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The walrus (''Odobenus rosmarus'') is a large flippered
marine mammal Marine mammals are aquatic mammals that rely on the ocean and other marine ecosystems for their existence. They include animals such as seals, whales, manatees, sea otters and polar bears. They are an informal group, unified only by their reli ...
with a discontinuous distribution about the
North Pole The North Pole, also known as the Geographic North Pole or Terrestrial North Pole, is the point in the Northern Hemisphere where the Earth's axis of rotation meets its surface. It is called the True North Pole to distinguish from the Mag ...
in the
Arctic Ocean The Arctic Ocean is the smallest and shallowest of the world's five major oceans. It spans an area of approximately and is known as the coldest of all the oceans. The International Hydrographic Organization (IHO) recognizes it as an ocean, a ...
and
subarctic The subarctic zone is a region in the Northern Hemisphere immediately south of the true Arctic, north of humid continental regions and covering much of Alaska, Canada, Iceland, the north of Scandinavia, Siberia, and the Cairngorms. Generally, ...
seas of the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
. The walrus is the only living species in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Odobenidae Odobenidae is a family of pinnipeds. The only living species is the walrus (''Odobenus rosmarus''). In the past, however, the group was much more diverse, and includes more than a dozen fossil genera. Taxonomy All genera, except ''Odobenus'', ...
and
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Odobenus''. This species is subdivided into two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
: the Atlantic walrus (''O. r. rosmarus''), which lives in the
Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
, and the Pacific walrus (''O. r. divergens''), which lives in the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
. Adult walrus are characterised by prominent
tusk Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with pigs and walruses, or, in the case of elephants, elongated incisors. Tusks share c ...
s and
whiskers Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to touch, sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are c ...
, and their considerable bulk: adult males in the Pacific can weigh more than and, among
pinniped Pinnipeds (pronounced ), commonly known as seals, are a widely distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant families Odobenidae (whose only living member is the walru ...
s, are exceeded in size only by the two species of
elephant seal Elephant seals are very large, oceangoing earless seals in the genus ''Mirounga''. Both species, the northern elephant seal (''M. angustirostris'') and the southern elephant seal (''M. leonina''), were hunted to the brink of extinction for oil ...
s. Walruses live mostly in shallow waters above the
continental shelves A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...
, spending significant amounts of their lives on the
sea ice Sea ice arises as seawater freezes. Because ice is less dense than water, it floats on the ocean's surface (as does fresh water ice, which has an even lower density). Sea ice covers about 7% of the Earth's surface and about 12% of the world's oce ...
looking for
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
bivalve mollusks to eat. Walruses are relatively long-lived, social animals, and they are considered to be a "
keystone species A keystone species is a species which has a disproportionately large effect on its natural environment relative to its abundance, a concept introduced in 1969 by the zoologist Robert T. Paine. Keystone species play a critical role in maintaini ...
" in the Arctic marine regions. The walrus has played a prominent role in the cultures of many
indigenous Arctic peoples Circumpolar peoples and Arctic peoples are umbrella terms for the various Indigenous peoples of the Arctic. Prehistory The earliest inhabitants of North America's central and eastern Arctic are referred to as the Arctic small tool tradition (AST) ...
, who have hunted the walrus for its meat, fat, skin, tusks, and bone. During the 19th century and the early 20th century, walruses were widely hunted and killed for their
blubber Blubber is a thick layer of vascularized adipose tissue under the skin of all cetaceans, pinnipeds, penguins, and sirenians. Description Lipid-rich, collagen fiber-laced blubber comprises the hypodermis and covers the whole body, except for pa ...
,
walrus ivory The walrus (''Odobenus rosmarus'') is a large flippered marine mammal with a discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. The walrus is the only living species in the fami ...
, and meat. The population of walruses dropped rapidly all around the Arctic region. Their population has rebounded somewhat since then, though the populations of Atlantic and Laptev walruses remain fragmented and at low levels compared with the time before human interference.


Etymology

The origin of the word ''walrus'' derives from a
Germanic language The Germanic languages are a branch of the Indo-European language family spoken natively by a population of about 515 million people mainly in Europe, North America, Oceania and Southern Africa. The most widely spoken Germanic language, Engli ...
, and it has been attributed largely to either the
Dutch language Dutch ( ) is a West Germanic language spoken by about 25 million people as a first language and 5 million as a second language. It is the third most widely spoken Germanic language The Germanic languages are a branch of the Indo-Europea ...
or
Old Norse Old Norse, Old Nordic, or Old Scandinavian, is a stage of development of North Germanic languages, North Germanic dialects before their final divergence into separate Nordic languages. Old Norse was spoken by inhabitants of Scandinavia and t ...
. Its first part is thought to derive from a word such as Old Norse ('whale') and the second part has been hypothesized to come from the Old Norse word ('horse'). For example, the
Old Norse Old Norse, Old Nordic, or Old Scandinavian, is a stage of development of North Germanic languages, North Germanic dialects before their final divergence into separate Nordic languages. Old Norse was spoken by inhabitants of Scandinavia and t ...
word means 'horse-whale' and is thought to have been passed in an inverted form to both Dutch and the dialects of northern Germany as and . An alternative theory is that it comes from the Dutch words 'shore' and 'giant'. The species name ''rosmarus'' is Scandinavian. The Norwegian manuscript ''
Konungs skuggsjá ''Konungs skuggsjá'' (Old Norse for "King's mirror"; Latin: ''Speculum regale'', modern Norwegian: ''Kongsspegelen'' (Nynorsk) or ''Kongespeilet'' (Bokmål)) is a Norwegian didactic text in Old Norse from around 1250, an example of speculum lite ...
'', thought to date from around AD 1240, refers to the walrus as in Iceland and in Greenland (walruses were by now extinct in Iceland and Norway, while the word evolved in Greenland). Several place names in Iceland, Greenland and Norway may originate from walrus sites: Hvalfjord, Hvallatrar and Hvalsnes to name some, all being typical walrus breeding grounds. The archaic English word for walrus—''morse''—is widely thought to have come from the
Slavic languages The Slavic languages, also known as the Slavonic languages, are Indo-European languages spoken primarily by the Slavic peoples and their descendants. They are thought to descend from a proto-language called Proto-Slavic, spoken during the Ear ...
, which in turn borrowed it from Finno-Ugric languages, and ultimately (according to
Ante Aikio Ante Aikio (Sámi: Luobbal Sámmol Sámmol Ánte; born 1977) is a Finnish linguist of Sámi origin who has been a professor of Sámi languages at the Sámi University of Applied Sciences in Kautokeino, Norway since 2015. Prior to this he has been a ...
) from an unknown
Pre-Finno-Ugric substrate Pre-Finno-Ugric substrate refers to substratum loanwords from unidentified non-Indo-European and non-Uralic languages that are found in various Finno-Ugric languages, most notably Sami. The presence of Pre-Finno-Ugric substrate in Sami language ...
language of Northern Europe. Compare () in Russian, in
Finnish Finnish may refer to: * Something or someone from, or related to Finland * Culture of Finland * Finnish people or Finns, the primary ethnic group in Finland * Finnish language, the national language of the Finnish people * Finnish cuisine See also ...
, in Northern Saami, and in French.
Olaus Magnus Olaus Magnus (October 1490 – 1 August 1557) was a Swedish writer, cartographer, and Catholic ecclesiastic. Biography Olaus Magnus (a Latin translation of his birth name Olof Månsson) was born in Linköping in October 1490. Like his elder ...
, who depicted the walrus in the ''
Carta Marina ''Carta marina et descriptio septentrionalium terrarum'' (Latin for ''Marine map and description of the Northern lands''; commonly abbreviated ''Carta marina'') is the first map of the Nordic countries to give details and place names, created by S ...
'' in 1539, first referred to the walrus as the , probably a Latinization of , and this was adopted by
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in his
binomial nomenclature In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
. The coincidental similarity between ''morse'' and the Latin word ('a bite') supposedly contributed to the walrus's reputation as a "terrible monster". The compound ''Odobenus'' comes from ' (
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
for 'teeth') and ' (Greek for 'walk'), based on observations of walruses using their tusks to pull themselves out of the water. The term in
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
means 'turning apart', referring to their tusks.


