Tepui Foliage-gleaner
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The tepui foliage-gleaner (''Syndactyla roraimae''), also known as the white-throated foliage-gleaner, is a species of bird in the Furnariinae subfamily of the ovenbird family Furnariidae. It is found in Brazil,
Guyana Guyana ( or ), officially the Cooperative Republic of Guyana, is a country on the northern mainland of South America. Guyana is an indigenous word which means "Land of Many Waters". The capital city is Georgetown. Guyana is bordered by the ...
and Venezuela.


Taxonomy and systematics

The tepui foliage-gleaner was originally described in genus ''
Philydor ''Philydor'' is a genus of foliage-gleaners, birds in the ovenbird family Furnariidae.Remsen, J.V., Jr (2017). Ovenbirds (Furnariidae). In: del Hoyo, J., Elliott, A., Sargatal, J., Christie, D.A. & de Juana, E. (eds.). Handbook of the Birds of ...
'' and then placed in genus ''
Automolus ''Automolus'' is a genus of bird in the ovenbird family Furnariidae. Taxonomy The genus ''Automolus'' was introduced in 1853 by the German naturalist Ludwig Reichenbach to accommodate the taxon ''Sphenura sulphurascens'' Lichtenstein, now tre ...
''. Publications in 2008 and 2011 showed that it belonged in ''Syndactyla''.Derryberry, E. P., S. Claramunt, G. Derryberry, R. T. Chesser, J. Cracraft, A. Aleixo, J. Pérez-Emán, J. V. Remsen, Jr., and R. T. Brumfield. (2011). Lineage diversification and morphological evolution in a large-scale continental radiation: the Neotropical ovenbirds and woodcreepers (Aves: Furnariidae). Evolution 65(10):2973–2986. The tepui foliage-gleaner has these four subspecies: *''S. r. paraquensis'' ( Phelps, WH & Phelps, WH Jr, 1947) *''S. r. duidae'' (
Chapman Chapman may refer to: Businesses * Chapman Entertainment, a former British television production company * Chapman Guitars, a guitar company established in 2009 by Rob Chapman * Chapman's, a Canadian ice cream and ice water products manufacturer ...
, 1939)
*''S. r. roraimae'' (
Hellmayr Carl Eduard Hellmayr (29 January 1878 in Vienna, Austria – 24 February 1944 in Orselina, Switzerland) was an Austrian ornithologist. Biography Hellmayr was born in Vienna and studied at the University of Vienna, although he did not complete hi ...
, 1917)
*''S. r. urutani'' (Phelps, WH Jr & Dickerman, 1980)


Description

The tepui foliage-gleaner is about long and weighs . It is the smallest member of its genus and has the most striking facial pattern. The sexes have the same plumage. Adults of the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
''S. r. roraimae'' have a white
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also ...
that contrasts with their blackish-brown face. Their crown is dark grayish brown, their back rich dark
rufescent Rufous () is a color that may be described as reddish-brown or brownish-red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a dia ...
brown, and their rump and uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are sm ...
chestnut-rufous. Their tail is chestnut-rufous. Their wing coverts are mostly rich dark rufescent brown, their
primary Primary or primaries may refer to: Arts, entertainment, and media Music Groups and labels * Primary (band), from Australia * Primary (musician), hip hop musician and record producer from South Korea * Primary Music, Israeli record label Works * ...
coverts darker brown, and their flight feathers a more rufescent brown. Their throat and malar area are white, their breast and belly medium brown, and their flanks and undertail coverts darker brown. Their iris is brown to dark brown, their maxilla black, their mandible highly variable from pinkish gray to black, and their legs and feet olive-yellow to greenish gray. Juveniles have an ochraeous supercilium, and more rufous underparts than adults, with dusky tips on the breast and belly feathers.Remsen, Jr., J. V. (2020). White-throated Foliage-gleaner (''Syndactyla roraimae''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.wtfgle1.01 retrieved August 30, 2023 Subspecies ''S. r. paraquensis'' has a paler and more yellowish brown back and a paler and more grayish olive breast and belly than the nominate. ''S. r. duidae'' has a more rufescent crown and underparts with a much brighter belly than the nominate. ''S. r. urutani'' is paler overall than the other subspecies, and is less rufous above and less ochraceous below.


Distribution and habitat

The tepui foliage-gleaner is found in the tepui region where eastern Venezuela, western Guyana, and northwestern Brazil meet. The subspecies' ranges are disjunct and they are found thus: *''S. r. paraquensis'': Cerro Paraque in south-central Venezuela's Amazonas state *''S. r. duidae'': several tepuis in Venezuela's Amazonas state, and
Cerro de la Neblina Cerro de la Neblina (lit. "Mountain of the Mist"), also known as Serra da Neblina in Brazil and Sierra de la Neblina in Venezuela, is a sandstone massif located in the northern Amazon Basin. It is a tilted, heavily eroded plateau, with a deep can ...
straddling the Venezuela/Brazil border *''S. r. roraimae'':
Mount Roraima Mount Roraima ( es, Monte Roraima; Tepuy Roraima; Cerro Roraima pt, Monte Roraima ) is the highest of the Pakaraima chain of tepuis (table-top mountain) or plateaux in South America. It is located at the junction of Venezuela, Brazil and Guyana. ...
straddling the junction of Venezuela, Guyana, and Brazil, and other tepuis in southeastern Venezuela and western Guyana *''S. r. urutani'': several tepuis in Venezuela's Bolívar state The tepui foliage-gleaner inhabits montane evergreen forest, wind-stunted woodland, and the transition zone between them. In elevation it mostly ranges between .


Behavior


Movement

The tepui foliage-gleaner is a year-round resident throughout its range.


Feeding

The tepui foliage-gleaner feeds on a variety of arthropods such as
Coleoptera Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
and
Orthoptera Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grassho ...
. It forages singly, in pairs, or in small family groups and often joins mixed-species feeding flocks. It mostly forages in the forest's understorey but will go up to the mid-storey. It typically stays low in the undergrowth but also seeks prey while clambering about in vine tangles and hitching along branches and up trunks. It gleans and probes for it prey among live and dead leaves, moss, and on branches. It occasionally hangs upside down while foraging.


Breeding

The tepui foliage-gleaner's breeding season in Guyana apparently ends in March or April, but nothing else is known about the species' breeding biology.


Vocalization

The tepui foliage-gleaner's song is "a long accelerating series of very harsh guttural 'jjza' notes, rising in pitch, usually preceded by a few single short notes". Its call is "a short, harsh 'chek' or 'tzik' ".


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed the tepui foliage-gleaner as being of Least Concern. It has a restricted range and an unknown population size, but the latter is believed to be stable. No immediate threats have been identified. It is considered uncommon to locally common and occurs in one Venezuelan national park.


References

{{Taxonbar, from=Q1267130 tepui foliage-gleaner Birds of the Tepuis tepui foliage-gleaner Taxonomy articles created by Polbot