. Neuroptera can be grouped together with the Megaloptera and Raphidioptera in th ...
and nine
Raphidioptera Snakeflies are a group of predatory insects comprising the order Raphidioptera with two extant families: Raphidiidae and Inocelliidae, consisting of roughly 260 species. In the past, the group had a much wider distribution than it does now; sn ...
. The 13 known
flea Flea, the common name for the order Siphonaptera, includes 2,500 species of small flightless insects that live as external parasites of mammals and birds. Fleas live by ingesting the blood of their hosts. Adult fleas grow to about long, a ...
species include parasites of the
Balkan snow vole The Balkan snow vole (''Dinaromys bogdanovi''), also known as Martino's snow vole, is the only member of the genus ''Dinaromys''. Eight subspecies of this vole have been recognized from southern parts of Europe. The genus name means "Dinaric mou ...
. The country has 68 identified
mayflies Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the orde ...
, including seven ''
Rhithrogena ''Rhithrogena'' is a genus of flatheaded mayflies in the family Heptageniidae The Heptageniidae (synonym: Ecdyonuridae) are a family of mayflies with over 500 described species mainly distributed in the Holarctic, Oriental, and Afrotropical r ...
'' species, all found in the Pena river. Known
Dictyoptera Dictyoptera (from Greek δίκτυον ''diktyon'' "net" and πτερόν ''pteron'' "wing") is an insect superorder that includes two extant orders of polyneopterous insects: the order Blattodea (termites and cockroaches together) and the ord ...
include 12
cockroach Cockroaches (or roaches) are a paraphyletic group of insects belonging to Blattodea, containing all members of the group except termites. About 30 cockroach species out of 4,600 are associated with human habitats. Some species are well-known as ...
es, four mantis, and two
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattode ...
s. Of the 14
lice Louse ( : lice) is the common name for any member of the clade Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera has variously been recognized as an order, infraorder, or a parvorder, as a result o ...
species known, two, '' Enderleinellus ferrisi'' and '' Schizophthirus gliris'', are parasites (of European ground squirrels and edible dormice respectively). '' Haploembia solieri'', which was found in Valandovo, is the only recorded
Embioptera The order (biology), order Embioptera, commonly known as webspinners or footspinners, are a small group of mostly Tropics, tropical and Subtropics, subtropical insects, classified under the Subclass (biology), subclass Pterygota. The order has ...
species. Other insect species include 97
Plecoptera Plecoptera is an order of insects, commonly known as stoneflies. Some 3,500 species are described worldwide, with new species still being discovered. Stoneflies are found worldwide, except Antarctica. Stoneflies are believed to be one of the mo ...
(10 of which are endemic), 64 Odonata, 49
Psocoptera Psocoptera are a paraphyletic group of insects that are commonly known as booklice, barklice or barkflies. The name Psocoptera has been replaced with Psocodea in recent literature, with the inclusion of the former order Phthiraptera into Psocode ...
, 42 Thysanoptera, five
Dermaptera Earwigs make up the insect order Dermaptera. With about 2,000 species in 12 families, they are one of the smaller insect orders. Earwigs have characteristic cerci, a pair of forcep-like pincers on their abdomen, and membranous wings folded ...
, three
Archaeognatha The Archaeognatha are an order of apterygotes, known by various common names such as jumping bristletails. Among extant insect taxa they are some of the most evolutionarily primitive; they appeared in the Middle Devonian period at about the sa ...
, two Mecoptera, one Strepsiptera ('' Hylecthrus rubi''), and one
Zygentoma Zygentoma are an order in the class Insecta, and consist of about 550 known species. The Zygentoma include the so-called silverfish or fishmoths, and the firebrats. A conspicuous feature of the order are the three long caudal filaments. The two ...
. There is no data for
Megaloptera Megaloptera is an order of insects. It contains the alderflies, dobsonflies and fishflies, and there are about 300 known species. The order's name comes from Ancient Greek, from ''mega-'' (μέγα-) "large" + ''pteryx'' (πτέρυξ) "wing ...
within the country.