Taxonomy and evolution

The walrus is a mammal in the order
Carnivora Carnivora is a Clade, monophyletic order of Placentalia, placental mammals consisting of the most recent common ancestor of all felidae, cat-like and canidae, dog-like animals, and all descendants of that ancestor. Members of this group are f ...
. It is the sole surviving member of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Odobenidae, one of three lineages in the
suborder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...
Pinnipedia Pinnipeds (pronounced ), commonly known as seals, are a widely distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant families Odobenidae (whose only living member is the wal ...
along with true seals ( Phocidae) and eared seals ( Otariidae). While there has been some debate as to whether all three lineages are
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
, i.e. descended from a single ancestor, or diphyletic, recent genetic evidence suggests all three descended from a
caniform Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include dogs (wolves, foxes, etc.), bears, raccoons, and mustelids. The Pinnipedia (seals, walruses and sea lions) are also assigned to this group. ...
ancestor most closely related to modern bears. Recent multigene analysis indicates the odobenids and otariids diverged from the phocids about 20–26 million years ago, while the odobenids and the otariids separated 15–20 million years ago. Odobenidae was once a highly diverse and widespread family, including at least twenty species in the subfamilies Imagotariinae, Dusignathinae and Odobeninae. The key distinguishing feature was the development of a squirt/suction feeding mechanism; tusks are a later feature specific to Odobeninae, of which the modern walrus is the last remaining (
relict A relict is a surviving remnant of a natural phenomenon. Biology A relict (or relic) is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas. Geology and geomorphology In geology, a r ...
) species. Two subspecies of walrus are widely recognized: the Atlantic walrus, ''O. r. rosmarus'' (Linnaeus, 1758) and the Pacific walrus, ''O. r. divergens'' (Illiger, 1815). Fixed genetic differences between the Atlantic and Pacific subspecies indicate very restricted gene flow, but relatively recent separation, estimated at 500,000 and 785,000 years ago. These dates coincide with the hypothesis derived from fossils that the walrus evolved from a tropical or subtropical ancestor that became isolated in the Atlantic Ocean and gradually adapted to colder conditions in the Arctic. From there, it presumably recolonized the
North Pacific Ocean North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating direction or geography. Etymology The word ''north'' is ...
during high glaciation periods in the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
via the Central American Seaway. An isolated population in the
Laptev Sea The Laptev Sea ( rus, мо́ре Ла́птевых, r=more Laptevykh; sah, Лаптевтар байҕаллара, translit=Laptevtar baỹğallara) is a marginal sea of the Arctic Ocean. It is located between the northern coast of Siberia, th ...
was considered by some authorities, including many Russian biologists and the canonical ''Mammal Species of the World'', to be a third subspecies, ''O. r. laptevi'' (Chapskii, 1940), but has since been determined to be of Pacific walrus origin.


Anatomy

While some outsized Pacific males can weigh as much as , most weigh between . An occasional male of the Pacific subspecies far exceeds normal dimensions. In 1909, a walrus hide weighing was collected from an enormous bull in
Franz Josef Land , native_name = , image_name = Map of Franz Josef Land-en.svg , image_caption = Map of Franz Josef Land , image_size = , map_image = Franz Josef Land location-en.svg , map_caption = Location of Franz Josef ...
, while in August 1910, Jack Woodson shot a walrus, harvesting its hide. Since a walrus's hide usually accounts for about 20% of its body weight, the total body mass of these two giants is estimated to have been at least . The Atlantic subspecies weighs about 10–20% less than the Pacific subspecies. Male Atlantic walrus weigh an average of . The Atlantic walrus also tends to have relatively shorter tusks and somewhat more flattened
snout A snout is the protruding portion of an animal's face, consisting of its nose, mouth, and jaw. In many animals, the structure is called a muzzle, rostrum, or proboscis. The wet furless surface around the nostrils of the nose of many mammals is c ...
. Females weigh about two-thirds as much as males, with the Atlantic females averaging , sometimes weighing as little as , and the Pacific female averaging . Length typically ranges from . Newborn walruses are already quite large, averaging in weight and in length across both sexes and subspecies. All told, the walrus is the third largest pinniped species, after the two
elephant seals Elephant seals are very large, oceangoing earless seals in the genus ''Mirounga''. Both species, the northern elephant seal (''M. angustirostris'') and the southern elephant seal (''M. leonina''), were hunted to the brink of extinction for oil ...
. Walruses maintain such a high body weight because of the blubber stored underneath their skin. This blubber keeps them warm and the fat provides energy to the walrus. The walrus's body shape shares features with both sea lions (
eared seal An eared seal or otariid or otary is any member of the marine mammal family Otariidae, one of three groupings of pinnipeds. They comprise 15 extant species in seven genera (another species became extinct in the 1950s) and are commonly known eith ...
s: Otariidae) and seals (
true seals The earless seals, phocids or true seals are one of the three main groups of mammals within the seal lineage, Pinnipedia. All true seals are members of the family Phocidae (). They are sometimes called crawling seals to distinguish them from th ...
: Phocidae). As with otariids, it can turn its rear flippers forward and move on all fours; however, its swimming technique is more like that of true seals, relying less on flippers and more on sinuous whole body movements. Also like phocids, it lacks external ears. The
extraocular muscles The extraocular muscles (extrinsic ocular muscles), are the seven extrinsic muscles of the human eye. Six of the extraocular muscles, the four recti muscles, and the superior and inferior oblique muscles, control movement of the eye and the ot ...
of the walrus are well-developed. This and its lack of orbital roof allow it to protrude its eyes and see in both a frontal and dorsal direction. However, vision in this species appears to be more suited for short-range.


