The palm weevil ''Rhynchophorus ferrugineus'' is one of two species of
snout beetle
The Curculionidae are a family of weevils, commonly called snout beetles or true weevils. They are one of the largest animal families, with 6,800 genera and 83,000 species described worldwide. They are the sister group to the family Brentidae.
T ...
known as the red palm weevil, Asian palm weevil or sago palm weevil. The adult beetles are relatively large, ranging between long, and are usually a rusty red colour—but many colour variants exist and have often been classified as different species (e.g., ''
Rhynchophorus vulneratus''). Weevil larvae can excavate holes in the trunks of
palm tree
The Arecaceae is a family of perennial flowering plants in the monocot order Arecales. Their growth form can be climbers, shrubs, tree-like and stemless plants, all commonly known as palms. Those having a tree-like form are called palm trees ...
s up to long, thereby weakening and eventually killing the host plant. As a result, the weevil is considered a major pest in palm plantations, including the
coconut palm
The coconut tree (''Cocos nucifera'') is a member of the palm tree family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or the f ...
,
date palm
''Phoenix dactylifera'', commonly known as date or date palm, is a flowering plant species in the palm family, Arecaceae, cultivated for its edible sweet fruit called dates. The species is widely cultivated across northern Africa, the Middle Eas ...
and
oil palm
''Elaeis'' () is a genus of palms containing two species, called oil palms. They are used in commercial agriculture in the production of palm oil. The African oil palm ''Elaeis guineensis'' (the species name ''guineensis'' referring to its co ...
.
Originally from
tropical Asia
Tropical Asia refers to the entirety of the areas in Asia with a tropical climate. These areas are of geographic and economic importance due to their natural resources and biodiversity, which include many species of agricultural value. There are ...
, the red palm weevil has spread to Africa and Europe, reaching the
Mediterranean
The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...
in the 1980s. It was first recorded in
Spain
, image_flag = Bandera de España.svg
, image_coat = Escudo de España (mazonado).svg
, national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond")
, national_anthem = (English: "Royal March")
, i ...
in 1994,
and in
France
France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
in 2006.
Additional infestations have been located in Malta, Italy (Tuscany, Sicily, Campania, Sardinia, Lazio, Marche, Puglia and Liguria), Croatia and Montenegro. It is also well established throughout most of Portugal, especially in the South. It also has established in Morocco, Tunisia, and other North African countries.
The weevil was first reported in the Americas on
Curaçao
Curaçao ( ; ; pap, Kòrsou, ), officially the Country of Curaçao ( nl, Land Curaçao; pap, Pais Kòrsou), is a Lesser Antilles island country in the southern Caribbean Sea and the Dutch Caribbean region, about north of the Venezuela coast ...
in January 2009 and sighted the same year in
Aruba
Aruba ( , , ), officially the Country of Aruba ( nl, Land Aruba; pap, Pais Aruba) is a constituent country of the Kingdom of the Netherlands physically located in the mid-south of the Caribbean Sea, about north of the Venezuela peninsula of ...
. It was reported in the United States at
Laguna Beach, California
Laguna Beach (; ''Laguna'', Spanish language, Spanish for "Lagoon") is a seaside resort city located in southern Orange County, California, in the United States. It is known for its mild year-round climate, scenic coves, environmental preservat ...
late in 2010 but this was a misidentification of the closely related species, ''
Rhynchophorus vulneratus'', and it did not become established.
Larvae of ''Rhynchophorus ferrugineus'' are considered a delicacy in
Southeast Asian cuisine
This is a list of Asian cuisines, by region. A cuisine is a characteristic style of cooking practices and traditions, usually associated with a specific culture or region. Asia, being the largest, most populous and culturally diverse continent, ...
. In some regions, however, larvae farming is strictly prohibited to prevent the potential devastation of plantation crops.
Taxonomy
Primarily due to the existence of numerous color forms across their ranges, the taxonomy and classification of red palm weevils has undergone a number of changes in understanding and circumscription. As such, the information in the literature should be viewed as a compilation of data which may apply to both species, depending primarily upon the biogeography; accordingly, the vast majority of publications presumably do refer to ''R. ferrugineus'' rather than ''vulneratus'', as the former is by far the most widely invasive. The most recent genus-level revision in 1966
recognized two species of red palm weevil, ''ferrugineus'' and ''vulneratus'', and for decades these were interpreted as separate taxa. A genetic study in 2004
concluded that ''vulneratus'' was not distinct from ''ferrugineus'', and treated them as synonyms, a view that was accepted until 2013, when yet another genetic study
came to the opposite conclusion, based on more comprehensive geographic sampling. Accordingly, the "red palm weevil" species that appeared in the US was ''vulneratus'' rather than ''ferrugineus'', though the latter is the invading species in all of the other global introductions.
