Tunicate
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Tunicate
A tunicate is a marine invertebrate animal, a member of the subphylum Tunicata (). It is part of the Chordata, a phylum which includes all animals with dorsal nerve cords and notochords (including vertebrates). The subphylum was at one time called Urochordata, and the term urochordates is still sometimes used for these animals. They are the only chordates that have lost their myomeric segmentation, with the possible exception of the 'seriation of the gill slits'. Some tunicates live as solitary individuals, but others replicate by budding and become colonies, each unit being known as a zooid. They are marine filter feeders with a water-filled, sac-like body structure and two tubular openings, known as siphons, through which they draw in and expel water. During their respiration and feeding, they take in water through the incurrent (or inhalant) siphon and expel the filtered water through the excurrent (or exhalant) siphon. Most adult tunicates are sessile, immobile and perman ...
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Chordata
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also Bilateral symmetry, bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit Metameric, metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cep ...
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Filter Feeder
Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feeding are clams, krill, sponges, baleen whales, and many fish (including some sharks). Some birds, such as flamingos and certain species of duck, are also filter feeders. Filter feeders can play an important role in clarifying water, and are therefore considered ecosystem engineers. They are also important in bioaccumulation and, as a result, as indicator organisms. Fish Most forage fish are filter feeders. For example, the Atlantic menhaden, a type of herring, lives on plankton caught in midwater. Adult menhaden can filter up to four gallons of water a minute and play an important role in clarifying ocean water. They are also a natural check to the deadly red tide. Extensive article on the role of menhaden in the ecosystem and possible resul ...
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Vertebrates
Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with currently about 69,963 species described. Vertebrates comprise such groups as the following: * jawless fish, which include hagfish and lampreys * jawed vertebrates, which include: ** cartilaginous fish (sharks, rays, and ratfish) ** bony vertebrates, which include: *** ray-fins (the majority of living bony fish) *** lobe-fins, which include: **** coelacanths and lungfish **** tetrapods (limbed vertebrates) Extant vertebrates range in size from the frog species ''Paedophryne amauensis'', at as little as , to the blue whale, at up to . Vertebrates make up less than five percent of all described animal species; the rest are invertebrates, which lack vertebral columns. The vertebrates traditionally include the hagfish, which do no ...
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Ascidiacea
Ascidiacea, commonly known as the ascidians, tunicates (in part), and sea squirts (in part), is a polyphyletic class in the subphylum Tunicata of sac-like marine invertebrate filter feeders. Ascidians are characterized by a tough outer "tunic" made of a polysaccharide. Ascidians are found all over the world, usually in shallow water with salinities over 2.5%. While members of the Thaliacea and Larvacea (Appendicularia) swim freely like plankton, sea squirts are sessile animals after their larval phase: they then remain firmly attached to their substratum, such as rocks and shells. There are 2,300 species of ascidians and three main types: solitary ascidians, social ascidians that form clumped communities by attaching at their bases, and compound ascidians that consist of many small individuals (each individual is called a zooid) forming colonies up to several meters in diameter. Sea squirts feed by taking in water through a tube, the oral siphon. The water enters the mouth ...
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Thaliacea
The Thaliacea comprise a class of marine animals within the subphylum Tunicata. Unlike their benthic relatives the ascidians, thaliaceans are free-floating (pelagic) for their entire lifespan. The group includes species with complex life cycles, with both solitary and colonial forms. Anatomy The three orders of thaliaceans are filter feeders. Pyrosomes are colonial animals, with multiple tiny ascidian-like zooids arranged in a cylinder closed at one end. All of the atrial siphons point inwards, emptying into a single, common cloaca in the centre of the cylinder. As the water exhaled by the zooids exits through a common opening, the water movement slowly propels the pyrosome through the sea. Salps and doliolids have a transparent barrel-shaped body through which they pump water, propelling them through the sea, and from which they extract food. The bulk of the body consists of the large pharynx. Water enters the pharynx through the large buccal siphon at the front end of the a ...
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Polycarpa Aurata
''Polycarpa aurata'', also known as the ox heart ascidian, the gold-mouth sea squirt or the ink-spot sea squirt, is a species of tunicate in the family Styelidae. Description ''Polycarpa aurata'' grows to a height of . It has an urn-shaped, hollow body with two siphons, one at the top and the other on the side. The body colour of this tunicate is white with purple and orange patches and purple lines. The inside is yellow or orange and this is visible around the rim of the siphons. Distribution This species is found in the tropical eastern Indian Ocean and the western Pacific Ocean. Its range includes the Philippines, Indonesia and northern Australia. Its depth range is . Biology Tunicates feed by drawing water in through the branchial siphon at the top, filtering out phytoplankton, bacteria and other food particles, before expelling the water through the atrial siphon at the side. Sometimes hydroids and algae grow on the outside of the tunicate and nudibranchs such as ''Nem ...