Tusks and dentition

While this was not true of all extinct walruses, the most prominent feature of the living species is its long tusks. These are elongated
canines Canine may refer to: Zoology and anatomy * a dog-like Canid animal in the subfamily Caninae ** ''Canis'', a genus including dogs, wolves, coyotes, and jackals ** Dog, the domestic dog * Canine tooth, in mammalian oral anatomy People with the surn ...
, which are present in both male and female walruses and can reach a length of 1 m (3 ft 3 in) and weigh up to 5.4 kg (12 lb). Tusks are slightly longer and thicker among males, which use them for fighting, dominance and display; the strongest males with the largest tusks typically dominate social groups. Tusks are also used to form and maintain holes in the ice and aid the walrus in climbing out of water onto ice. Tusks were once thought to be used to dig out prey from the seabed, but analyses of abrasion patterns on the tusks indicate they are dragged through the sediment while the upper edge of the snout is used for digging. While the
dentition Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ...
of walruses is highly variable, they generally have relatively few teeth other than the tusks. The maximal number of teeth is 38 with dentition formula: , but over half of the teeth are rudimentary and occur with less than 50% frequency, such that a typical dentition includes only 18 teeth


Vibrissae (whiskers)

Surrounding the tusks is a broad mat of stiff bristles ("mystacial
vibrissae Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are coarser ...
"), giving the walrus a characteristic whiskered appearance. There can be 400 to 700 vibrissae in 13 to 15 rows reaching 30 cm (12 in) in length, though in the wild they are often worn to much shorter lengths due to constant use in foraging. The vibrissae are attached to muscles and are supplied with blood and nerves, making them highly sensitive organs capable of differentiating shapes thick and wide.


Skin

Aside from the vibrissae, the walrus is sparsely covered with fur and appears bald. Its skin is highly wrinkled and thick, up to around the neck and shoulders of males. The blubber layer beneath is up to thick. Young walruses are deep brown and grow paler and more cinnamon-colored as they age. Old males, in particular, become nearly pink. Because skin blood vessels constrict in cold water, the walrus can appear almost white when swimming. As a
secondary sexual characteristic Secondary sex characteristics are features that appear during puberty in humans, and at sexual maturity in other animals. These characteristics are particularly evident in the sexual dimorphism, sexually dimorphic phenotypic traits that distinguis ...
, males also acquire significant nodules, called "bosses", particularly around the neck and shoulders. The walrus has an air sac under its throat which acts like a flotation bubble and allows it to bob vertically in the water and sleep. The males possess a large baculum (penis bone), up to in length, the largest of any land mammal, both in absolute size and relative to body size.


Life history


Reproduction

Walruses live to about 20–30 years old in the wild. The males reach sexual maturity as early as seven years, but do not typically mate until fully developed at around 15 years of age. They rut from January through April, decreasing their food intake dramatically. The females begin ovulating as soon as four to six years old. The females are diestrous, coming into heat in late summer and also around February, yet the males are fertile only around February; the potential fertility of this second period is unknown. Breeding occurs from January to March, peaking in February. Males aggregate in the water around ice-bound groups of estrous females and engage in competitive vocal displays. The females join them and copulate in the water.
Gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregna ...
lasts 15 to 16 months. The first three to four months are spent with the
blastula Blastulation is the stage in early animal embryonic development that produces the blastula. In mammalian development the blastula develops into the blastocyst with a differentiated inner cell mass and an outer trophectoderm. The blastula (from ...
in suspended development before it implants itself in the uterus. This strategy of delayed implantation, common among pinnipeds, presumably evolved to optimize both the mating season and the birthing season, determined by ecological conditions that promote newborn survival. Calves are born during the spring migration, from April to June. They weigh at birth and are able to swim. The mothers nurse for over a year before weaning, but the young can spend up to five years with the mothers. Walrus milk contains higher amounts of fats and protein compared to land animals but lower compared to phocid seals. This lower fat content in turn causes a slower growth rate among calves and a longer nursing investment for their mothers. Because ovulation is suppressed until the calf is weaned, females give birth at most every two years, leaving the walrus with the lowest reproductive rate of any pinniped.


Migration

The rest of the year (late summer and fall), walruses tend to form massive aggregations of tens of thousands of individuals on rocky beaches or outcrops. The migration between the ice and the beach can be long-distance and dramatic. In late spring and summer, for example, several hundred thousand Pacific walruses migrate from the
Bering Sea The Bering Sea (, ; rus, Бе́рингово мо́ре, r=Béringovo móre) is a marginal sea of the Northern Pacific Ocean. It forms, along with the Bering Strait, the divide between the two largest landmasses on Earth: Eurasia and The Ameri ...
into the
Chukchi Sea Chukchi Sea ( rus, Чуко́тское мо́ре, r=Chukotskoye more, p=tɕʊˈkotskəjə ˈmorʲɪ), sometimes referred to as the Chuuk Sea, Chukotsk Sea or the Sea of Chukotsk, is a marginal sea of the Arctic Ocean. It is bounded on the west b ...
through the relatively narrow Bering Strait.