=Crustaceans

= Of the 490
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
species found, 133 are considered endemic. Within the class Branchiopoda, the most numerous order is Diplostraca, which has 86 species. ''
Alona smirnovi ''Alona smirnovi'' is a species of crustacean in family Chydoridae. It is endemic to North Macedonia. Its natural habitat is inland karst Karst is a topography formed from the dissolution of soluble rocks such as limestone, dolomite, and gypsu ...
'' is endemic to Lake Ohrid. Of the other orders, there are six species of
Anostraca Anostraca is one of the four orders of crustaceans in the class Branchiopoda; its members are referred to as fairy shrimp. They live in vernal pools and hypersaline lakes across the world, and they have even been found in deserts, ice-covered m ...
, including ''
Chirocephalus pelagonicus ''Chirocephalus pelagonicus'' is a species of crustaceans in the family Chirocephalidae. It is endemic to North Macedonia North Macedonia, ; sq, Maqedonia e Veriut, (Macedonia before February 2019), officially the Republic of North Ma ...
'' which is endemic to wetlands in the
Pelagonia Pelagonia ( mk, Пелагонија, Pelagonija; el, Πελαγονíα, Pelagonía) is a geographical region of Macedonia named after the ancient kingdom. Ancient Pelagonia roughly corresponded to the present-day municipalities of Bitola, Pri ...
plain. The two species of order Notostraca (which contains only the Triopsidae family) present are '' Lepidurus apus'' and ''
Triops cancriformis ''Triops cancriformis'', European tadpole shrimp or tadpole shrimp, is a species of tadpole shrimp found in Europe to the Middle East and India. Due to habitat destruction, many populations have recently been lost across its European range, s ...
''. All 172 representatives of the
ostracod Ostracods, or ostracodes, are a class of the Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 70,000 species (only 13,000 of which are extant) have been identified, grouped into several orders. They are small crustaceans, typic ...
s are from the order
Podocopida The Podocopida are an order of ostracods in the subclass Podocopa. It is the most diverse of the four orders of ostracods, and also has a rich fossil record. Taxonomy The following suborders and unassigned taxa are contained in the order Podoco ...
. Numerous genera within this order have high levels of endemism. Of the class
Maxillopoda Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group can ...
, 146 species are
copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...
s: 60 Cyclopoida, 54 Harpacticoida, 30
Calanoida Calanoida is an order of copepods, a group of arthropods commonly found as zooplankton. The order includes around 46 families with about 1800 species of both marine and freshwater copepods between them. Description Calanoids can be distinguis ...
, and two Poecilostomatoida. Both Cyclopoida and Harpacticoida contain several endemic genera. 12 of these copepods are known to be parasitic, found infecting fish in the major lakes. Other Maxillopoda include three
Branchiura The family Argulidae, whose members are commonly known as carp lice or fish lice, are parasitic crustaceans in the class Ichthyostraca. It is the only family in the monotypic subclass Branchiura and the order Arguloida, although a second family, ...
, all from the genus ''
Argulus ''Argulus'' is a genus of fish lice in the family Argulidae The family Argulidae, whose members are commonly known as carp lice or fish lice, are parasitic crustaceans in the class Ichthyostraca. It is the only family in the monotypic subclas ...
'' within the class
Argulidae The family Argulidae, whose members are commonly known as carp lice or fish lice, are parasitic crustaceans in the class Ichthyostraca. It is the only family in the monotypic subclass Branchiura and the order Arguloida, although a second family, ...
. One, ''
Argulus foliaceus ''Argulus foliaceus'', also known as the common fish louse, is a species of fish lice in the family Argulidae. It is "the most common and widespread native argulid in the Palaearctic" and "one of the most widespread crustacean ectoparasites of ...
'', is a parasite of at least three fish species. There is one representative of the
Pentastomida The Pentastomida are an enigmatic group of parasitic arthropods commonly known as tongue worms due to the resemblance of the species of the genus ''Linguatula'' to a vertebrate tongue; molecular studies point to them being degenerate crustaceans. ...
known: ''
Linguatula serrata ''Linguatula serrata'' is a species of cosmopolitan zoonotic parasite, belonging to the tongueworm order Pentastomida. They are wormlike parasites of the respiratory systems of vertebrates. They live in the nasopharyngeal region of mammals. Cat ...
'', which is a human parasite. There are representatives of four orders of
Malacostraca Malacostraca (from New Latin; ) is the largest of the six classes of crustaceans, containing about 40,000 living species, divided among 16 orders. Its members, the malacostracans, display a great diversity of body forms and include crabs, lobst ...
.
Isopoda Isopoda is an order of crustaceans that includes woodlice and their relatives. Isopods live in the sea, in fresh water, or on land. All have rigid, segmented exoskeletons, two pairs of antennae, seven pairs of jointed limbs on the thorax, an ...
is represented by 50 species, of which about a third are endemic.
Amphipoda Amphipoda is an order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods range in size from and are mostly detritivores or scavengers. There are more than 9,900 amphipod species so far desc ...
is represented by 47 species, with five genera (''
Bogidiella ''Bogidiella'' is a genus of crustacean in the family Bogidiellidae Bogidiellidae is a family of amphipod crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, bra ...
'', '' Gammarus'', '' Niphargus'', '' Hadzia'', and '' Ignolfiella'') containing many endemic species.
Decapoda The Decapoda or decapods (literally "ten-footed") are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crabs, lobsters, crayfish, shrimp and prawns. Most decapods are scavengers. The order is esti ...
is represented by five species: two Astacidae (''
Austropotamobius torrentium ''Austropotamobius torrentium'', also called the stone crayfish, is a European species of fresh water, freshwater crayfish in the family Astacidae. It is mostly found in tributaries of the Danube, having originated in the northern part of the Bal ...
'' and '' Astacus astacus''), two
Potamidae Potamidae is a family (biology), family of freshwater crabs. It includes more than 650 species and nearly 100 genera, which are placed into two subfamilies: Potaminae and Potamiscinae. Subfamily Potaminae The Potaminae Ortmann, 1896 are distrib ...
(''
Potamon fluviatile ''Potamon fluviatile'' is a freshwater crab found in or near wooded streams, rivers and lakes in Southern Europe. It is an omnivore with broad ecological tolerances, and adults typically reach in size during their 10–12 year lifespan. They inh ...
'' and ''
Potamon ibericum ''Potamon ibericum'' is a Eurasian species of freshwater crab. It is an omnivore that feeds on land, but returns regularly to the water, and can survive short periods of drought in burrows and under stones. Its range (biology), natural range stre ...
''), and one Atyidae ('' Atyaephyra stankoi'', which was found in Lake Dojran).
Bathynellacea Bathynellacea is an order of crustaceans which live interstitially in groundwater. Some species can tolerate low salt concentrations, and at least one African species is a thermophile, living in hot springs and tolerating temperatures up to . Ba ...
is represented by three species: '' Bathynella natans'', '' Parabathynella stygia'', and '' Bathynella chappuisi''. There are around 100 species of Myriapoda, including 18 endemic millipedes. There are 21 identified species of Entognatha, including 11
Collembola Springtails (Collembola) form the largest of the three lineages of modern hexapods that are no longer considered insects (the other two are the Protura and Diplura). Although the three orders are sometimes grouped together in a class called Ento ...
, eight Protura and two Diplura.


