Scaphognathinae
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Scaphognathinae
Rhamphorhynchidae is a group of early pterosaurs named after ''Rhamphorhynchus'', that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley.Seeley, H.G. (1870). "The Orithosauria: An Elementary Study of the Bones of Pterodactyles." Cambridge, 135 p. Members of the group possess no more than 11 pairs of teeth in the rostrum, a deltopectoral crest that is constricted at the base but expanded at the distal end, and a bent phalange on the fifth toe. Rhamphorhynchidae traditionally contains two subfamilies: the Rhamphorhynchinae and the Scaphognathinae. While not recovered as distinct clades by all analyses, there do appear to be traits uniting members of each group. Rhamphorhynchines are more common, were lightly built, and had jaws ending in pointed tips that contained more teeth, which are often procumbent (pointed forward). Scaphognathines are comparatively quite rare, were more robust skeletally, and had shorter wing proportions. The br ...
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Klobiodon
''Klobiodon'' is a genus of rhamphorhynchid pterosaur from the Middle Jurassic Taynton Limestone Formation of Oxfordshire, England. Etymology The type species of ''Klobiodon'' is ''Klobiodon rochei''. The generic name ''Klobiodon'' means "small cage tooth," from the Greek κλωβίον, ''klobion'', "little cage", and ὀδών, ''odon'', "tooth", in reference to the large anterior laniaries that appear to form a fish grab, while the specific name ''rochei'' honors the comic book artist Nick Roche for his anatomically correct designs inspired by dinosaurs. History The holotype of ''Klobiodon rochei'', NHMUK PV OR 47991, was first mentioned by George Robert Waterhouse (1878) as part of the collection of the British Museum of Natural History, where he mentions Richard Owen intended to name the specimen ''Pterodactylus raptor'' in an unpublished manuscript. It had been donated by Robert Marsham. Richard Lydekker (1888) referred the specimen to ''Rhamphorhynchus depressirostris'' ( ...
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Fenghuangopterus
''Fenghuangopterus'' is a genus of basal pterosaur that lived in northeastern China during the Middle Jurassic. The type species ''Fenghuangopterus lii'' was in 2010 described and named by Lü Junchang ''et al.'' The generic name is derived from the Fenghuang Mountain and a Latinized Ancient Greek ''pteron'', "wing". The specific name honors Li Xiumei, who donated the fossil. It is known from a single relatively complete, though badly crushed, fossil skeleton, holotype CYGB-0037, recovered from the Tiaojishan Formation of Liaoning Province, about 160 million years old. ''Fenghuangopterus'' is a member of the rhamphorhynchid subfamily Scaphognathinae, which had previously been known only from the Late Jurassic and includes the close relatives ''Scaphognathus'', ''Sordes'' and ''Harpactognathus''. Description ''Fenghuangopterus'' was similar to other scaphognathines in its short, blunt skull with a large antorbital fenestra, and widely spaced, vertically oriented teeth (as oppos ...
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Harpactognathus
''Harpactognathus'' (meaning "seizing/grasping jaw") is a genus of pterosaur found in the Late Jurassic-age Morrison Formation of Albany County, Wyoming, United States. It is based on NAMAL 101, a partial skull consisting of the snout, recovered from near Bone Cabin Quarry in 1996. The specific name honors the discoverer, Joe Gentry, a volunteer for the Western Paleontological Laboratories, in Lehi, Utah.Carpenter, K., Unwin, D.M., Cloward, K., Miles, C.A., and Miles, C. (2003). A new scaphognathine pterosaur from the Upper Jurassic Formation of Wyoming, USA. In: Buffetaut, E., and Mazin, J.- M. (eds.). ''Evolution and Palaeobiology of Pterosaurs.'' Geological Society of London, Special Publications 217:45-54. The holotype of ''Harpactognathus'' consists of a partial rostrum. Its describers found it to be most similar to ''Scaphognathus'' among pterosaurs, albeit substantially larger (estimated skull length of 280–300 mm (11–12 in), estimated wingspan of at least 2 ...
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Dolicorhamphus
''Dolicorhamphus'' is an extinct genus of pterosaurs from the Middle Jurassic Taynton Limestone Formation and Fuller's Earth Formations of England. The genus contains two species, ''D. bucklandii'' and ''D. depressirostris''. Both species of ''Dolicorhamphus'' were assigned to the wastebasket genus '' Rhamphocephalus'', considered a "rhamphorhynchoid" pterosaur. However, the type species of ''Rhamphocephalus'', ''R. prestwichii'', was reinterpreted as a thalattosuchian by O'Sullivan and Martill on 2018, but ''"R." depressirostris'' and ''"R." bucklandii'' were clearly pterosaurian; the former was considered to be in the Scaphognathinae while the latter was only referred to a more general Rhamphorhynchidae. Andres (2021) resurrected the genus ''Dolicorhamphus'' for the two, finding it to be a valid genus of pterosaurs closely related to ''Klobiodon'', although he did not assign a type species. This generic assignment has since been followed by other researchers. ''D. bucklandi'' i ...
