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''Sordes'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of small
pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
from the late
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
( Oxfordian/
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 154.8 ±0.8 Ma and 149.2 ±0.7 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
)
Karabastau Svita The Karabastau Formation () is a geological Formation (geology), formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle Jurassic, Middle to Late Jurassic. It is an important locality for insect f ...
of
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a landlocked country primarily in Central Asia, with a European Kazakhstan, small portion in Eastern Europe. It borders Russia to the Kazakhstan–Russia border, north and west, China to th ...
. This genus was named in 1971 by Aleksandr Grigorevich Sharov. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Sordes pilosus''. The genus name is
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
for "filth" or "scum"; but Sharov translates it as "", which means "devil" or "evil spirit", so the intended translation is "hairy devil" (the specific name is Latin for "hairy"; despite ''sordes'' being feminine, it has not yet been emended to ''pilosa'').


Discovery

''Sordes'' is based on the
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
PIN 2585/3, which consists of a crushed relatively complete skeleton on a slab. It was found in the 1960s at the foothills of the Karatau in
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a landlocked country primarily in Central Asia, with a European Kazakhstan, small portion in Eastern Europe. It borders Russia to the Kazakhstan–Russia border, north and west, China to th ...
. Sharov had already referred a
paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype (biology), isotype ...
or second specimen: PIN 2470/1, again a fairly complete skeleton on a slab. By 2003 another six specimens had been discovered.


Description

''Sordes'' had a 0.63 m (2 ft) wingspan. The wings were relatively short. ''Sordes'' had, according to Sharov and Unwin, wing membranes attached to the legs and a membrane between the legs. It had a short neck. It had a long tail, accounting for over half its length, with at the end an elongated vane.


Skull and dentition

It had a slender, not round, head with moderately long, pointed jaws. The skull was about 8 cm (3.2 in) long. Unlike many pterosaurs, it had no head crest. The teeth in the frontal half of the jaws are large and pointed to facilitate prey capture. The teeth beyond these in the rear half of the jaw are much smaller and more numerous than those at the front, suggesting that they were more for crushing. Together these two types of teeth indicate specialisation for prey that was difficult to catch yet required some effort to eat. Likely contenders are invertebrates with tougher exoskeletons, or amphibians that were slippery to catch and then required some crunching before they could be swallowed.


Pycnofibers

The
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
shows remains of the soft parts, such as membranes and hair-like filaments. This was the first unequivocal proof that pterosaurs had a layer of hair-like filaments covering their bodies, later named
pycnofibres Pterosaurs are an extinct clade of flying reptiles in the Order (biology), order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosau ...
. The pycnofibres served as insulation, an indication the group was
warm-blooded Warm-blooded is a term referring to animal species whose bodies maintain a temperature higher than that of their environment. In particular, homeothermic species (including birds and mammals) maintain a stable body temperature by regulating ...
, and provided a streamlined flight profile. The pycnofibres are present in two main types: longer at the extreme part of the wing membrane and shorter near the body. In the 1990s, David Unwin argued that both types were essentially not hairs but reinforcing fibres of the flight membranes.Elgin, R.A., Hone, D.W.E., and Frey, E. 2011. The extent of the pterosaur flight membrane. Acta Palaeontologica Polonica 56 (1): 99–111

/ref> Later he emphasized that "hair" in the form of pycnofibres was indeed present on the body, after the find of new specimens clearly showing this.


Classification

''Sordes'' has been assigned to the family
Rhamphorhynchidae Rhamphorhynchidae is a group of early pterosaurs named after '' Rhamphorhynchus'', that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley.Seeley, H.G. (1870). "The Orithosauria: An Elementary Study ...
. These were among the earliest of the pterosaurs, evolving in the late
Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
and surviving to the late Jurassic. According to Unwin, within Rhamphorhynchidae ''Sordes'' belonged to the Scaphognathinae. Other researchers however, such as
Alexander Kellner Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian geologist and paleontologist who is a leading expert in the field of studying pterosaurs. His research has focused mainly on fossil reptiles from the Cretaceous Period, i ...
and Lü Junchang, have produced
cladistic Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...
analyses showing that ''Sordes'' was much more basal, and not a rhamphorhynchid.


See also

*
List of pterosaur genera This list of pterosaurs is a comprehensive listing of all Genus, genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considere ...
*
Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and Biological taxonomy, taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic Era (ge ...


References


External links


Artistic reconstruction of a ''Sordes pilosus'' at the Fossilsmith StudiosA fossil of a ''Sordes'' at paleo.ru
{{Portalbar, Paleontology Late Jurassic pterosaurs of Asia Novialoids Taxa named by Aleksandr Grigorevich Sharov Fossil taxa described in 1971