Fish

The total number of fish species differs between sources, potentially due to different classifications regarding what counts as a species. One calculation identifies 85
Actinopterygii Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...
and two lampreys. Of these, 27 are endemic, and 19 are invasive. Some fish are endemic to certain lakes or rivers. Lake Ohrid has 21 native species, of which eight are endemic, and seven are introduced species. Lake Prespa has 11 native species, of which eight are endemic, and 12 introduced species. Lake Dojran has 12 native species, one of which is endemic, along with two introduced species. Of the Actinopterygii, the
European sea sturgeon The European sea sturgeon (''Acipenser sturio''), also known as the Atlantic sturgeon or common sturgeon, is a species of sturgeon native to Europe. It was formerly abundant, being found in coastal habitats all over Europe. It is anadromous and b ...
, the
European eel The European eel (''Anguilla anguilla'') is a species of eel, a snake-like, catadromous fish. They are normally around and rarely reach more than , but can reach a length of up to in exceptional cases. Eels have been important sources of fo ...
, and '' Alburnus macedonicus'' are considered critically endangered, and the Prespa minnow and ''
Salmo peristericus ''Salmo peristericus'', or the Prespa troutKoutseri I et al. (2010Species Action Plan for the Endemic Prespa Trout, ''Salmo peristericus'': A Conservation Toolwww.balwois.com is a variety of trout, a freshwater fish in the family Salmonidae. It i ...
'' are considered endangered. A further 10 species are vulnerable, one near threatened, and 10 data deficient. Salmonidae species in Orhid Lake are endangered, while the
common carp The Eurasian carp or European carp (''Cyprinus carpio''), widely known as the common carp, is a widespread freshwater fish of eutrophic waters in lakes and large rivers in Europe and Asia.Fishbase''Cyprinus carpio'' Linnaeus, 1758/ref>Arkive The ...
is endangered in Lake Prespa. The two lamprey species are the Ukrainian brook lamprey and ''
Eudontomyzon stankokaramani ''Eudontomyzon stankokaramani'', the Drin brook lamprey, is a non-predatory, freshwater resident species of lamprey found in the Drin river system of Albania and Kosovo and the basins of Lakes Ohrid Ohrid ( mk, Охрид ) is a city in Nort ...
''. The first lives in the Vardar's drainage basin, while the second lives in the Adriatic Sea basin. Both are protected by law.


Amphibians

The
amphibian Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...
species found comprise nine
frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
s, and five
salamander Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten ...
s. Three of these have been categorized as endangered, three as vulnerable, three as near threatened, and five as least concern. Considering the effects of disease and habitat loss, that proportion of threatened amphibian species is roughly in line with the state of amphibians worldwide. None of these species are endemic to the country, although two are endemic to the Balkans.


Reptiles

The 32 species of
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s present are made up of 16
snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...
s, 12
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s, and four
tortoise Tortoises () are reptiles of the family Testudinidae of the order Testudines (Latin: ''tortoise''). Like other turtles, tortoises have a turtle shell, shell to protect from predation and other threats. The shell in tortoises is generally hard, ...
s. Within North Macedonia, 11 of these species are widespread, 10 are restricted to certain habitats, and 11 have very limited range. While none of these are endemic to the country, two are endemic to the Balkans. One snake species present is the
Caspian whipsnake The Caspian whipsnake (''Dolichophis caspius'', sometimes also ''Coluber caspius'') also known as the large whipsnake (among various other species in genus ''Dolichophis''/'' Coluber''), is a common species of whipsnake found in the Balkans ...
, Europe's largest snake. Of those with IUCN Red List assessments, seven of them are endangered, six are vulnerable, eight are near threatened, and ten are of least concern. The globally vulnerable ''
Vipera ursinii :''Common names: meadow viper, Ursini's viper, Mallow D, Ludwig D, Nilson G (2003). ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company. . meadow adder,Brown, John Haynes (1973). ''To ...
'', one of three
Viperidae The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...
in North Macedonia, has strict protection, while another 22 species have some level of protection.
Hermann's tortoise Hermann's tortoise (''Testudo hermanni'') is a species of tortoise. Two subspecies are known: the western Hermann's tortoise (''T. h. hermanni'' ) and the eastern Hermann's tortoise (''T. h. boettgeri'' ). Sometimes mentioned as a subspecies, ...
, found in forest and meadows, are expected to lost at least one third of their population over the next 75 years.


Birds

There are 318
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
species with confirmed sightings, and an additional 16 for which there are less reliable claims. None are endemic. 215 of these species nest in the country. 106 bird species have some level of protection under law. At least eight nesting species have become locally extinct, while another 7–15 no longer nest in the country. For many species, fewer than 100 nesting couples remain.


