Sauroplites
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''Sauroplites'' (meaning "saurian hoplite") is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of herbivorous
ankylosaurian Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the Early
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
of
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
.


Discovery and naming

In 1930, the Swedish paleontologist
Anders Birger Bohlin Dr. Anders Birger Bohlin (26 March 1898 – 28 November 1990) was a Swedes, Swedish palaeontologist. As well as his work on dinosaurs and prehistoric mammals, Bohlin was part of the group that established the existence of Peking Man ''(Sinanthr ...
during the Swedish-Chinese expeditions of
Sven Hedin Sven Anders Hedin, KNO1kl RVO,Wennerholm, Eric (1978) ''Sven Hedin – En biografi'', Bonniers, Stockholm (19 February 1865 – 26 November 1952) was a Swedish geographer, topographer, explorer, photographer, travel writer and illustrator ...
discovered an ankylosaurian fossil near Tebch in
Inner Mongolia Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an autonomous region of the People's Republic of China. Its border includes most of the length of China's border with the country of Mongolia. Inner Mongolia also accounts for a ...
.B. Bohlin, 1953, ''Fossil reptiles from Mongolia and Kansu. Reports from the Scientific Expedition to the North-western Provinces of China under Leadership of Dr. Sven Hedin. VI. Vertebrate Palaeontology 6.'' The Sino-Swedish Expedition Publications 37, 113 pp The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
''Sauroplites scutiger'' was named and described by Bohlin in 1953. The generic name is derived from Greek ''sauros'' or ''saura'', "lizard", and ''hoplites'', "
hoplite Hoplites ( ) ( grc, ὁπλίτης : hoplítēs) were citizen-soldiers of Ancient Greece, Ancient Greek Polis, city-states who were primarily armed with spears and shields. Hoplite soldiers used the phalanx formation to be effective in war with ...
, armed foot soldier". The specific name is
new Latin New Latin (also called Neo-Latin or Modern Latin) is the revival of Literary Latin used in original, scholarly, and scientific works since about 1500. Modern scholarly and technical nomenclature, such as in zoological and botanical taxonomy ...
for "shield bearer", in reference to the body armour. At first generally accepted as valid, even though a diagnosis had originally not been provided, ''Sauroplites'' was later often considered a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
'' because it is based on fragmentary material. Some believed it might actually be a specimen of another ankylosaur, ''
Shamosaurus ''Shamosaurus'' is an extinct genus of herbivorous basal ankylosaurid ankylosaur from Early Cretaceous ( Aptian to Albian stage) deposits of Höövör, Mongolia. Discovery and naming In 1977, a Soviet-Mongolian expedition discovered the ske ...
''. However, in 2014
Victoria Megan Arbour Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
discovered a clear unique trait,
autapomorphy In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...
: the sacral or pelvic shield shows rosettes with a large central osteoderm surrounded by a single ring of smaller scutes. Other species have multiple or irregular rings. She concluded that ''Sauroplites'' was a valid
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
.Arbour, Victoria Megan, 2014, ''Systematics, evolution, and biogeography of the ankylosaurid dinosaurs''. Ph.D thesis, University of Alberta The specimens were not given an inventory number and are today lost, though some casts are present in the
American Museum of Natural History The American Museum of Natural History (abbreviated as AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. In Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 26 inter ...
as specimen AMNH 2074. They were found in a layer of the
Zhidan Group The Zhidan Group is a geological formation in China whose strata date back to the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution (Earl ...
, probably dating from the
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
to
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
stages Stage or stages may refer to: Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly British theatre newspaper * S ...
. The carcass had been deposited on its back and the bones had been eroded away, apart from some ribs and perhaps a piece of an
ischium The ischium () form ...
, leaving parts of the body armour in a largely articulated position.


Description

The large central osteoderms of the rosettes are rather flat and have a diameter of ten centimetres. More to the front oval osteoderms, with an asymmetric low keel, cover the back, with a length of up to forty centimetres. Thirty centimetre osteoderms cover the sides.


Classification

Bohlin placed ''Sauroplites'' in the
Ankylosauridae Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Pal ...
. However, Arbour (2014) considered it possible that it was a member of the
Nodosauridae Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, wer ...
in view of the possession of a sacral shield consisting of fused rosettes which is unknown with unequivocal ankylosaurids, even though nodosaurid remains from Asia are rare and contentious. In a
cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
analysis she performed, ''Sauroplites'' was recovered as a nodosaurid. In 2018, Rivera-Sylva and colleagues suggested that ''Sauroplites'' belonged to a clade of basal nodosaurids that also contained ''
Dongyangopelta ''Dongyangopelta'' (meaning "Dongyang shield") is an extinct genus of nodosaurid, ankylosaurian, dinosaur known from the Early to Late Cretaceous Chaochuan Formation (Albian or Cenomanian stage) of Dongyang, Zhejiang Province, China. The genus c ...
'' and ''
Mymoorapelta ''Mymoorapelta'' (Meaning "Vannetta Moore and Pete and Marilyn Mygatt's shield" after a combination of the names of the discoverers of the Mygatt-Moore Quarry that fossils were originally collected from, and pɛltə "shield") is a monospecific ...
'', Their cladogram is shown below:


See also

*
Timeline of ankylosaur research This timeline of ankylosaur research is a chronological listing of events in the history of paleontology focused on the ankylosaurs, quadrupedal herbivorous dinosaurs who were protected by a covering bony plates and spikes and sometimes by a club ...


References

{{Taxonbar, from=Q144163 Nodosaurids Early Cretaceous dinosaurs of Asia Taxa named by Birger Bohlin Paleontology in Inner Mongolia Ornithischian genera