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Invertebrate
Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate subphylum Vertebrata. Familiar examples of invertebrates include arthropods, mollusks, annelids, echinoderms and cnidarians. The majority of animal species are invertebrates; one estimate puts the figure at 97%. Many invertebrate taxa have a greater number and variety of species than the entire subphylum of Vertebrata. Invertebrates vary widely in size, from 50  μm (0.002 in) rotifers to the 9–10 m (30–33 ft) colossal squid. Some so-called invertebrates, such as the Tunicata and Cephalochordata, are more closely related to vertebrates than to other invertebrates. This makes the invertebrates paraphyletic, so the term has little meaning in taxonomy. Etymology The word "invertebrate" comes from the Latin word ''vertebra'', whi ...
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Doliolida
The Doliolida are an order of small marine animals of the subphylum Tunicata. They are in the class Thaliacea, which also includes the salps and pyrosomes. The doliolid body is small, typically 1–2 mm long, and barrel-shaped; it features two wide siphons, one at the front and the other at the back end, and eight or nine circular muscle strands reminiscent of barrel bands. Like all tunicates, except for the predatory tunicate, they are filter feeders. Unlike the related class Ascidiacea, which are sessile, but like the class Appendicularia, they are free-swimming plankton; cilia pump water through the body which drives them forward. As the water passes through, small particles and plankton on which the animal feeds are strained from the water by the gill slits. Doliolids can also move by contracting the muscular bands around the body creating a temporary water jet that thrusts them forward or backward quite quickly. The Doliolida have a complicated life cycle that i ...
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Yarnemia
''Yarnemia ascidiformis'' is a fossil tentatively classified as a tunicate. While ''Y. ascidiformis'' looks similar to tunicates, the oldest unequivocal tunicate, ''Shankouclava'' dates to the Cambrian period, while ''Y. ascidiformis'' is Ediacaran in age. Etymology The generic name ''Yarnemia'' comes from the village of Yarnema near which the first specimens were found. The specific epithet, ''ascidiformis'', refers to the likeness to ascidians. See also Other Ediacaran biota tentatively identified as tunicates include * ''Ausia fenestrata'' * ''Burykhia hunti ''Burykhia hunti'' is a Precambrian fossil from the White Sea region of Russia dating to . It is considered of ascidian affinity, due to the sac-like morphology and a series of distinctly perforated bands reminiscent of a tunicate pharynx. If ' ...'' References Ediacaran life Tunicate genera Prehistoric chordate genera Fossil taxa described in 1984 {{paleo-deuterostome-stub ...
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Larvaceans
Larvaceans, class Appendicularia, are solitary, free-swimming tunicates found throughout the world's oceans. Like most tunicates, larvaceans are filter feeders. Unlike most other tunicates, they live in the pelagic zone, specifically in the photic zone, or sometimes deeper. They are transparent planktonic animals, generally less than in body length, excluding the tail. Anatomy The adult larvaceans resemble the tadpole-like larvae of most tunicates. Like a common tunicate larva, the adult Appendicularia have a discrete trunk and tail. Larvaceans produce a "house" made of mucopolysaccharides and cellulose. In most species, the house surrounds the animal like a bubble. Even for species in which the house does not completely surround the body, such as ''Fritillaria (tunicate), Fritillaria'', the house is always present and attached to at least one surface. These houses are discarded and replaced regularly as the animal grows in size and its filters become clogged; in ''Oikopleura'' ...
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Notochord
In anatomy, the notochord is a flexible rod which is similar in structure to the stiffer cartilage. If a species has a notochord at any stage of its life cycle (along with 4 other features), it is, by definition, a chordate. The notochord consists of inner, vacuolated cells covered by fibrous and elastic sheaths, lies along the anteroposterior axis (''front to back''), is usually closer to the dorsal than the ventral surface of the embryo, and is composed of cells derived from the mesoderm. The most commonly cited functions of the notochord are: as a midline tissue that provides directional signals to surrounding tissue during development, as a skeletal (structural) element, and as a vertebral precursor. In lancelets the notochord persists throughout life as the main structural support of the body. In tunicates the notochord is present only in the larval stage, being completely absent in the adult animal. In these invertebrate chordates, the notochord is not vacuolated. In all ...
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Shankouclava
''Shankouclava'' is an extinct genus of tunicate. It represents the oldest, unequivocal genus of tunicates, from 520 million years ago. It has been found in the Lower Cambrian Maotianshan Shale at Shankou village, Anning, near Kunming (South China). Each of the eight specimens found and used for description were isolated, suggesting that the genus was solitary and not colonial. The generic name is composed of the fossil locality, Shankou, and the Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ... word ''clava'' (club-shaped). ''Shankouclava'' had a soft, sac-like body that was elongated and pointed proximally. The body lengths of individuals vary from 2 cm (0.8 in) to 4 cm (1.6 in). References Tunicate genera Prehistoric chordate genera Fossil tax ...
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