Mammals

At least 85 mammal species inhabit North Macedonia, of which four are endemic to the Balkans: the
western broad-toothed field mouse The western broad-toothed field mouse (''Apodemus epimelas'') is a species of rodent in the genus ''Apodemus'' from southeastern Europe. It is related to '' A. mystacinus'', which occurs further to the east. It is found in Albania, Bosnia and ...
, the
Balkan snow vole The Balkan snow vole (''Dinaromys bogdanovi''), also known as Martino's snow vole, is the only member of the genus ''Dinaromys''. Eight subspecies of this vole have been recognized from southern parts of Europe. The genus name means "Dinaric mou ...
, Felten's vole, and the
Balkan mole The Balkan mole (''Talpa stankovici'') is a species of mammal in the family Talpidae. It is found in Montenegro, Albania, North Macedonia and Greece, including the island of Corfu Corfu (, ) or Kerkyra ( el, Κέρκυρα, Kérkyra, , ; ; ...
. The
chamois The chamois (''Rupicapra rupicapra'') or Alpine chamois is a species of goat-antelope native to mountains in Europe, from west to east, including the Alps, the Dinarides, the Tatra and the Carpathian Mountains, the Balkan Mountains, the Ril ...
sub-species ''Rupicapra rupicapra balcanica'' and lynx sub-species the Balkan lynx are endemic to the Balkans, and have core populations within North Macedonia. It has been suggested that the country's European ground squirrel population be considered its own sub-species. Eight mammal species are considered invasive. The Balkan lynx persists around the Albanian-Macedonian border. Its low population leads it to being rarely seen, and it is the subject of recovery efforts. The lynx has a reputation for killing livestock, despite there being very few recorded incidents. There are perhaps fewer than 100 spread across North Macedonia and Albania.
Brown bear The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is kno ...
s live in mountainous forests in the west of the country. Around 160-200 individuals inhabit these forests, part of the larger Dinaric-
Pindos The Pindus (also Pindos or Pindhos; el, Πίνδος, Píndos; sq, Pindet; rup, Pindu) is a mountain range located in Northern Greece and Southern Albania. It is roughly 160 km (100 miles) long, with a maximum elevation of 2,637 metres ...
population. About 70 are found in Mavrovo National Park. The population of
wolves The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
numbered around 800–1000 as of 2013. Eurasian otters can be found throughout most waterways, although they are absent from a couple of areas with particularly high levels of pollution. Golden jackals were considered extinct in the area by the 1960s, but became re-established in forested mountainous areas, especially in the west, during the 21st century. An individual raccoon dog, a species which was introduced to European Russia in the 20th century, was found dead in the north of the country, indicating the species is spreading southwards. Herbivores include
roe deer The roe deer (''Capreolus capreolus''), also known as the roe, western roe deer, or European roe, is a species of deer. The male of the species is sometimes referred to as a roebuck. The roe is a small deer, reddish and grey-brown, and well-adapt ...
, chamois,
wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species is ...
, and European hares. European ground squirrels are found on Mount Mokra. The
European wildcat The European wildcat (''Felis silvestris'') is a small wildcat species native to continental Europe, Scotland, Turkey and the Caucasus. It inhabits forests from the Iberian Peninsula, Italy, Central and Eastern Europe to the Caucasus. Its fur is ...
, Balkan lynx, Eurasian otter, brown bear, European ground squirrel, and Balkan snow vole have strict protection under national legislation, and another 10 mammal species have lesser levels of protection. Three bats, two rodents, and one mustelid are listed as threatened in the IUCN Red List.


Algae

Knowledge of
algal Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...
species in North Macedonia remains patchy, with inconsistent research and several taxa that remain not fully identified. There are 2095 identified forms of algae, with the vast majority being
diatom A diatom (Neo-Latin ''diatoma''), "a cutting through, a severance", from el, διάτομος, diátomos, "cut in half, divided equally" from el, διατέμνω, diatémno, "to cut in twain". is any member of a large group comprising sev ...
s. It is thought at least 10% of these are endemic, with at least 194 having been so far identified as such. Most research on algae has taken place in Lakes Ohrid and Prespa. Lake Ohrid by itself holds 798 identified taxa, with 158 being endemic to just that lake.