Ecology


Range and habitat

The majority of the population of the Pacific walrus spends its summers north of the Bering Strait in the
Chukchi Sea Chukchi Sea ( rus, Чуко́тское мо́ре, r=Chukotskoye more, p=tɕʊˈkotskəjə ˈmorʲɪ), sometimes referred to as the Chuuk Sea, Chukotsk Sea or the Sea of Chukotsk, is a marginal sea of the Arctic Ocean. It is bounded on the west b ...
of the Arctic Ocean along the northern coast of eastern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
, around
Wrangel Island Wrangel Island ( rus, О́стров Вра́нгеля, r=Ostrov Vrangelya, p=ˈostrəf ˈvrangʲɪlʲə; ckt, Умӄиԓир, translit=Umqiḷir) is an island of the Chukotka Autonomous Okrug, Russia. It is the 91st largest island in the w ...
, in the
Beaufort Sea The Beaufort Sea (; french: Mer de Beaufort, Iñupiaq: ''Taġiuq'') is a marginal sea of the Arctic Ocean, located north of the Northwest Territories, the Yukon, and Alaska, and west of Canada's Arctic islands. The sea is named after Sir Fr ...
along the northern shore of
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
south to
Unimak Island Unimak Island ( ale, Unimax, russian: Унимак) is the largest island in the Aleutian Islands chain of the U.S. state of Alaska. Geography It is the easternmost island in the Aleutians and, with an area of , the ninth largest island in the U ...
, and in the waters between those locations. Smaller numbers of males summer in the
Gulf of Anadyr The Gulf of Anadyr, or Anadyr Bay (russian: Анадырский залив), is a large bay on the Bering Sea in far northeast Siberia. It has a total surface area of Location The bay is roughly rectangular and opens to the southeast. The corn ...
on the southern coast of the Siberian
Chukchi Peninsula The Chukchi Peninsula (also Chukotka Peninsula or Chukotski Peninsula; russian: Чуко́тский полуо́стров, ''Chukotskiy poluostrov'', short form russian: Чуко́тка, ''Chukotka''), at about 66° N 172° W, is the eastern ...
, and in Bristol Bay off the southern coast of Alaska, west of the
Alaska Peninsula The Alaska Peninsula (also called Aleut Peninsula or Aleutian Peninsula, ale, Alasxix̂; Sugpiaq: ''Aluuwiq'', ''Al'uwiq'') is a peninsula extending about to the southwest from the mainland of Alaska and ending in the Aleutian Islands. The ...
. In the spring and fall, walruses congregate throughout the Bering Strait, reaching from the western coast of Alaska to the Gulf of Anadyr. They winter over in the
Bering Sea The Bering Sea (, ; rus, Бе́рингово мо́ре, r=Béringovo móre) is a marginal sea of the Northern Pacific Ocean. It forms, along with the Bering Strait, the divide between the two largest landmasses on Earth: Eurasia and The Ameri ...
along the eastern coast of Siberia south to the northern part of the
Kamchatka Peninsula The Kamchatka Peninsula (russian: полуостров Камчатка, Poluostrov Kamchatka, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and we ...
, and along the southern coast of Alaska. A 28,000-year-old fossil walrus was dredged up from the bottom of
San Francisco Bay San Francisco Bay is a large tidal estuary in the U.S. state of California, and gives its name to the San Francisco Bay Area. It is dominated by the big cities of San Francisco, San Jose, and Oakland. San Francisco Bay drains water from a ...
, indicating that Pacific walruses ranged that far south during the last Ice Age. Commercial harvesting reduced the population of the Pacific walrus to between 50,000 and 100,000 in the 1950s-1960s. Limits on commercial hunting allowed the population to increase to a peak in the 1970s-1980s, but subsequently, walrus numbers have again declined. Early aerial censuses of Pacific walrus conducted at five-year intervals between 1975 and 1985 estimated populations of above 220,000 in each of the three surveys. In 2006, the population of the Pacific walrus was estimated to be around 129,000 on the basis of an aerial census combined with satellite tracking. There were roughly 200,000 Pacific walruses in 1990. The much smaller population of Atlantic walruses ranges from the Canadian Arctic, across
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...
,
Svalbard Svalbard ( , ), also known as Spitsbergen, or Spitzbergen, is a Norwegian archipelago in the Arctic Ocean. North of mainland Europe, it is about midway between the northern coast of Norway and the North Pole. The islands of the group range ...
, and the western part of Arctic Russia. There are eight hypothetical subpopulations of Atlantic walruses, based largely on their geographical distribution and movements: five west of Greenland and three east of Greenland. The Atlantic walrus once ranged south to
Sable Island Sable Island (french: île de Sable, literally "island of sand") is a small Canadian island situated southeast of Halifax, Nova Scotia, and about southeast of the closest point of mainland Nova Scotia in the North Atlantic Ocean. The island i ...
,
Nova Scotia Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. Nova Scotia is Latin for "New Scotland". Most of the population are native Eng ...
, and as late as the 18th century was found in large numbers in the Greater Gulf of St. Lawrence region, sometimes in colonies of up to 7,000 to 8,000 individuals. This population was nearly eradicated by commercial harvest; their current numbers, though difficult to estimate, probably remain below 20,000. In April 2006, the Canadian Species at Risk Act listed the population of the northwestern Atlantic walrus in
Quebec Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirtee ...
, New Brunswick,
Nova Scotia Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. Nova Scotia is Latin for "New Scotland". Most of the population are native Eng ...
, Newfoundland and Labrador as having been eradicated in Canada. A genetically distinct population existed in Iceland that was wiped out after Norse settlement around 1213–1330 AD. The isolated population of Laptev Sea walruses is confined year-round to the central and western regions of the
Laptev Sea The Laptev Sea ( rus, мо́ре Ла́птевых, r=more Laptevykh; sah, Лаптевтар байҕаллара, translit=Laptevtar baỹğallara) is a marginal sea of the Arctic Ocean. It is located between the northern coast of Siberia, th ...
, the eastmost regions of the
Kara Sea The Kara Sea (russian: Ка́рское мо́ре, ''Karskoye more'') is a marginal sea, separated from the Barents Sea to the west by the Kara Strait and Novaya Zemlya, and from the Laptev Sea to the east by the Severnaya Zemlya archipelago. ...
, and the westmost regions of the
East Siberian Sea The East Siberian Sea ( rus, Восто́чно-Сиби́рское мо́ре, r=Vostochno-Sibirskoye more) is a marginal sea in the Arctic Ocean. It is located between the Arctic Cape to the north, the coast of Siberia to the south, the New Si ...
. The current population of these walruses has been estimated to be between 5,000 and 10,000. Even though walruses can dive to depths beyond 500 meters, they spend most of their time in shallow waters (and the nearby ice floes) hunting for food. In March 2021, a single walrus, nicknamed
Wally the Walrus Wally the Walrus, also known as Wally the Wandering Walrus, is a male arctic walrus who attracted much media attention for appearing, and hauling out, during 2021 in several locations across the coast of western Europe, mainly Ireland and Brit ...
, was sighted at
Valentia Island Valentia Island () is one of Ireland's most westerly points. It lies off the Iveragh Peninsula in the southwest of County Kerry. It is linked to the mainland by the Maurice O'Neill Memorial Bridge at Portmagee. A car ferry also departs from R ...
,
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
, far south of its typical range, potentially due to having fallen asleep on an iceberg that then drifted south towards Ireland. Days later, a walrus, thought to be the same animal, was spotted on the
Pembrokeshire Pembrokeshire ( ; cy, Sir Benfro ) is a Local government in Wales#Principal areas, county in the South West Wales, south-west of Wales. It is bordered by Carmarthenshire to the east, Ceredigion to the northeast, and the rest by sea. The count ...
coast,
Wales Wales ( cy, Cymru ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by England to the Wales–England border, east, the Irish Sea to the north and west, the Celtic Sea to the south west and the ...
. In June 2022, a single walrus was sighted on the shores of the
Baltic Sea The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden and the North and Central European Plain. The sea stretches from 53°N to 66°N latitude and from ...
- at
Rügen Rügen (; la, Rugia, ) is Germany's largest island. It is located off the Pomeranian coast in the Baltic Sea and belongs to the state of Mecklenburg-Western Pomerania. The "gateway" to Rügen island is the Hanseatic city of Stralsund, where ...
Island,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
,
Mielno Mielno (german: Groß Möllen or ) is a resort town in Koszalin County, West Pomeranian Voivodeship, in north-western Poland. It is the seat of the gmina (administrative district) called Gmina Mielno. It lies approximately north-west of Koszali ...
,
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...
and
Skälder Bay Skälder Bay ( sv, Skälderviken) is a bay in Skåne County, Sweden. It is an inlet of the Kattegat on the western coast of Scania, located between the Bjäre Peninsula to the north (which separates it from Laholm Bay) and the Kullen Peninsula ...
,
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
. In July 2022, there was a report of a lost, starving walrus (nicknamed as Stena) in the coastal waters of the towns of
Hamina Hamina (; sv, Fredrikshamn, , Sweden ) is a List of cities in Finland, town and a Municipalities of Finland, municipality of Finland. It is located approximately east of the country's capital Helsinki, in the Kymenlaakso Regions of Finland, reg ...
and
Kotka Kotka (; ; la, Aquilopolis) is a city in the southern part of the Kymenlaakso province on the Gulf of Finland. Kotka is a major port and industrial city and also a diverse school and cultural city, which was formerly part of the old Kymi parish. ...
in
Kymenlaakso Kymenlaakso ( sv, Kymmenedalen; " Kymi/Kymmene Valley") is a region in Finland. It borders the regions of Uusimaa, Päijät-Häme, South Savo and South Karelia and Russia (Leningrad Oblast). Its name means literally ''The Valley of River Kymi'' ...
,
Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of B ...
, that, despite rescue attempts, died of starvation when the rescuers tried to transport it to the
Korkeasaari Zoo Korkeasaari Zoo ( fi, Korkeasaaren eläintarha, previously known as ''Helsinki Zoo'') is the largest zoo in Finland, located on the island of Korkeasaari in Helsinki. The zoo was first opened in 1889, originally showcasing common animals from Fi ...
for treatment.