Distribution
Range
The native range of this species is considered to include Bangladesh, Cambodia, China, India, Japan, Laos, Pakistan, Philippines, Sri Lanka, Taiwan, and Vietnam; records from Indonesia, Malaysia, Myanmar, Singapore, and Thailand largely or exclusively refer to ''R. vulneratus''.
''R. ferrugineus'' has now been reported and confirmed from Albania, Algeria, Aruba, Bahrain, Bosnia and Herzegovina, Croatia, Curaçao, Cyprus, Egypt, France (incl. Corsica), Greece, Israel, Italy (incl. Sicily and Sardinia), Jordan, Kuwait, Libya, Malta, Monaco, Montenegro, Morocco, Oman, Palestine, Portugal (incl. Madeira), Qatar, Saudi Arabia, Slovenia, Spain (incl. the Balearic and Canary islands), Syria, Tunisia, Turkey, and United Arab Emirates. Records from Australia, Papua New Guinea, Samoa, the Solomon Islands, and Vanuatu
have not been confirmed and are likely to be specimens of ''
Rhynchophorus bilineatus
''Rhynchophorus bilineatus'', common name Black Palm Weevil, is a species of beetles belonging to the family Curculionidae.
Description
''Rhynchophorus bilineatus'' can reach a body length of about 40mm. These large beetles are considered a maj ...
'', a closely related species indigenous to the region.
CABI ISC range list
=Africa
=
* absent from
Algeria
)
, image_map = Algeria (centered orthographic projection).svg
, map_caption =
, image_map2 =
, capital = Algiers
, coordinates =
, largest_city = capital
, relig ...
*
Djibouti
Djibouti, ar, جيبوتي ', french: link=no, Djibouti, so, Jabuuti officially the Republic of Djibouti, is a country in the Horn of Africa, bordered by Somalia to the south, Ethiopia to the southwest, Eritrea in the north, and the Red ...
*
Egypt
Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Mediter ...
*
Libya
Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to Egypt–Libya bo ...
*
Mauritania
Mauritania (; ar, موريتانيا, ', french: Mauritanie; Berber: ''Agawej'' or ''Cengit''; Pulaar: ''Moritani''; Wolof: ''Gànnaar''; Soninke:), officially the Islamic Republic of Mauritania ( ar, الجمهورية الإسلامية ...
*
Morocco
Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
*
Tunisia
)
, image_map = Tunisia location (orthographic projection).svg
, map_caption = Location of Tunisia in northern Africa
, image_map2 =
, capital = Tunis
, largest_city = capital
, ...
*
Nigeria
Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
=Asia
=
*
Bahrain
Bahrain ( ; ; ar, البحرين, al-Bahrayn, locally ), officially the Kingdom of Bahrain, ' is an island country in Western Asia. It is situated on the Persian Gulf, and comprises a small archipelago made up of 50 natural islands and an ...
*
Bangladesh
Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
*
Cambodia
Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand t ...
*
China
China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
since 1997, first detection in Guangdong.
[ .] The model of Ge et al 2015 predicts a wide area of invasion based on
climate
Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologic ...
suitability, throughout the
south
South is one of the cardinal directions or Points of the compass, compass points. The direction is the opposite of north and is perpendicular to both east and west.
Etymology
The word ''south'' comes from Old English ''sūþ'', from earlier Pro ...
, north to the
east
East or Orient is one of the four cardinal directions or points of the compass. It is the opposite direction from west and is the direction from which the Sun rises on the Earth.
Etymology
As in other languages, the word is formed from the fa ...
and west to central China.
**
Fujian
Fujian (; alternately romanized as Fukien or Hokkien) is a province on the southeastern coast of China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its capi ...
**
Guangdong
Guangdong (, ), alternatively romanized as Canton or Kwangtung, is a coastal province in South China on the north shore of the South China Sea. The capital of the province is Guangzhou. With a population of 126.01 million (as of 2020) ...
since 1997
**
Guangxi
Guangxi (; ; Chinese postal romanization, alternately romanized as Kwanghsi; ; za, Gvangjsih, italics=yes), officially the Guangxi Zhuang Autonomous Region (GZAR), is an Autonomous regions of China, autonomous region of the People's Republic ...
**
Hainan
Hainan (, ; ) is the smallest and southernmost province of the People's Republic of China (PRC), consisting of various islands in the South China Sea. , the largest and most populous island in China,The island of Taiwan, which is slightly l ...