Diversity and endemism hotspots

Within North Macedonia, there are multiple areas that are likely high in endemism. Lake Ohrid is an extensively studied area, and its diversity is reflected in a high number of endemic species. Lake Prespa lies quite close to Lake Ohrid, and the two lakes are connected
hydrologically Hydrology () is the scientific study of the movement, distribution, and management of water on Earth and other planets, including the water cycle, water resources, and environmental watershed sustainability. A practitioner of hydrology is call ...
. While Lake Prespa has fewer species than Lake Ohrid and while the two lakes share similar species compositions, Lake Prespa has its own endemic species and species more closely related to more western water bodies than to Lake Ohrid. Within Lake Ohrid alone there are over 100 insects, 75 flatworms (35 endemic to the lake and nearby waterbodies), 72 gastropods (56 endemic), 52 ostracods (33 endemic), 49
rotifer The rotifers (, from the Latin , "wheel", and , "bearing"), commonly called wheel animals or wheel animalcules, make up a phylum (Rotifera ) of microscopic and near-microscopic pseudocoelomate animals. They were first described by Rev. John H ...
s, 43 Acari, 36 oligochaetes (17 endemic), 36 copepods (six endemic), 31 Cladocera (one endemic), 30 endemic
ciliate The ciliates are a group of alveolates characterized by the presence of hair-like organelles called cilia, which are identical in structure to flagellum, eukaryotic flagella, but are in general shorter and present in much larger numbers, with a ...
s, 24 leeches (12 endemic), 24 nematodes (three endemic), 14 amoebas, 13 bivalves (two endemic), 10–11 Amphipoda (9 endemic), four isopods (3 endemic), four sponges, and two decapods. The ''
Ancylus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. (2 ...
'' genus species in this lake are monophyletic, likely evolving within the lake itself. The same situation is true for leeches from the genus ''Dina''. Lake Prespa is less well studied, but is known to have at over 100 insects, 90 crustaceans, 60 rotifers, 50 flatworms, 36 molluscs (27 of which are snails), 35 annelids, and three sponges. Seven of these snails are endemic, as is the mollusc ''
Pisidium ''Pisidium'' is a genus of very small or minute freshwater clams known as pill clams or pea clams, aquatic bivalve molluscs in the family Sphaeriidae, the pea clams and fingernail clams. In some bivalve classification systems, the family Spha ...
maasseni''. ''
Radix In a positional numeral system, the radix or base is the number of unique digits, including the digit zero, used to represent numbers. For example, for the decimal/denary system (the most common system in use today) the radix (base number) is t ...
'' snail species in both Ohrid and Lake Prespas are related, and there are endemic species within springs near the
Monastery of Saint Naum The Monastery of Saint Naum ( mk, Манастир „Свети Наум“) is an Eastern Orthodox monastery in North Macedonia, named after the medieval Bulgarian writer and enlightener Saint Naum who founded it. It is situated along Lake Ohri ...
that are unrelated to the species found in the lakes. Lake Dojran has 17 rotifers. It is expected that mountainous areas will also contain a number of endemic species. North Macedonia contains many areas of ice age refugia, which retain significant plant diversity. Plant endemism and sub-endemism is high around mountainous areas. Mammal diversity is highest in the mountainous west of the country. Caves in the west, especially within the drainage basins of the Radika, Galichica, Jakupica, and Poreche rivers, are thought to have rates of invertebrate endemism of around 90%. 57 species of
stygofauna Stygofauna are any fauna that live in groundwater systems or aquifers, such as caves, fissures and vugs. Stygofauna and troglofauna are the two types of subterranean fauna (based on life-history). Both are associated with subterranean environments ...
are known, including 14 pseudoscorpions, 12 beetles, and 10 isopods.


Human influence

There is significant interaction between the people of North Macedonia and wildlife, especially in rural areas. This continues despite rural areas decreasing in population by 2.2% from 2005 to 2010. Foraging for wild plants remains a common activity for those in rural areas. Around 700 plant species are considered medicinal or aromatic, with 220 frequently used. A quarter of known fungi species are edible, and some fungi and lichens are collected for use and export. Carnivores, such as bears, wolves, and lynx, are viewed negatively in some areas. Bears and wolves are known to attack livestock such as sheep, cattle, and goats. In some instances wolves have been reported to hunt up to 38% of sheep flocks, and an even higher percentage of goats. Foxes and wild cats are reported to kill poultry. There are reports of bear, wolf, and even lynx and jackal attacks on humans, although none of these attacks resulted in deaths. Such conflict is expected to decrease given the number of sheep in the country halved following a 1996
foot-and-mouth disease Foot-and-mouth disease (FMD) or hoof-and-mouth disease (HMD) is an infectious and sometimes fatal viral disease that affects cloven-hoofed animals, including domestic and wild bovids. The virus causes a high fever lasting two to six days, followe ...
outbreak and a resultant drop in the value of sheep products. Bears and wild boars are also known to damage agricultural produce. Some species have cultural relevance. The lynx is depicted on a denar coin. Storks are considered to bring luck in local folklore, especially with regard to child-bearing.


