''Tetrapanax papyrifer'', the rice-paper plant (''通草—tong cao''), is an

evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...

shrub in the

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Araliaceae The Araliaceae are a family of flowering plants composed of about 43 genera and around 1500 species consisting of primarily woody plants and some herbaceous plants. The morphology of Araliaceae varies widely, but it is predominantly distinguish ...

, the sole

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

in the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

''Tetrapanax''. The specific epithet is frequently misspelled as "papyriferum", "papyriferus", or "papyrifera". It is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...

Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...

, but widely cultivated in

East Asia East Asia is the eastern region of Asia, which is defined in both Geography, geographical and culture, ethno-cultural terms. The modern State (polity), states of East Asia include China, Japan, Mongolia, North Korea, South Korea, and Taiwan. ...

and sometimes in other

tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...

regions as well. The species was once included in the genus ''

Fatsia ''Fatsia'' is a small genus of three species of evergreen shrubs in the family Araliaceae native to southern Japan and Taiwan. They typically have stout, sparsely branched stems bearing spirally-arranged, large leathery, palmately lobed leav ...

'' as ''Fatsia papyrifera''. A second species, ''Tetrapanax tibetanus'', is now regarded as a synonym of '' Merrilliopanax alpinus''.

Description

''Tetrapanax'' grows between 3–7 metres tall, usually with unbranched stems 2 cm in diameter, and bearing a rosette of large leaves atop the crown; the top of the plant can visually, albeit superficially, appear similar to a number of plants belonging to '' Arecaceae'' (the palm family). The leaves are carried on 40–60 cm petioles; the orbicular leaf blade measures from 30–50 cm across, with anywhere from 5-12 deeply palmately lobes, the central lobes being larger and Y-forked near the end. It spreads extensively, by sprouts and runners from the

root In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the su ...

system, underground. The

inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...

is a large panicle of hemispherical or globular

umbel In botany, an umbel is an inflorescence that consists of a number of short flower stalks (called pedicels) that spread from a common point, somewhat like umbrella ribs. The word was coined in botanical usage in the 1590s, from Latin ''umbella'' "p ...

s, at the end of the stem. The

flowers A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...

have 4-6 small, white petals. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...

is a small berry.

Uses

''Tetrapanax papyrifer'' is used in

traditional Chinese medicine Traditional Chinese medicine (TCM) is an alternative medical practice drawn from traditional medicine in China. It has been described as "fraught with pseudoscience", with the majority of its treatments having no logical mechanism of acti ...

. The

pith Pith, or medulla, is a tissue in the stems of vascular plants. Pith is composed of soft, spongy parenchyma cells, which in some cases can store starch. In eudicotyledons, pith is located in the center of the stem. In monocotyledons, it ext ...

from the stem is used to make a substance commonly known as

rice paper "Rice paper" has many varieties such as rice paper made from tree bark to make drawing and writing paper or from rice flour and tapioca flour and then mixed with salt and water to produce a thin rice cake and dried to become harder and paper-like ...

, but more properly termed pith paper. The species is cultivated as an ornamental specimen plant, and has received the

Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...

's Award of Garden Merit.

Cultivars

The

cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...

'Rex' is a semi-evergreen shrub or tree with huge palmate leaves. Classified by the Royal Horticultural Society as H4, it is evergreen in mild locations,

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...

where temperatures fall below freezing, and herbaceous with more prolonged freezing. It prefers a sheltered position in full sun or partial shade.

Gallery

File:Tetrapanax papyrifer Jardin des Plantes.jpg , At the Jardin des plantes de

Paris Paris () is the Capital city, capital and List of communes in France with over 20,000 inhabitants, most populous city of France, with an estimated population of 2,165,423 residents in 2019 in an area of more than 105 km² (41 sq mi), ma ...

, France File:Tetrapanax_papyrifer_Leaf.jpg, Leaf File:Tetrapanax papyrifer4.jpg, Inflorescence

References

External links

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Chinese Botanical Paintings, ''Tetrapanax papyriferum'' (Hook.) Koch
{{Taxonbar, from1=Q8085880, from2=Q712166 Araliaceae Monotypic Apiales genera Endemic flora of Taiwan