**
Jiangsu
Jiangsu (; ; pinyin: Jiāngsū, Postal romanization, alternatively romanized as Kiangsu or Chiangsu) is an Eastern China, eastern coastal Provinces of the People's Republic of China, province of the China, People's Republic of China. It is o ...
**
Tibet
Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
**
Yunnan
Yunnan , () is a landlocked Provinces of China, province in Southwest China, the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is ...
**
Zhejiang
Zhejiang ( or , ; , also romanized as Chekiang) is an eastern, coastal province of the People's Republic of China. Its capital and largest city is Hangzhou, and other notable cities include Ningbo and Wenzhou. Zhejiang is bordered by Jiang ...
*
Georgia
Georgia most commonly refers to:
* Georgia (country), a country in the Caucasus region of Eurasia
* Georgia (U.S. state), a state in the Southeast United States
Georgia may also refer to:
Places
Historical states and entities
* Related to the ...
*
Hong Kong
Hong Kong ( (US) or (UK); , ), officially the Hong Kong Special Administrative Region of the People's Republic of China ( abbr. Hong Kong SAR or HKSAR), is a city and special administrative region of China on the eastern Pearl River Delt ...
*
India
India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
**
Andaman and Nicobar Islands
The Andaman and Nicobar Islands is a union territory of India consisting of 572 islands, of which 37 are inhabited, at the junction of the Bay of Bengal and the Andaman Sea. The territory is about north of Aceh in Indonesia and separated f ...
**
Andhra Pradesh
Andhra Pradesh (, abbr. AP) is a state in the south-eastern coastal region of India. It is the seventh-largest state by area covering an area of and tenth-most populous state with 49,386,799 inhabitants. It is bordered by Telangana to the ...
**
Assam
Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...
**
Bihar
Bihar (; ) is a state in eastern India. It is the 2nd largest state by population in 2019, 12th largest by area of , and 14th largest by GDP in 2021. Bihar borders Uttar Pradesh to its west, Nepal to the north, the northern part of West Be ...
**
Daman and Diu
Daman and Diu (; ) was a former union territory in northwestern India. With an area of , it was the smallest administrative subdivision of India on the mainland. The territory comprised two districts, Damaon and Dio island, geographically sep ...
**
Goa
Goa () is a state on the southwestern coast of India within the Konkan region, geographically separated from the Deccan highlands by the Western Ghats. It is located between the Indian states of Maharashtra to the north and Karnataka to the ...
**
Gujarat
Gujarat (, ) is a state along the western coast of India. Its coastline of about is the longest in the country, most of which lies on the Kathiawar peninsula. Gujarat is the fifth-largest Indian state by area, covering some ; and the ninth ...
**
Karnataka
Karnataka (; ISO: , , also known as Karunāḍu) is a state in the southwestern region of India. It was formed on 1 November 1956, with the passage of the States Reorganisation Act. Originally known as Mysore State , it was renamed ''Karnat ...
**
Kerala
Kerala ( ; ) is a state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile regions of Cochin, Malabar, South ...
**
Maharashtra
Maharashtra (; , abbr. MH or Maha) is a states and union territories of India, state in the western India, western peninsular region of India occupying a substantial portion of the Deccan Plateau. Maharashtra is the List of states and union te ...
**
Meghalaya
Meghalaya (, or , meaning "abode of clouds"; from Sanskrit , "cloud" + , "abode") is a states and union territories of India, state in northeastern India. Meghalaya was formed on 21 January 1972 by carving out two districts from the state of As ...
**
Odisha
Odisha (English: , ), formerly Orissa ( the official name until 2011), is an Indian state located in Eastern India. It is the 8th largest state by area, and the 11th largest by population. The state has the third largest population of ...
**
Tamil Nadu
Tamil Nadu (; , TN) is a States and union territories of India, state in southern India. It is the List of states and union territories of India by area, tenth largest Indian state by area and the List of states and union territories of India ...
**
Tripura
Tripura (, Bengali: ) is a state in Northeast India. The third-smallest state in the country, it covers ; and the seventh-least populous state with a population of 36.71 lakh ( 3.67 million). It is bordered by Assam and Mizoram to the east a ...
**
Uttar Pradesh
Uttar Pradesh (; , 'Northern Province') is a state in northern India. With over 200 million inhabitants, it is the most populated state in India as well as the most populous country subdivision in the world. It was established in 1950 ...
**
West Bengal
West Bengal (, Bengali: ''Poshchim Bongo'', , abbr. WB) is a state in the eastern portion of India. It is situated along the Bay of Bengal, along with a population of over 91 million inhabitants within an area of . West Bengal is the fourt ...
* absent from
Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
*
Iran
Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
*
Iraq
Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq ...