Threats

Data deficiency is a significant issue for assessing environmental threats. For example, knowledge about
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
in lacking. The invasive species that are relatively well studied are generally those that directly affect commercial resources, such as the Colorado potato beetle affecting crops and the
Prussian carp The Prussian carp, silver Prussian carp or Gibel carp (''Carassius gibelio''), is a member of the family Cyprinidae, which includes many other fish, such as the common carp, goldfish, and the smaller minnows. It is a medium-sized cyprinid, and d ...
reducing native fish populations in Lake Dojran. Marsh draining, agricultural
land use Land use involves the management and modification of natural environment or wilderness into built environment such as settlements and semi-natural habitats such as arable fields, pastures, and managed woods. Land use by humans has a long h ...
change, and mining have significantly altered the landscape.
Hydropower Hydropower (from el, ὕδωρ, "water"), also known as water power, is the use of falling or fast-running water to Electricity generation, produce electricity or to power machines. This is achieved by energy transformation, converting the Pot ...
dams have been placed in gorges that host rare and endemic plants. ''Lysimachia thyrsiflora'' became locally extinct due to the creation of
Mavrovo Lake Mavrovo Lake ( mk, Мавровско Езеро, translit=Mavrovsko ezero) is situated in the Mavrovo region, in the Mavrovo and Rostuša municipality of the Republic of North Macedonia, less than 100 km from Skopje. It is an important to ...
. Dams in the Black Drim have blocked eel migration to the
Sargasso Sea The Sargasso Sea () is a region of the Atlantic Ocean bounded by four currents forming an ocean gyre. Unlike all other regions called seas, it has no land boundaries. It is distinguished from other parts of the Atlantic Ocean by its charac ...
. Smaller watercourses are at risk of diversion and depletion due to demand for irrigation.
Water pollution Water pollution (or aquatic pollution) is the contamination of water bodies, usually as a result of human activities, so that it negatively affects its uses. Water bodies include lakes, rivers, oceans, aquifers, reservoirs and groundwater. Water ...
remains an issue. The most polluted river, the Vardar, is polluted by around of waste annually, including 75000 tons of solid particles, 5000 tons of
nitrogen Nitrogen is the chemical element with the symbol N and atomic number 7. Nitrogen is a nonmetal and the lightest member of group 15 of the periodic table, often called the pnictogens. It is a common element in the universe, estimated at se ...
, and 1000 tons of
phosphorus Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ear ...
. Heavy industry contributes to such pollution in both surface and groundwater.
Marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...
es are a habitat under considerable threat, not just from land use changes but from climate change, and lowland marshes may disappear completely from the country. This wetland conversion is threatening local plant species. In addition to water pollution and water flow modification, fish are threatened by illegal fishing and invasive species. On land, many plant species are threatened by over-harvesting. '' Gentiana lutea'' is rare due to over-collection for medical purposes.
Illegal logging Illegal logging is the harvest, transportation, purchase or sale of timber in violation of laws. The harvesting procedure itself may be illegal, including using corrupt means to gain access to forests; extraction without permission, or from a pro ...
is also a threat, reducing the habitat of animals such as lynx. Poaching can be an issue, with traps and poison used to catch animals such as wild boar. Poaching increased after
independence Independence is a condition of a person, nation, country, or state in which residents and population, or some portion thereof, exercise self-government, and usually sovereignty, over its territory. The opposite of independence is the statu ...
, due to perceived high fees for hunting within one of the 249 hunting grounds and a lack of punishment, despite poaching a lynx theoretically being punishable by eight years in jail. Wolf hunting is legal. As they are considered pests, the government pays a bounty of 700 denar per wolf killed. Between 100 and 200 wolves are killed per year, mostly through chance encounters during the hunting of other animals. Brown bears are still sometimes poached, and while hunting them was banned in 1996, their population has not increased significantly since then. Poaching has also caused a decline in the chamois population.


Conservation

Of the 22,500 species that occur in the country, over 800 are considered to be endemic. Some species listed as endemic nonetheless have ranges which may stretch slightly outside North Macedonia's borders. For example, species found only in the three major lakes are considered endemic, despite the lakes being shared with neighbouring countries. North Macedonia contains 33 pan-European habitat types regarded as endangered under the Bern Convention's
Emerald network The Emerald network is a network of Areas of Special Conservation Interest to conserve wild flora and fauna and their natural habitats of Europe, which was launched in 1989 by the Council of Europe as part of its work under the Convention on the ...
. The network covers 35 areas encompassing 29% of the country. The same network includes 167 species that require specific conservation measures: 7 invertebrates, 13 fish, 3 amphibians, 7 reptiles, 115 birds, 17 mammals, and 5 plants. The government of North Macedonia has classified threats to biological diversity into 249 items, 17 of which were considered to have a very high priority. The root causes behind these 17 priority threats include poor policymaking, inconsistent enforcement of laws and regulations, poverty, low public awareness, and
climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
. 18 habitats are thought to be vulnerable to climate change, along with 58 plant species and 224 animal species. 65 bird species are listed under the European Union's Birds Directive Annex I, while 15 migratory bird species are listed under Annex I of the Convention on the Conservation of Migratory Species of Wild Animals. 24 reptiles are listed in the Bern Convention and 25 in the European Union Habitats directive appendices. Eight amphibian species are protected by law, while also falling under the Bern Convention and European Union Habitats directive. 34 fish species have protection under Macedonian law.
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
has identified 24 Important Bird and Biodiversity Areas, while
Plantlife International Plantlife is the international conservation membership charity working to secure a world rich in wild plants and fungi. It is the only UK membership charity dedicated to conserving wild plants and fungi in their natural habitats and helping peo ...
has identified 42 Important Plant Areas. The Vulture Conservation Project has been operated by the Macedonian Ecological Society since 2003. The Balkan Lynx Recovery Programme (or Programme for Balkan Lynx Recovery) began in 2006. From 2007 to 2014, the government allocated between 4 and 9 million denar each year for environmental protection. However, most funding comes from external bodies, such as the Global Environment Facility, the European Union (including pre-accession assistance), and bilateral contributions. Cross-border projects exist with Albania, Bulgaria, and Greece. Species that have increased in number from low bases of local extirpation include ''Gentiana pneumonanthe'', ''Ranunculus lingua'', ''
Salvinia natans ''Salvinia natans'' (commonly known as floating fern, floating watermoss, floating moss, or commercially, water butterfly wings) is an annual floating aquatic fern, which can appear superficially similar to moss. It is found throughout the worl ...
'', '' Nuphar lutea'', and '' Menyanthes trifoliata''.