Diet

Walruses prefer shallow shelf regions and forage primarily on the sea floor, often from sea ice platforms. They are not particularly deep divers compared to other pinnipeds; the deepest dives in a study of Atlantic walrus near
Svalbard Svalbard ( , ), also known as Spitsbergen, or Spitzbergen, is a Norwegian archipelago in the Arctic Ocean. North of mainland Europe, it is about midway between the northern coast of Norway and the North Pole. The islands of the group range ...
were only 31±17 m (102 ft) but a more recent study recorded dives exceeding 500 m (1640 ft) in
Smith Sound Smith Sound ( da, Smith Sund; french: Détroit de Smith) is an uninhabited Arctic sea passage between Greenland and Canada's northernmost island, Ellesmere Island. It links Baffin Bay with Kane Basin and forms part of the Nares Strait. On the ...
, between NW Greenland and Arctic Canada - in general peak dive depth can be expected to depend on prey distribution and seabed depth. The walrus has a diverse and opportunistic diet, feeding on more than 60 genera of marine organisms, including
shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...
, crabs,
tube worms A tubeworm is any worm-like sessility (zoology), sessile invertebrate that anchors its tail to an underwater surface and secretes around its body a mineral tube, into which it can withdraw its entire body. Tubeworms are found among the following t ...
, soft
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and sec ...
s,
tunicate A tunicate is a marine invertebrate animal, a member of the subphylum Tunicata (). It is part of the Chordata, a phylum which includes all animals with dorsal nerve cords and notochords (including vertebrates). The subphylum was at one time ca ...
s,
sea cucumber Sea cucumbers are echinoderms from the class Holothuroidea (). They are marine animals with a leathery skin and an elongated body containing a single, branched gonad. Sea cucumbers are found on the sea floor worldwide. The number of holothuria ...
s, various
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s (such as
snail A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gastro ...
s,
octopus An octopus ( : octopuses or octopodes, see below for variants) is a soft-bodied, eight- limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttle ...
es, and
squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting t ...
), some types of slow-moving
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
, and even parts of other pinnipeds. However, it prefers benthic bivalve mollusks, especially
clam Clam is a common name for several kinds of bivalve molluscs. The word is often applied only to those that are edible and live as infauna, spending most of their lives halfway buried in the sand of the seafloor or riverbeds. Clams have two she ...
s, for which it forages by grazing along the sea bottom, searching and identifying prey with its sensitive
vibrissae Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are coarser ...
and clearing the murky bottoms with jets of water and active flipper movements. The walrus sucks the meat out by sealing its powerful lips to the organism and withdrawing its piston-like tongue rapidly into its mouth, creating a vacuum. The walrus palate is uniquely vaulted, enabling effective suction. The diet of the Pacific walrus consist almost exclusively of benthic invertebrates (97 percent). Aside from the large numbers of organisms actually consumed by the walrus, its foraging has a large peripheral impact on benthic communities. It disturbs ( bioturbates) the sea floor, releasing nutrients into the water column, encouraging mixing and movement of many organisms and increasing the patchiness of the
benthos Benthos (), also known as benthon, is the community of organisms that live on, in, or near the bottom of a sea, river, lake, or stream, also known as the benthic zone. Seal tissue has been observed in a fairly significant proportion of walrus stomachs in the Pacific, but the importance of seals in the walrus diet is under debate. There have been isolated observations of walruses preying on seals up to the size of a bearded seal. Rarely, incidents of walruses preying on seabirds, particularly the
Brünnich's guillemot The thick-billed murre or Brünnich's guillemot (''Uria lomvia'') is a bird in the auk family (Alcidae). This bird is named after the Danish zoologist Morten Thrane Brünnich. The very deeply black North Pacific subspecies ''Uria lomvia arra'' i ...
(''Uria lomvia''), have been documented. Walruses may occasionally prey on ice-entrapped
narwhal The narwhal, also known as a narwhale (''Monodon monoceros''), is a medium-sized toothed whale that possesses a large "tusk" from a protruding canine tooth. It lives year-round in the Arctic waters around Greenland, Canada and Russia. It is o ...
s and scavenge on whale carcasses but there is little evidence to prove this.