*
Israel
Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
*
Japan
Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
**
Honshu
, historically called , is the largest and most populous island of Japan. It is located south of Hokkaidō across the Tsugaru Strait, north of Shikoku across the Inland Sea, and northeast of Kyūshū across the Kanmon Straits. The island separ ...
**
Kyushu
is the third-largest island of Japan's five main islands and the most southerly of the four largest islands ( i.e. excluding Okinawa). In the past, it has been known as , and . The historical regional name referred to Kyushu and its surroun ...
**
Ryukyu Islands
The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Taiwan: the Ōsumi, Tokara, Amami, Okinawa, and Sakishima Islands (further divided into the Miyako and Yaeyama Islands), with Yonaguni ...
*
Jordan
Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...
*
Kuwait
Kuwait (; ar, الكويت ', or ), officially the State of Kuwait ( ar, دولة الكويت '), is a country in Western Asia. It is situated in the northern edge of Eastern Arabia at the tip of the Persian Gulf, bordering Iraq to the nort ...
*
Lebanon
Lebanon ( , ar, لُبْنَان, translit=lubnān, ), officially the Republic of Lebanon () or the Lebanese Republic, is a country in Western Asia. It is located between Syria to the north and east and Israel to the south, while Cyprus li ...
*
Laos
Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
* absent from
Malaysia
Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
*
Myanmar
Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
*
Oman
Oman ( ; ar, عُمَان ' ), officially the Sultanate of Oman ( ar, سلْطنةُ عُمان ), is an Arabian country located in southwestern Asia. It is situated on the southeastern coast of the Arabian Peninsula, and spans the mouth of t ...
*
Pakistan
Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
*
Philippines
The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no),
* bik, Republika kan Filipinas
* ceb, Republika sa Pilipinas
* cbk, República de Filipinas
* hil, Republ ...
*
Qatar
Qatar (, ; ar, قطر, Qaṭar ; local vernacular pronunciation: ), officially the State of Qatar,) is a country in Western Asia. It occupies the Qatar Peninsula on the northeastern coast of the Arabian Peninsula in the Middle East; it sh ...
*
Saudi Arabia
Saudi Arabia, officially the Kingdom of Saudi Arabia (KSA), is a country in Western Asia. It covers the bulk of the Arabian Peninsula, and has a land area of about , making it the fifth-largest country in Asia, the second-largest in the A ...
* absent from
Singapore
Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Southeast Asia. It lies about one degree of latitude () north of the equator, off the southern tip of the Malay Peninsula, borde ...
*
Sri Lanka
Sri Lanka (, ; si, ශ්රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
*
Syria
Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
*
Taiwan
Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
*
Thailand
Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
*
Turkey
Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
*
United Arab Emirates
The United Arab Emirates (UAE; ar, اَلْإِمَارَات الْعَرَبِيَة الْمُتَحِدَة ), or simply the Emirates ( ar, الِْإمَارَات ), is a country in Western Asia (The Middle East). It is located at th ...
*
Vietnam
Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
*
Yemen
Yemen (; ar, ٱلْيَمَن, al-Yaman), officially the Republic of Yemen,, ) is a country in Western Asia. It is situated on the southern end of the Arabian Peninsula, and borders Saudi Arabia to the Saudi Arabia–Yemen border, north and ...
**
Socotra
Socotra or Soqotra (; ar, سُقُطْرَىٰ ; so, Suqadara) is an island of the Republic of Yemen in the Indian Ocean, under the ''de facto'' control of the UAE-backed Southern Transitional Council, a secessionist participant in Yemen’s ...
=Europe
=
*
Albania
Albania ( ; sq, Shqipëri or ), or , also or . officially the Republic of Albania ( sq, Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic and Ionian Seas within the Mediterranean Sea and shares ...
* absent from
Austria
Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
* absent from
Belgium
Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to th ...
*
Bosnia and Herzegovina
Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and often known informally as Bosnia, is a country at the crossroads of south and southeast Europe, located in the Balkans. Bosnia and H ...
*
Bulgaria
Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Macedon ...
*
Croatia
, image_flag = Flag of Croatia.svg
, image_coat = Coat of arms of Croatia.svg
, anthem = "Lijepa naša domovino"("Our Beautiful Homeland")
, image_map =
, map_caption =
, capit ...
*
Cyprus
Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is geo ...
* absent from
Denmark
)
, song = ( en, "King Christian stood by the lofty mast")
, song_type = National and royal anthem
, image_map = EU-Denmark.svg
, map_caption =
, subdivision_type = Sovereign state
, subdivision_name = Danish Realm, Kingdom of Denmark
...
* absent from
Finland
Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of B ...