Legal framework

Environmental protection became part of national law for the first time in 1963, under Article 32 of the Constitution of the Socialist Republic of Macedonia. This was maintained in the 1991 Constitution of the Republic of Macedonia under Article 56. The constitution obliges citizens to promote and protect the environment. Most environmental laws are set at a national level, with
Assembly of North Macedonia The Assembly of the Republic of North Macedonia ( mk, Собрание на Република Северна Македонија, sq, Kuvendi i Republikës së Maqedonisë së Veriut), or the Sobranie ( mk, Собрание, "Assembly"), is t ...
having a commission for transport, communications, and environment. The Ministry of Environment and Physical Planning (MoEPP) handles government implementation of laws and regulations. In 2007, the Administration of Environment was established within the MoEPP. In 2014 the State Inspectorate of Environment and Nature became a separate legal entity within the MoEPP. The also plays a role in ensuring the environment is utilised sustainably. The country is working to integrate European Union environmental legislation into its national laws, and to meet the Aichi targets. IUCN Red List classification and methodology is specifically referenced by the Law on Nature Protection of the Republic of North Macedonia ("Official Gazette" no. 67/04, as amended). While the MoEPP is responsible for nature protecting and monitoring, natural resources, such as forests, waters, and
game animals Game or quarry is any wild animal hunted for animal products (primarily meat), for recreation (" sporting"), or for trophies. The species of animals hunted as game varies in different parts of the world and by different local jurisdictions, th ...
remain under the jurisdiction of other ministries. This overlap in jurisdiction has complicated ecosystem management, and the implementation of the European Union Acquis communautaire into North Macedonian Law. Different aspects of environmental monitoring are carried out by a number of different ministries. However, the development of a biodiversity strategy was an early example of effective cooperation between ministries. The Law on Protection of Ohrid, Prespa and Lake Dojrans provided specific protection for the three large lakes starting in 1977. The Law on Environment and Nature Protection and Promotion was instituted in 1996. The Law on Nature Protection was established in 2004 to consolidate previous laws, such as the Law on Natural Rarities (1973) and the Law on Protection of Ohrid, Prespa and Lake Dojrans (1977), and the Law on Protection of National Parks (1980). It also transposes parts of the Acquis communautaire, namely the Habitats Directive, the Birds Directive, and the Regulation on the Protection of Species of Wild Fauna and Flora by Regulating Trade Therein. The use of natural resources such as wild plants and animal parts is regulated under the Law on Nature Protection. National parks issue permits for the collection of wild resources from their forests. This law also legislates to protect the environment from invasive species. Forests are regulated under the Spatial Plan of the Republic of Macedonia (2004), the Strategy for Sustainable Forestry Development in the Republic of Macedonia (2006), the Law on Reproductive Material of Forest Tree Species (2007), and the Law on Forests (2009). Species important for agriculture have their own protection under Article 78 of the Law on Agriculture and Rural Development. 2666 samples from 89 species were stored in a dedicated Gene Bank at the Institute of Agriculture as of 2013. This seed bank is regulated under the Law on Seeds and Seedlings. The government is considering establishing a gene bank for native flora, after a previous attempt at Botanical Garden of the Institute of Biology at the Faculty of Natural Sciences and Mathematics, Ss. Cyril and Methodius University of Skopje, ended after one year. The Law on Waters transfers the European Union
Water Framework Directive The Water Framework Directive 2000/60/EC is an EU directive which commits European Union member states to achieve good qualitative and quantitative status of all water bodies (including marine waters up to one nautical mile from shore) by 2015. ...
into national law, and directs the management of waters, shorelands, and wetlands. Other laws relevant to water management include the Law on Water Management Companies and the Law on Water Communities. The government has published a National Strategy for Waters to guide its actions, and is preparing specific action plans for various lakes and river systems.
Commercial fishing Commercial fishing is the activity of catching fish and other seafood for commercial profit, mostly from wild fisheries. It provides a large quantity of food to many countries around the world, but those who practice it as an industry must often ...
is regulated under the Law on Fishery and Aquaculture. A transboundary plan for fisheries management in Lake Prespa is under development. A joint management plan is being put in place for Lake Ohrid. The first National Biodiversity Strategy and Action Plan was adopted in 2004, alongside the Law on Nature Protection. Only 56% of its proposed actions were implemented. The 2018–2023 National Biodiversity Strategy and Action Plan was adopted on 13 March 2020 during the government's 58th session, and contains 19 specific targets. Another current plan is the National Strategy for Nature Protection for 2017–2027. The first National Environment Action Plan was put in place in 2006. It and the subsequent Second National Environmental Action Plans direct the government's anti-
pollution Pollution is the introduction of contaminants into the natural environment that cause adverse change. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the ...
efforts.
Pelister National Park Pelister National Park is a national park in the Municipality of Bitola, North Macedonia. The park is located in the Baba Mountain massif and covers an area of . The altitude of the park varies between above sea level and is filled with exqu ...
, established in 1948, was the first
national park A national park is a nature park, natural park in use for conservation (ethic), conservation purposes, created and protected by national governments. Often it is a reserve of natural, semi-natural, or developed land that a sovereign state dec ...
in
Yugoslavia Yugoslavia (; sh-Latn-Cyrl, separator=" / ", Jugoslavija, Југославија ; sl, Jugoslavija ; mk, Југославија ;; rup, Iugoslavia; hu, Jugoszlávia; rue, label=Pannonian Rusyn, Югославия, translit=Juhoslavija ...
. Mavrovo National Park followed in 1949, and Mount Galičica became the in 1958. The Law on Protection of National Parks was implemented in 1980. Other protected areas include "National Monuments" (
Smolare Falls Smolare Falls ( Macedonian: Смоларски Водопад) is the tallest waterfall in the Republic of North Macedonia, falling from a height of 39.5 metres (129.5 feet). It is located above the village of Smolare in the Municipality of ...
, Markovi Kuli,
Kuklica Kuklica ( mk, Куклица) is a small village in the municipality of Kratovo, North Macedonia North Macedonia, ; sq, Maqedonia e Veriut, (Macedonia before February 2019), officially the Republic of North Macedonia,, is a country ...
, Lovki-Golemo, Slatinski Izvor cave, Lake Prespa, Lake Dojran, and the Vevchani Springs), "Strict Nature Reserves" (only Ploche Litotelmi), "Nature Parks" ( Ezerani at Lake Prespa), and "Natural Rarities" ( Dona Duka Cave). Individual species can also be declared natural rarities, and this status has been granted to the
sycamore tree Sycamore is a name which has been applied to several types of trees, but with somewhat similar leaf forms. The name derives from the ancient Greek ' (''sūkomoros'') meaning "fig-mulberry". Species of trees known as sycamore: * ''Acer pseudoplata ...
. Protected areas have been established in an ad hoc manner, and for a variety of purposes, resulting in no coherent network or strategy. The Law on Nature Protection divided them into six categories. Altogether, the 86 protected sites cover 230,083 ha, or 8.9% of the country (4.5% national parks, 3.0% monuments, 1.4% other protected areas). While these national parks have established management systems, other protected areas lack specific plans and dedicated scientific studies. A fourth park, Shar Mountain National Park, was created in 2021. There are two classes of protected species under North Macedonian law, "Strictly Protected" and "Protected". National protection measures exist for "Strictly Protected" species, but not yet for species identified as "Protected". Legislation has been enacted to control the trade of wildlife and wildlife parts. The Constitution treats game animals as
common goods Common goods (also called common-pool resources) are defined in economics as goods that are rivalrous and non-excludable. Thus, they constitute one of the four main types based on the criteria: * whether the consumption of a good by one pers ...
, which are given special protection. Current regulation is established under the 2012 Law on Hunting. 133 wild species are classified as game: 110 birds and 23 mammals. There are 112 hunting grounds for large game, and 144 for small game. These are all divided between 11 hunting management areas under the General Hunting Management Master Plan. The most hunted species include hares, partridges, pheasants, wild boar, and chamois. Previously unprotected, bears were classified as a game species in 1988, gaining hunting law protections in 1996. 762 wolves, 76 deer, 521 chamois, over 6000 wild boar, over 32,000 rabbits, 252 birds of prey, and 1500 waterfowl were recorded as having been shot between 2003 and 2012. A few hunted species have seen steep reductions in their population. Some deer populations have been reduced by up to 93%. The legal list of protected and strictly protected species has not kept up to date with taxonomy, for example by double-counting species which had previously been assigned multiple names.