Predators

Due to its great size and tusks, the walrus has only two natural predators: the
orca The orca or killer whale (''Orcinus orca'') is a toothed whale belonging to the oceanic dolphin family, of which it is the largest member. It is the only Extant taxon, extant species in the genus ''Orcinus'' and is recognizable by its black ...
and the
polar bear The polar bear (''Ursus maritimus'') is a hypercarnivorous bear whose native range lies largely within the Arctic Circle, encompassing the Arctic Ocean, its surrounding seas and surrounding land masses. It is the largest extant bear specie ...
. The walrus does not, however, comprise a significant component of either of these predators' diets. Both the orca and the polar bear are also most likely to prey on walrus calves. The polar bear often hunts the walrus by rushing at beached aggregations and consuming the individuals crushed or wounded in the sudden exodus, typically younger or infirm animals. The bears also isolate walruses when they
overwinter Overwintering is the process by which some organisms pass through or wait out the winter season, or pass through that period of the year when "winter" conditions (cold or sub-zero temperatures, ice, snow, limited food supplies) make normal activi ...
and are unable to escape a charging bear due to inaccessible diving holes in the ice. However, even an injured walrus is a formidable opponent for a polar bear, and direct attacks are rare. Armed with its ivory tusks, walruses have been known to fatally injure polar bears in battles if the latter follows the other into the water, where the bear is at a disadvantage. Polar bear–walrus battles are often extremely protracted and exhausting, and bears have been known to break away from the attack after injuring a walrus. Orcas regularly attack walruses, although walruses are believed to have successfully defended themselves via counterattack against the larger cetacean. However, orcas have been observed successfully attacking walruses with few or no injuries.