*
France
France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
**
Corsica
Corsica ( , Upper , Southern ; it, Corsica; ; french: Corse ; lij, Còrsega; sc, Còssiga) is an island in the Mediterranean Sea and one of the 18 regions of France. It is the fourth-largest island in the Mediterranean and lies southeast of ...
*
Greece
Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
**
Crete
Crete ( el, Κρήτη, translit=, Modern: , Ancient: ) is the largest and most populous of the Greek islands, the 88th largest island in the world and the fifth largest island in the Mediterranean Sea, after Sicily, Sardinia, Cyprus, and ...
*
Italy
Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
**
Sardinia
Sardinia ( ; it, Sardegna, label=Italian, Corsican and Tabarchino ; sc, Sardigna , sdc, Sardhigna; french: Sardaigne; sdn, Saldigna; ca, Sardenya, label=Algherese and Catalan) is the second-largest island in the Mediterranean Sea, after ...
**
Sicily
(man) it, Siciliana (woman)
, population_note =
, population_blank1_title =
, population_blank1 =
, demographics_type1 = Ethnicity
, demographics1_footnotes =
, demographi ...
*
Malta
Malta ( , , ), officially the Republic of Malta ( mt, Repubblika ta' Malta ), is an island country in the Mediterranean Sea. It consists of an archipelago, between Italy and Libya, and is often considered a part of Southern Europe. It lies ...
*
Montenegro
)
, image_map = Europe-Montenegro.svg
, map_caption =
, image_map2 =
, capital = Podgorica
, coordinates =
, largest_city = capital
, official_languages = M ...
* absent from the
Netherlands
)
, anthem = ( en, "William of Nassau")
, image_map =
, map_caption =
, subdivision_type = Sovereign state
, subdivision_name = Kingdom of the Netherlands
, established_title = Before independence
, established_date = Spanish Netherl ...
* absent from
Poland
Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...
*
Portugal
Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
**
Madeira
)
, anthem = ( en, "Anthem of the Autonomous Region of Madeira")
, song_type = Regional anthem
, image_map=EU-Portugal_with_Madeira_circled.svg
, map_alt=Location of Madeira
, map_caption=Location of Madeira
, subdivision_type=Sovereign st ...
*
Russia
Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
**
Southern Russia
Southern Russia or the South of Russia (russian: Юг России, ''Yug Rossii'') is a colloquial term for the southernmost geographic portion of European Russia generally covering the Southern Federal District and the North Caucasian Feder ...
* eradicated from
Slovenia
Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, an ...
*
Spain
, image_flag = Bandera de España.svg
, image_coat = Escudo de España (mazonado).svg
, national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond")
, national_anthem = (English: "Royal March")
, i ...
**
Balearic Islands
The Balearic Islands ( es, Islas Baleares ; or ca, Illes Balears ) are an archipelago in the Balearic Sea, near the eastern coast of the Iberian Peninsula. The archipelago is an autonomous community and a province of Spain; its capital is ...
** eradicated from the
Canary Islands
The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
* absent from
Ukraine
Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
* absent from the
United Kingdom
The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
** absent from
England
England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
=North America
=
*
Aruba
Aruba ( , , ), officially the Country of Aruba ( nl, Land Aruba; pap, Pais Aruba) is a constituent country of the Kingdom of the Netherlands physically located in the mid-south of the Caribbean Sea, about north of the Venezuela peninsula of ...
*
Curaçao
Curaçao ( ; ; pap, Kòrsou, ), officially the Country of Curaçao ( nl, Land Curaçao; pap, Pais Kòrsou), is a Lesser Antilles island country in the southern Caribbean Sea and the Dutch Caribbean region, about north of the Venezuela coast ...
*
Netherlands Antilles
nl, In vrijheid verenigd"Unified by freedom"
, national_anthem =
, common_languages = Dutch English Papiamento
, demonym = Netherlands Antillean
, capital = Willemstad
, year_start = 1954
, year_end = 2010
, date_start = 15 December
, ...
* eradicated from the
United States
The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
** eradicated from
California
California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
=Oceania
=
*
Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
unassessed
** absent from
New South Wales
)
, nickname =
, image_map = New South Wales in Australia.svg
, map_caption = Location of New South Wales in AustraliaCoordinates:
, subdivision_type = Country
, subdivision_name = Australia
, established_title = Before federation
, es ...
** absent from the
Northern Territory
The Northern Territory (commonly abbreviated as NT; formally the Northern Territory of Australia) is an states and territories of Australia, Australian territory in the central and central northern regions of Australia. The Northern Territory ...
**
Queensland
)
, nickname = Sunshine State
, image_map = Queensland in Australia.svg
, map_caption = Location of Queensland in Australia
, subdivision_type = Country
, subdivision_name = Australia
, established_title = Before federation
, established_ ...
unassessed
** absent from
South Australia
South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...