International conventions

North Macedonia ratified the Convention on Biological Diversity in 1997, the
United Nations Framework Convention on Climate Change The United Nations Framework Convention on Climate Change (UNFCCC) established an international environmental treaty to combat "dangerous human interference with the climate system", in part by stabilizing greenhouse gas concentrations in th ...
in 1997, the United Nations Convention to Combat Desertification in 2002, and the
Kyoto Protocol The Kyoto Protocol was an international treaty which extended the 1992 United Nations Framework Convention on Climate Change (UNFCCC) that commits state parties to reduce greenhouse gas emissions, based on the scientific consensus that (part ...
in 2004. North Macedonia is also a party to the Convention on the Conservation of Migratory Species of Wild Animals, along with the specific Agreement on the Conservation of Populations of European Bats and Agreement on the Conservation of African-Eurasian Migratory Waterbirds. Two areas totalling have been designated wetlands of international importance under the
Ramsar Convention The Ramsar Convention on Wetlands of International Importance Especially as Waterfowl Habitat is an international treaty for the conservation and sustainable use of Ramsar sites (wetlands). It is also known as the Convention on Wetlands. It i ...
: Lake Prespa in 1995 and Dorjan Lake in 2007. However, there is currently no national programme for wetlands conservation, despite obligations under the convention. Lake Ohrid was designated a UNESCO
World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for h ...
under the criteria for nature in 1979. Slatinski Izvor cave and the Markovi Kuli landscape were tentatively enrolled in 2004.


References


External links


Balkan lynx recovery programme

Biodiversity Strategy and Action Plan of the Republic of Macedonia (2004)
{{DEFAULTSORT:Wildlife Of North Macedonia Biota of North Macedonia
North Macedonia North Macedonia, ; sq, Maqedonia e Veriut, (Macedonia before February 2019), officially the Republic of North Macedonia,, is a country in Southeast Europe. It gained independence in 1991 as one of the successor states of Socialist Feder ...