''Jianchangnathus'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of basal

pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...

from the

Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...

Tiaojishan Formation The Tiaojishan Formation is a geological formation in Hebei and Liaoning, People's Republic of China, dating to the middle-late Jurassic period (Bathonian- Oxfordian stages). It is known for its exceptionally preserved fossils, including those of ...

of northeastern

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

Naming

''Jianchangnathus'' was first described and named by Cheng Xin,

Wang Xiaolin Wang may refer to: Names * Wang (surname) (王), a common Chinese surname * Wāng (汪), a less common Chinese surname * Titles in Chinese nobility * A title in Korean nobility * A title in Mongolian nobility Places * Wang River in Thailand ...

, Jiang Shunxing and Alexander W.A. Kellner in

2012 File:2012 Events Collage V3.png, From left, clockwise: The passenger cruise ship Costa Concordia lies capsized after the Costa Concordia disaster; Damage to Casino Pier in Seaside Heights, New Jersey as a result of Hurricane Sandy; People gather ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Jianchangnathus robustus''. The generic name combines a reference to

Jianchang County Jianchang () is a county of Huludao City in the southwest of Liaoning province, China. It is the largest division of Huludao, with an area of , and population of 600,000, located in mountainous terrain west of that city, serviced by China Nationa ...

with a

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

γνάθος, ''gnathos'', "jaw". The specific name means "robust" in

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

. ''Jianchangnathus'' is known from a single fossil skeleton,

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

IVPP The Institute of Vertebrate Paleontology and Paleoanthropology (IVPP; ) of China is a research institution and collections repository for fossils, including many dinosaur and pterosaur specimens (many from the Yixian Formation). As its name suggest ...

V16866, recovered near Linglongta, in Jianchang County.

Description

Autapomorphies In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...

of ''Jiangchangnathus'' include: a convex top margin of the lower jaw; a large front branch of the

jugal The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anatomy ...

; and the first three pairs of teeth of the lower jaws pointing strongly forwards. Its describers found it to share several features with ''

Scaphognathus ''Scaphognathus'' was a pterosaur that lived around Germany during the Late Jurassic. It had a wingspan of 0.9 m (3 ft). Naming The first known ''Scaphognathus'' specimen was described in 1831 by August Goldfuss who mistook the taill ...

'', including a high front end of the lower jaws, a pear-shaped lower temporal fenestra with the broad end below and teeth in the

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...

of the upper jaw that have a space equal to that of three toothsockets between them. Additionally, undescribed fossils of a pterosaur referred to ''Jianchangnathus'' suggest that the color of its pycnofibers was brown.Li, Q., J.A. Clarke, K.-Q. Gao, C.-F. Zhou, Q. Meng, D. Li, L. D'Alba, and M.D. Shawkey. 2014. Melanosome evolution indicates a key physiological shift within feathered dinosaurs. Nature 507: 350-353.

Phylogeny

''Jianchangnathus'' was assigned by the describers to the Scaphognathidae.

References

{{Taxonbar, from=Q2464965 Middle Jurassic pterosaurs of Asia Rhamphorhynchids Fossil taxa described in 2012 Fossils of China

Naming

Description

Phylogeny

See also

References