** absent from
Tasmania
)
, nickname =
, image_map = Tasmania in Australia.svg
, map_caption = Location of Tasmania in AustraliaCoordinates:
, subdivision_type = Country
, subdi ...
** absent from
Victoria
Victoria most commonly refers to:
* Victoria (Australia), a state of the Commonwealth of Australia
* Victoria, British Columbia, provincial capital of British Columbia, Canada
* Victoria (mythology), Roman goddess of Victory
* Victoria, Seychelle ...
** absent from
Western Australia
Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
* absent from
Papua New Guinea
Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
*
Samoa
Samoa, officially the Independent State of Samoa; sm, Sāmoa, and until 1997 known as Western Samoa, is a Polynesian island country consisting of two main islands (Savai'i and Upolu); two smaller, inhabited islands (Manono Island, Manono an ...
unassessed
*
Solomon Islands
Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capita ...
unassessed
*
Vanuatu
Vanuatu ( or ; ), officially the Republic of Vanuatu (french: link=no, République de Vanuatu; bi, Ripablik blong Vanuatu), is an island country located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of no ...
unassessed
Hosts
This species of red palm weevil is reported to attack 19 palm species
and is the worst such pest in the world.
Although the weevil was first reported on
coconut
The coconut tree (''Cocos nucifera'') is a member of the palm tree family ( Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or the ...
in Southeast Asia, it has gained a foothold on
date palm
''Phoenix dactylifera'', commonly known as date or date palm, is a flowering plant species in the palm family, Arecaceae, cultivated for its edible sweet fruit called dates. The species is widely cultivated across northern Africa, the Middle Eas ...
over the last two decades in several Middle Eastern countries, and then expanded its range to Africa and Europe.
This expansion has been due to the movement of infested planting material from contaminated to uninfected areas.
In the Mediterranean region, the red palm weevil also severely damages ''
Phoenix canariensis
''Phoenix canariensis'', the Canary Island date palm or pineapple palm, is a species of flowering plant in the palm family Arecaceae, native to the Canary Islands off the coast of Morocco. It is a relative of ''Phoenix dactylifera'', the true ...
''. Currently, the pest is reported in almost 15% of the global coconut-growing countries and in nearly 50% of the date palm-growing countries.
All known hosts of ''R. ferrugineus'' as compiled from sources by CABI ISC are: ''
Areca catechu
''Areca catechu'' is a species of palm which grows in much of the tropical Pacific, Asia, and parts of east Africa. The palm is believed to have originated in the Philippines, but is widespread in cultivation and is considered naturalized in so ...
'', ''
Arenga pinnata
''Arenga pinnata'' (syn. ''Arenga saccharifera'') is an economically important feather palm native to tropical Asia, from eastern India east to Malaysia, Indonesia, and the Philippines in the east. Common names include sugar palm, areng palm (al ...
'', ''
Borassus flabellifer
''Borassus flabellifer'', commonly known as doub palm, palmyra palm, tala or tal palm, toddy palm, wine palm or ice apple, is native to South Asia (especially in Bangladesh & South India) and Southeast Asia. It is reportedly naturalized in Socot ...
'', ''
Brahea armata
''Brahea armata'', commonly known as Mexican blue palm or blue hesper palm, is a large evergreen tree of the palm family Arecaceae, endemic to Baja California, Mexico. It is widely planted as an ornamental.
Description
It grows to a height o ...
'', ''
Brahea edulis
''Brahea'' is a genus of palms in the family Arecaceae. They are commonly referred to as hesper palms and are endemic to Mexico and Central America.Govaerts, R. & Dransfield, J. (2005). World Checklist of Palms: 1-223. The Board of Trustees of th ...
'', ''
Butia capitata
''Butia capitata'', also known as jelly palm, is a '' Butia'' palm native to the states of Minas Gerais and Goiás in Brazil. It is known locally as ''coquinho-azedo'' or ''butiá'' in (northern) Minas Gerais.Fruits of Butia capitata (Mart.) Becc ...
'', ''
Calamus
Calamus may refer to:
Botany and zoology
* ''Calamus'' (fish), a genus of fish in the family Sparidae
* ''Calamus'' (palm), a genus of rattan palms
* Calamus, the hollow shaft of a feather, also known as the quill
* '' Acorus calamus'', the swe ...
merrillii'', ''
Caryota cumingii
''Caryota'' is a genus of palm trees. They are often known as fishtail palms because of the shape of their leaves. There are about 13 species native to Asia ( China, India, Indonesia, etc.), northern Australia, and the South Pacific. One of ...
