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Autapomorphy
In phylogenetics, an autapomorphy is a distinctive feature, known as a Synapomorphy, derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or Outgroup (cladistics), outgroup taxa, not even those most closely related to the focal taxon (which may be a species, family (biology), family or in general any clade). It can therefore be considered as an apomorphy in relation to a single taxon. The word ''autapomorphy'', introduced in 1950 by German entomology, entomologist Willi Hennig, is derived from the Greek language, Greek words αὐτός, ''autos'' "self"; ἀπό, ''apo'' "away from"; and μορφή, ''morphḗ'' = "shape". Discussion Because autapomorphies are only present in a single taxon, they do not convey information about relationship. Therefore, autapomorphies are not useful to infer phylogenetic relationships. However, autapomorphy, like synapomorphy and plesiomorphy is a relative concept depending on the ta ...
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Apomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel Phenotypic trait, character or character state that has evolution, evolved from its ancestral form (or Plesiomorphy and symplesiomorphy, plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxon, taxa and is therefore Hypothesis#Scientific hypothesis, hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies Homology (biology), homology. Examples of apomorphy are the presence of Terrestrial locomotion#Posture, erect gait, fur, Evolution of mammalian auditory ossicles, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a Terrestrial locomotion#Posture, sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word ...
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Taxon
In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and given a particular ranking, especially if and when it is accepted or becomes established. It is very common, however, for taxonomists to remain at odds over what belongs to a taxon and the criteria used for inclusion, especially in the context of rank-based (" Linnaean") nomenclature (much less so under phylogenetic nomenclature). If a taxon is given a formal scientific name, its use is then governed by one of the nomenclature codes specifying which scientific name is correct for a particular grouping. Initial attempts at classifying and ordering organisms (plants and animals) were presumably set forth in prehistoric times by hunter-gatherers, as suggested by the fairly sophisticated folk taxonomies. Much later, Aristotle, and later st ...
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Ophidia
__FORCETOC__ Ophidia (also known as Pan-Serpentes) is a group of squamate reptiles including modern snakes and reptiles more closely related to snakes than to other living groups of lizards. Ophidia was defined as the "most recent common ancestor of ''Pachyrhachis'' and Serpentes (modern snakes), and all its descendants" by Lee and Caldwell (1998: 1551). The clade name Ophidia derives from the Ancient Greek word (), meaning "small snake". Evolution Modern snakes are thought to have evolved from either burrowing or aquatic lizards during the mid-Cretaceous period, and the earliest known fossils date to around 112 Ma ago. However, the relationship between modern snake and more primitive snake ancestors, many of which retained hind limbs, is less clear. While many of these "stem-snakes" are known from Mesozoic fossils, some of them may be descendants of the earliest true snakes rather than more primitive lineages. Below is a cladogram modified from a study by Wilson ''et al.'' ...
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Symplesiomorphy
In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, and synapomorphy all mean a trait shared between species because they share an ancestral species. Apomorphic and synapomorphic characteristics convey much information about evolutionary clades and can be used to define taxa. However, plesiomorphic and symplesiomorphic characteristics cannot. The term ''symplesiomorphy'' was introduced in 1950 by German entomologist Willi Hennig. Examples A backbone is a plesiomorphic trait shared by birds and mammals, and does not help in placing an animal in one or the other of these two clades. Birds and mammals share this trait because both clades are descended from the same far distant ancestor. Other clades, e.g. snakes, lizards, turtles, fish, frogs, all have backbones and none are either birds n ...
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Homology (biology)
In biology, homology is similarity in anatomical structures or genes between organisms of different taxa due to shared ancestry, ''regardless'' of current functional differences. Evolutionary biology explains homologous structures as retained heredity from a common descent, common ancestor after having been subjected to adaptation (biology), adaptive modifications for different purposes as the result of natural selection. The term was first applied to biology in a non-evolutionary context by the anatomist Richard Owen in 1843. Homology was later explained by Charles Darwin's theory of evolution in 1859, but had been observed before this from Aristotle's biology onwards, and it was explicitly analysed by Pierre Belon in 1555. A common example of homologous structures is the forelimbs of vertebrates, where the bat wing development, wings of bats and origin of avian flight, birds, the arms of primates, the front flipper (anatomy), flippers of whales, and the forelegs of quadrupedalis ...
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Homoplasy
Homoplasy, in biology and phylogenetics, is the term used to describe a feature that has been gained or lost independently in separate lineages over the course of evolution. This is different from homology, which is the term used to characterize the similarity of features that can be parsimoniously explained by common ancestry. Homoplasy can arise from both similar selection pressures acting on adapting species, and the effects of genetic drift. Most often, homoplasy is viewed as a similarity in morphological characters. However, homoplasy may also appear in other character types, such as similarity in the genetic sequence, life cycle types or even behavioral traits. Etymology The term homoplasy was first used by Ray Lankester in 1870. The corresponding adjective is either ''homoplasic'' or ''homoplastic''. It is derived from the two Ancient Greek words (), meaning "similar, alike, the same", and (), meaning "to shape, to mold". Parallelism and convergence Parallel and ...
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Convergent Evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last common ancestor of those groups. The cladistic term for the same phenomenon is Cladogram#Homoplasies, homoplasy. The recurrent evolution of flight is a classic example, as flying pterygota, insects, birds, pterosaurs, and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are ''analogous'', whereas ''homology (biology), homologous'' structures or traits have a common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions. The opposite of convergence is divergent evolution, where related species evolve different trai ...
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Phenotypic Trait
A phenotypic trait, simply trait, or character state is a distinct variant of a phenotypic characteristic of an organism; it may be either inherited or determined environmentally, but typically occurs as a combination of the two.Lawrence, Eleanor (2005) ''Henderson's Dictionary of Biology''. Pearson, Prentice Hall. For example, having eye color is a ''character'' of an organism, while blue, brown and hazel versions of eye color are ''traits''. The term ''trait'' is generally used in genetics, often to describe the phenotypic expression of different combinations of alleles in different individual organisms within a single population, such as the famous purple vs. white flower coloration in Gregor Mendel's pea plants. By contrast, in systematics, the term ''character state'' is employed to describe features that represent fixed diagnostic differences among taxa, such as the absence of tails in great apes, relative to other primate groups. Definition A phenotypic trait is ...
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Cladistics
Cladistics ( ; from Ancient Greek 'branch') is an approach to Taxonomy (biology), biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived (phylogenetics), derived characteristics (synapomorphies) that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose Phenotypic trait, character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used Paraphyly, paraphyletically, outside of cladistics, e.g. as a 'E ...
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