Relationship with humans


Conservation

In the 18th and 19th centuries, the walrus was heavily exploited by American and European
sealers Sealer may refer either to a person or ship engaged in seal hunting, or to a sealant; associated terms include: Seal hunting * Sealer Hill, South Shetland Islands, Antarctica * Sealers' Oven, bread oven of mud and stone built by sealers around 180 ...
and whalers, leading to the near-
extirpation Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
of the Atlantic subspecies. As early as 1871 traditional hunters were expressing concern about the numbers of walrus being hunted by whaling fleets. Commercial walrus harvesting is now outlawed throughout its range, although Chukchi,
Yupik Yupik may refer to: * Yupik peoples, a group of indigenous peoples of Alaska and the Russian Far East * Yupik languages, a group of Eskimo-Aleut languages Yupꞌik (with the apostrophe) may refer to: * Yup'ik people The Yup'ik or Yupiaq (sg ...
and
Inuit Inuit (; iu, ᐃᓄᐃᑦ 'the people', singular: Inuk, , dual: Inuuk, ) are a group of culturally similar indigenous peoples inhabiting the Arctic and subarctic regions of Greenland, Labrador, Quebec, Nunavut, the Northwest Territories ...
peoples are permitted to kill small numbers towards the end of each summer. Traditional hunters used all parts of the walrus. The meat, often preserved, is an important winter nutrition source; the flippers are fermented and stored as a delicacy until spring; tusks and bone were historically used for tools, as well as material for handicrafts; the oil was rendered for warmth and light; the tough hide made rope and house and boat coverings; and the intestines and gut linings made waterproof parkas. While some of these uses have faded with access to alternative technologies, walrus meat remains an important part of local diets, and tusk carving and engraving remain a vital art form. According to
Adolf Erik Nordenskiöld Nils Adolf Erik Nordenskiöld (18 November 183212 August 1901) was a Finland-Swedish aristocrat, geologist, mineralogist and Arctic explorer. He was a member of the Fenno-Swedish Nordenskiöld family of scientists and held the title of a friher ...
, European hunters and Arctic explorers found walrus meat not particularly tasty, and only ate it in case of necessity; however walrus
tongue The tongue is a muscular organ (anatomy), organ in the mouth of a typical tetrapod. It manipulates food for mastication and swallowing as part of the digestive system, digestive process, and is the primary organ of taste. The tongue's upper surfa ...
was a delicacy. File:Walrus hunter 1911.jpg, Hunter sitting on dozens of walruses killed for their tusks, 1911 File:PolarBearWalrusTuskCarving.jpg, alt=Photo of section of tusk, Walrus tusk
scrimshaw Scrimshaw is scrollwork, engravings, and carvings done in bone or ivory. Typically it refers to the artwork created by whalers, engraved on the byproducts of whales, such as bones or cartilage. It is most commonly made out of the bones and teeth ...
made by Chukchi artisans depicting polar bears attacking walruses, on display in the Magadan Regional Museum,
Magadan Magadan ( rus, Магадан, p=məɡɐˈdan) is a port town and the administrative center of Magadan Oblast, Russia, located on the Sea of Okhotsk in Nagayev Bay (within Taui Bay) and serving as a gateway to the Kolyma region. History Maga ...
, Russia File:Walrus in Marineland, Ontario, Canada - 2017 (37223504105).jpg, Trained walrus in captivity at Marineland File:Valrossarna på Skansen matas av djurskötare och herre i paletå och hög hatt - Nordiska Museet - NMA.0048348.jpg, Walrus being fed at
Skansen Skansen (; "the Sconce") is the oldest open-air museum and zoo in Sweden located on the island Djurgården in Stockholm, Sweden. It was opened on 11 October 1891 by Artur Hazelius (1833–1901) to show the way of life in the different parts of S ...
in
Stockholm Stockholm () is the Capital city, capital and List of urban areas in Sweden by population, largest city of Sweden as well as the List of urban areas in the Nordic countries, largest urban area in Scandinavia. Approximately 980,000 people liv ...
,
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
, 1908
Walrus hunts are regulated by resource managers in
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
, the United States, Canada, and
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...
(self-governing country in the
Kingdom of Denmark The Danish Realm ( da, Danmarks Rige; fo, Danmarkar Ríki; kl, Danmarkip Naalagaaffik), officially the Kingdom of Denmark (; ; ), is a sovereign state located in Northern Europe and Northern North America. It consists of Denmark, metropolitan ...
), and representatives of the respective hunting communities. An estimated four to seven thousand Pacific walruses are harvested in
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
and in Russia, including a significant portion (about 42%) of struck and lost animals. Several hundred are removed annually around Greenland. The sustainability of these levels of harvest is difficult to determine given uncertain population estimates and parameters such as
fecundity Fecundity is defined in two ways; in human demography, it is the potential for reproduction of a recorded population as opposed to a sole organism, while in population biology, it is considered similar to fertility, the natural capability to pr ...
and
mortality Mortality is the state of being mortal, or susceptible to death; the opposite of immortality. Mortality may also refer to: * Fish mortality, a parameter used in fisheries population dynamics to account for the loss of fish in a fish stock throug ...
. The ''Boone and Crockett Big Game Record'' book has entries for Atlantic and Pacific walrus. The recorded largest tusks are just over 30 inches and 37 inches long respectively. The effects of
global climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
are another element of concern. The extent and thickness of the pack ice has reached unusually low levels in several recent years. The walrus relies on this ice while giving birth and aggregating in the reproductive period. Thinner pack ice over the Bering Sea has reduced the amount of resting habitat near optimal feeding grounds. This more widely separates lactating females from their calves, increasing nutritional stress for the young and lower reproductive rates. Reduced coastal sea ice has also been implicated in the increase of stampeding deaths crowding the shorelines of the
Chukchi Sea Chukchi Sea ( rus, Чуко́тское мо́ре, r=Chukotskoye more, p=tɕʊˈkotskəjə ˈmorʲɪ), sometimes referred to as the Chuuk Sea, Chukotsk Sea or the Sea of Chukotsk, is a marginal sea of the Arctic Ocean. It is bounded on the west b ...
between eastern Russia and western Alaska. Analysis of trends in ice cover published in 2012 indicate that Pacific walrus populations are likely to continue to decline for the foreseeable future, and shift further north, but that careful conservation management might be able to limit these effects. Currently, two of the three walrus subspecies are listed as "least-concern" by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
, while the third is "data deficient". The Pacific walrus is not listed as "depleted" according to the
Marine Mammal Protection Act The Marine Mammal Protection Act (MMPA) was the first act of the United States Congress to call specifically for an ecosystem approach to wildlife management. Authority MMPA was signed into law on October 21, 1972, by President Richard Nixon ...
nor as "threatened" or "endangered" under the
Endangered Species Act The Endangered Species Act of 1973 (ESA or "The Act"; 16 U.S.C. § 1531 et seq.) is the primary law in the United States for protecting imperiled species. Designed to protect critically imperiled species from extinction as a "consequence of ec ...
. The Russian Atlantic and Laptev Sea populations are classified as Category 2 (decreasing) and Category 3 (rare) in the Russian Red Book. Global trade in
walrus ivory The walrus (''Odobenus rosmarus'') is a large flippered marine mammal with a discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. The walrus is the only living species in the fami ...
is restricted according to a
CITES CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
Appendix 3 listing. In October 2017, the
Center for Biological Diversity The Center for Biological Diversity is a nonprofit membership organization known for its work protecting endangered species through legal action, scientific petitions, creative media and grassroots activism. It was founded in 1989 by Kieran Suckl ...
announced they would sue the
U.S. Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with othe ...
to force it to classify the Pacific Walrus as a threatened or endangered species. In 1952, walruses in
Svalbard Svalbard ( , ), also known as Spitsbergen, or Spitzbergen, is a Norwegian archipelago in the Arctic Ocean. North of mainland Europe, it is about midway between the northern coast of Norway and the North Pole. The islands of the group range ...
were nearly gone due to ivory hunting over a 300 years period, but the Norwegian government banned their commercial hunting and the walruses began to rebound in 2006, making their population increase to 2,629.