'', ''
Caryota maxima
''Caryota'' is a genus of palm trees. They are often known as fishtail palms because of the shape of their leaves. There are about 13 species native to Asia ( China, India, Indonesia, etc.), northern Australia, and the South Pacific. One of ...
'', ''
Caryota urens'', ''
Chamaerops humilis
''Chamaerops'' is a genus of flowering plants in the family Arecaceae. The only currently fully accepted species is ''Chamaerops humilis'', variously called European fan palm or the Mediterranean dwarf palm. It is one of the most hardy palms, ...
'', ''
Cocos nucifera
The coconut tree (''Cocos nucifera'') is a member of the palm tree family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or the f ...
'', ''
Corypha umbraculifera
''Corypha umbraculifera'', the talipot palm, is a species of palm native to eastern and southern India and Sri Lanka. It is also grown in Cambodia, Myanmar, China, Thailand and the Andaman Islands. It is a flowering plant with the largest inf ...
'', ''
Corypha utan
''Corypha utan'', the cabbage palm, buri palm or gebang palm, is a species of palm native to Asia and Oceania.
Description
It grows up to tall, and, on the York Peninsula of Queensland, up to 1.5 meters (4' 11") thick (exceeded only by Boras ...
'', ''
Elaeis guineensis
''Elaeis guineensis'' is a species of palm commonly just called oil palm but also sometimes African oil palm or macaw-fat. It is the principal source of palm oil. It is native to west and southwest Africa, specifically the area between Angola an ...
'', ''
Howea forsteriana
''Howea forsteriana'', the Kentia palm, thatch palm or palm court palm, is a species of flowering plant in the palm family, Arecaceae, endemic to Lord Howe Island in Australia. It is also widely grown on Norfolk Island. It is a relatively slow-g ...
'', ''
Jubaea chilensis
''Jubaea'' is a genus of palms with one species, ''Jubaea chilensis'' or ''Jubaea spectabilis'', commonly known in English as the Chilean wine palm or Chile cocopalm, and palma chilena in Spanish. It is native to southwestern South America and i ...
'', ''
Livistona chinensis
''Livistona chinensis'', the Chinese fan palm or fountain palm, is a species of subtropical palm tree of east Asia. It is native to southern Japan, Taiwan, the Ryukyu Islands, southeastern China and Hainan. In Japan, two notable populations occ ...
'', ''
L. decora'', ''
Metroxylon sagu
''Metroxylon sagu'', the true sago palm, is a species of palm in the genus '' Metroxylon'', native to tropical southeastern Asia.
The tree is a major source of sago starch.
Description
True sago palm is a suckering (multiple-stemmed) palm, ...
'', ''
Phoenix canariensis
''Phoenix canariensis'', the Canary Island date palm or pineapple palm, is a species of flowering plant in the palm family Arecaceae, native to the Canary Islands off the coast of Morocco. It is a relative of ''Phoenix dactylifera'', the true ...
'', ''
P. dactylifera'' (date palm), ''
P. sylvestris'', ''
Roystonea regia
''Roystonea regia'', commonly known as the Cuban royal palm or Florida royal palm, is a species of palm that is native to Mexico, parts of Central America and the Caribbean, and southern Florida. A large and attractive palm, it has been planted ...
'', ''
Sabal palmetto
''Sabal palmetto'' (, '' SAY-bəl''), also known as cabbage palm, cabbage palmetto, sabal palm, blue palmetto, Carolina palmetto, common palmetto, Garfield's tree, and swamp cabbage, is one of 15 species of palmetto palm.
It is native to the So ...
'', ''
Trachycarpus fortunei
''Trachycarpus fortunei'', the Chinese windmill palm, windmill palm or Chusan palm, is a species of hardy evergreen palm tree in the family Arecaceae, native to parts of China, Japan, Myanmar and India.
Description
Growing to tall, ''Trachyca ...
'', ''
Washingtonia filifera
''Washingtonia filifera'', the desert fan palm, California fan palm, or California palm,Flora of North America Association. ''Flora of North America: North of Mexico Volume 22: Magnoliophyta: Alismatidae, Arecidae, Commelinidae (in Part), and Zi ...
'', and ''
W. robusta''.
Lab studies have reared the insect on diets of ''
Agave americana
''Agave americana'', common names century plant, maguey, or American aloe, is a species of flowering plant in the family Asparagaceae, native to Mexico and the United States in Texas. It is cultivated worldwide as an ornamental plant, and has b ...