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Angustinaripterus
''Angustinaripterus'' was a basal pterosaur, belonging to the breviquartossan family Rhamphorhynchidae (more specifically within the subfamily Rhamphorhynchinae) and discovered at Dashanpu near Zigong in the Sichuan province of China. Discovery and etymology ''Angustinaripterus'' was named in 1983 by He Xinlu. The type species is ''Angustinaripterus longicephalus''. The genus name is derived from Latin ''angustus'', "narrow" and ''naris'', "nostril", combined with Latinized Greek ''pteron'', "wing". The specific name is derived from Latin ''longus'', "long", and Greek ''kephale'', "head". The holotype, ZDM T8001, is a single skull with lower jaws, found in 1981 by researchers from the Zigong Historical Museum of the Salt Industry, in the Xiashaximiao Formation (Bathonian). Description The skull of ''Angustinaripterus'', of which the left side is severely damaged, is very elongated and flat. The back part is missing; in its preserved state it has a length of ; the total le ...
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Early Jurassic
The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma (million years ago), and ends at the start of the Middle Jurassic 174.1 Ma. Certain rocks of marine origin of this age in Europe are called "Lias Group, Lias" and that name was used for the period, as well, in 19th-century geology. In southern Germany rocks of this age are called Black Jurassic. Origin of the name Lias There are two possible origins for the name Lias: the first reason is it was taken by a geologist from an England, English quarryman's dialect pronunciation of the word "layers"; secondly, sloops from north Cornwall, Cornish ports such as Bude would sail across the Bristol Channel to the Vale of Glamorgan to load up with rock from coastal limestone quarries (lias limestone from S ...
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Scaphognathus
''Scaphognathus'' was a pterosaur that lived around Germany during the Late Jurassic. It had a wingspan of 0.9 m (3 ft). Naming The first known ''Scaphognathus'' specimen was described in 1831 by August Goldfuss who mistook the tailless specimen for a new ''Pterodactylus'' species: ''P. crassirostris''. The specific name means "fat snout" in Latin. This specimen was an incomplete adult with a 0.9 m (3 ft) wingspan recovered from the Solnhofen strata near Eichstätt. In 1858 Johann Wagner referred the species to ''Rhamphorhynchus''. After recognising the fundamentally different snout shape, Wagner, after previous failed attempts by Leopold Fitzinger and Christoph Gottfried Andreas Giebel, who used preoccupied names, in 1861 named a distinct genus: ''Scaphognathus'', derived from Greek ''skaphe'', "boat" or "tub", and ''gnathos'', "jaw", in reference to the blunt shape of the lower jaws. In the early twentieth century, the "rhamphorhynchoid" nature of ''S. c ...
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Jianchangnathus
''Jianchangnathus'' is an extinct genus of basal pterosaur from the Middle Jurassic Tiaojishan Formation of northeastern China. Naming ''Jianchangnathus'' was first described and named by Cheng Xin, Wang Xiaolin, Jiang Shunxing and Alexander W.A. Kellner in 2012 and the type species is ''Jianchangnathus robustus''. The generic name combines a reference to Jianchang County with a Greek γνάθος, ''gnathos'', "jaw". The specific name means "robust" in Latin. ''Jianchangnathus'' is known from a single fossil skeleton, holotype IVPP V16866, recovered near Linglongta, in Jianchang County. Description Autapomorphies of ''Jiangchangnathus'' include: a convex top margin of the lower jaw; a large front branch of the jugal; and the first three pairs of teeth of the lower jaws pointing strongly forwards. Its describers found it to share several features with ''Scaphognathus'', including a high front end of the lower jaws, a pear-shaped lower temporal fenestra with the broa ...
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Jianchangopterus
''Jianchangopterus'' is a genus of scaphognathine rhamphorhynchid pterosaur from the Middle Jurassic of western Liaoning, China. ''Jianchangopterus'' is known from a nearly complete skeleton with skull preserved. It was collected from the Tiaojishan Formation. It was first named (after Jianchang County) by Lü Junchang and Bo Xue in 2011 and the type species is ''Jianchangopterus zhaoianus''. See also * List of pterosaur genera * Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs w ... References Middle Jurassic pterosaurs of Asia Rhamphorhynchids Fossil taxa described in 2011 Taxa named by Lü Junchang Paleontology in Liaoning {{Pterosaur-stub ...
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Phylogenetics
In biology, phylogenetics (; from Greek language, Greek wikt:φυλή, φυλή/wikt:φῦλον, φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, Protein, protein Amino acid, amino acid sequences, or Morphology (biology), morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An un ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Family (biology)
Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". What belongs to a family—or if a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, but in plants, they can be characterized on the basis of both vegetative and reproductive features of plant species. Taxonomists often take different positions about descriptions, and there may be no broad consensus across the scientific community for some time. The publishing of new data and opini ...
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