Culture


Folklore

The walrus plays an important role in the religion and
folklore Folklore is shared by a particular group of people; it encompasses the traditions common to that culture, subculture or group. This includes oral traditions such as tales, legends, proverbs and jokes. They include material culture, ranging ...
of many
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
peoples. Skin and bone are used in some ceremonies, and the animal appears frequently in legends. For example, in a Chukchi version of the widespread myth of the Raven, in which
Raven A raven is any of several larger-bodied bird species of the genus ''Corvus''. These species do not form a single taxonomic group within the genus. There is no consistent distinction between "crows" and "ravens", common names which are assigned t ...
recovers the sun and the moon from an evil spirit by seducing his daughter, the angry father throws the daughter from a high cliff and, as she drops into the water, she turns into a walrus – possibly the original walrus. According to various legends, the tusks are formed either by the trails of mucus from the weeping girl or her long braids. This myth is possibly related to the Chukchi myth of the old walrus-headed woman who rules the bottom of the sea, who is in turn linked to the Inuit goddess Sedna. Both in Chukotka and
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
, the
aurora borealis An aurora (plural: auroras or aurorae), also commonly known as the polar lights, is a natural light display in Earth's sky, predominantly seen in high-latitude regions (around the Arctic and Antarctic). Auroras display dynamic patterns of br ...
is believed to be a special world inhabited by those who died by violence, the changing rays representing deceased souls playing ball with a walrus head. File:Ivorymasks.jpg, alt=Photo of two masks: In the center is the image of a face, surrounded by a ring, in turn surrounded by eight white rectangular pieces., Walrus ivory masks made by
Yupik Yupik may refer to: * Yupik peoples, a group of indigenous peoples of Alaska and the Russian Far East * Yupik languages, a group of Eskimo-Aleut languages Yupꞌik (with the apostrophe) may refer to: * Yup'ik people The Yup'ik or Yupiaq (sg ...
in
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
File:Briny Beach.jpg, alt=Drawing of walrus, and square-headed men, both perched on rocks, with ocean and cliffs in background,
John Tenniel Sir John Tenniel (; 28 February 182025 February 1914)Johnson, Lewis (2003), "Tenniel, John", ''Grove Art Online, Oxford Art Online'', Oxford University Press. Web. Retrieved 12 December 2016. was an English illustrator, graphic humorist and pol ...
's illustration for
Lewis Carroll Charles Lutwidge Dodgson (; 27 January 1832 – 14 January 1898), better known by his pen name Lewis Carroll, was an English author, poet and mathematician. His most notable works are ''Alice's Adventures in Wonderland'' (1865) and its sequel ...
's poem "
The Walrus and the Carpenter "The Walrus and the Carpenter" is a narrative poem by Lewis Carroll that appears in his book ''Through the Looking-Glass'', published in December 1871. The poem is recited in chapter four, by Tweedledum and Tweedledee to Alice. The poem is co ...
" File:Hugo-de-Groot-Nederlandtsche-jaerboeken MG 0188.tif, Dutch explorers fight a walrus on the coast of
Novaya Zemlya Novaya Zemlya (, also , ; rus, Но́вая Земля́, p=ˈnovəjə zʲɪmˈlʲa, ) is an archipelago in northern Russia. It is situated in the Arctic Ocean, in the extreme northeast of Europe, with Cape Flissingsky, on the northern island, ...
, 1596
Most of the distinctive 12th-century
Lewis Chessmen The Lewis chessmen ( no, Lewisbrikkene; gd, Fir-Tàilisg; sco, Lewis chesmen) or Uig chessmen, named after the island or the bay where they were found, are a group of distinctive 12th-century chess pieces, along with other game pieces, most o ...
from northern Europe are carved from walrus ivory, though a few have been found to be made of whales' teeth.


Literature

Because of its distinctive appearance, great bulk, and immediately recognizable whiskers and tusks, the walrus also appears in the popular cultures of peoples with little direct experience with the animal, particularly in English children's literature. Perhaps its best-known appearance is in
Lewis Carroll Charles Lutwidge Dodgson (; 27 January 1832 – 14 January 1898), better known by his pen name Lewis Carroll, was an English author, poet and mathematician. His most notable works are ''Alice's Adventures in Wonderland'' (1865) and its sequel ...
's whimsical poem "
The Walrus and the Carpenter "The Walrus and the Carpenter" is a narrative poem by Lewis Carroll that appears in his book ''Through the Looking-Glass'', published in December 1871. The poem is recited in chapter four, by Tweedledum and Tweedledee to Alice. The poem is co ...
" that appears in his 1871 book ''
Through the Looking-Glass ''Through the Looking-Glass, and What Alice Found There'' (also known as ''Alice Through the Looking-Glass'' or simply ''Through the Looking-Glass'') is a novel published on 27 December 1871 (though indicated as 1872) by Lewis Carroll and the ...
''. In the poem, the
eponymous An eponym is a person, a place, or a thing after whom or which someone or something is, or is believed to be, named. The adjectives which are derived from the word eponym include ''eponymous'' and ''eponymic''. Usage of the word The term ''epon ...
antiheroes An antihero (sometimes spelled as anti-hero) or antiheroine is a main character in a story who may lack conventional heroic qualities and attributes, such as idealism, courage, and morality. Although antiheroes may sometimes perform action ...
use trickery to consume a great number of
oyster Oyster is the common name for a number of different families of salt-water bivalve molluscs that live in marine or brackish habitats. In some species, the valves are highly calcified, and many are somewhat irregular in shape. Many, but not al ...
s. Although Carroll accurately portrays the biological walrus's appetite for bivalve mollusks, oysters, primarily nearshore and
intertidal The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of Marine habitat, habitats with var ...
inhabitants, these organisms in fact comprise an insignificant portion of its diet in captivity. The "walrus" in the cryptic Beatles song "
I Am the Walrus "I Am the Walrus" is a song by the English rock band the Beatles from their 1967 television film ''Magical Mystery Tour''. Written by John Lennon and credited to Lennon–McCartney, it was released as the B-side to the single "Hello, Goodbye" a ...
" is a reference to the Lewis Carroll poem. Another appearance of the walrus in literature is in the story "The White Seal" in
Rudyard Kipling Joseph Rudyard Kipling ( ; 30 December 1865 – 18 January 1936)''The Times'', (London) 18 January 1936, p. 12. was an English novelist, short-story writer, poet, and journalist. He was born in British India, which inspired much of his work. ...
's ''
The Jungle Book ''The Jungle Book'' (1894) is a collection of stories by the English author Rudyard Kipling. Most of the characters are animals such as Shere Khan the tiger and Baloo the bear, though a principal character is the boy or "man-cub" Mowgli, ...
'', where it is the "old Sea Vitch—the big, ugly, bloated, pimpled, fat-necked, long-tusked walrus of the North Pacific, who has no manners except when he is asleep".


See also

*
Elephant seal Elephant seals are very large, oceangoing earless seals in the genus ''Mirounga''. Both species, the northern elephant seal (''M. angustirostris'') and the southern elephant seal (''M. leonina''), were hunted to the brink of extinction for oil ...


References


Further reading

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External links

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Biologist Tracks Walruses Forced Ashore As Ice Melts
– audio report by ''
NPR National Public Radio (NPR, stylized in all lowercase) is an American privately and state funded nonprofit media organization headquartered in Washington, D.C., with its NPR West headquarters in Culver City, California. It differs from other ...
''
Thousands Of Walruses Crowd Ashore Due To Melting Sea Ice
– video by ''
National Geographic ''National Geographic'' (formerly the ''National Geographic Magazine'', sometimes branded as NAT GEO) is a popular American monthly magazine published by National Geographic Partners. Known for its photojournalism, it is one of the most widely ...
''
Voices in the Sea – Sounds of the Walrus
{{good article Pinnipeds of the Arctic Ocean Mammals described in 1758 Mammals of Asia Mammals of Canada Mammals of Greenland Mammals of North America Mammals of Russia Mammals of the United States Mammals of Europe Marine mammals Pinnipeds of North America Odobenids Taxa named by Carl Linnaeus Holarctic fauna