'' and ''
Saccharum officinarum
''Saccharum officinarum'' is a large, strong-growing species of grass in the genus '' Saccharum''. Its stout stalks are rich in sucrose, a simple sugar which accumulates in the stalk internodes. It originated in New Guinea, and is now cultivated ...
'', but these findings have not been observed in the wild. There is evidence that the weevil prefers the 'Sukkary' cultivar of date palm to other cultivars.
The palm species ''
Washingtonia filifera
''Washingtonia filifera'', the desert fan palm, California fan palm, or California palm,Flora of North America Association. ''Flora of North America: North of Mexico Volume 22: Magnoliophyta: Alismatidae, Arecidae, Commelinidae (in Part), and Zi ...
'' and ''
Chamaerops humilis
''Chamaerops'' is a genus of flowering plants in the family Arecaceae. The only currently fully accepted species is ''Chamaerops humilis'', variously called European fan palm or the Mediterranean dwarf palm. It is one of the most hardy palms, ...
'' may be moderately resistant to the red palm weevil, though both are known hosts.
Life cycle
![Rhynchophorus_ferrugineus_larva](https://upload.wikimedia.org/wikipedia/commons/8/8d/Rhynchophorus_ferrugineus_larva.JPG)
This weevil usually infests palms younger than twenty years.
While the adult causes some damage through feeding, it is the burrowing of the larva into the heart of the palm that can cause the greatest mortality of trees. The adult female lays approximately two hundred eggs on new growth in the crown of the palm, at the base of young leaves, or in open lesions on the plant. The egg hatches into a white, legless
larva
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle.
The ...
. The larva will feed on the soft fibres and terminal buds, tunneling through the internal tissue of the tree for about a month. The larvae can occasionally grow to a length of .
At
pupation
A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in thei ...
, the larva will leave the tree and form a cocoon built of dry palm fibers in leaf litter at the base of the tree. The total life cycle takes about 3-4 months.
Oviposition
After fertilization, the adult female can lay between 300 and 500 eggs. They lay in holes they produced while searching for food, or take advantage of the cracks or wounds in a recently cut palm. At oviposition, females bend upward and the tarsi are anchored to the tissue with the spines of the third pair of legs to push the ovipositor into the tough palm tissue. After laying, the female protects and secures the eggs with a secretion that rapidly hardens around the eggs. On average, females produce 210 eggs per clutch, most of which hatch over a period of 3 days. The eggs are white, cylindrical, glossy, oval shaped, and measure . The back of these eggs possess special 'gill cover' structures that provide the developing insect with oxygen.
Larvae
The neonate larvae are yellow-white, segmented, legless, and have a chitinous head capsule that is a darker brown than the rest of the body. They have powerful horizontal conical jaws which they use to burrow from the axils of the leaves to the crown, where they feed voraciously. Upon completion of larval development, the larva will sometimes emerge from the trunk of the tree, and build a pupal case of fiber extracted from the galleries inside the palm. The larva will then undergo metamorphosis into an adult. The larva will also weave a pupal case at the base of the palm fronds within the frond itself or at the centre of the base of the plant.
Adult
The adult insect is an excellent flier and is able to travel great distances.
While they prefer to attack palms that are already infested or weakened by other stresses, they will colonize healthy palms.
Predators, diseases, and parasites
''R. ferrugineus'' is predated by ''
Chelisoches morio'', infected by a
cytoplasmic polyhedrosis virus and ''
Metarhizium pingshaense'', and parasitized by ''
Heterorhabditis indicus'', ''
Hypoaspis'' spp., ''
Praecocilenchus ferruginophorus'', ''
Scolia erratica'', ''
Steinernema carpocapsae
''Steinernema carpocapsae'' is an entomopathogenic nematode and a member of the family Steinernematidae. It is a parasitic roundworm that has evolved an insect-killing symbiosis with bacteria, and kills its hosts within a few days of infection ...
'', and ''
Steinernema riobravis''.
Behaviour
![Rh](https://upload.wikimedia.org/wikipedia/commons/8/88/Rh._ferrugineus_treatment_02.jpg)
The behaviour and bionomics of the red palm weevil have been investigated in the field. The biology and life history of the weevil have been investigated on five diets, including banana, sugarcane, squash fruit, apple, and palm crown. The rate of development showed great variation depending on the host used. Developmental time was shortest on palm crown, followed by banana, and then squash fruit and apple, and longest on sugarcane. Egg production was the highest on palm crown, being 338 ± 37.24 eggs/female followed by banana, squash fruit, and apple, and the lowest (117. ± 18.9 eggs/female) on sugarcane.
Studies show that this insect is attracted by ethyl acetate, 2-methoxy.4.vinylphenol, gamma-nonanoic lactone, 4SSS-ferrugineol, 50H and 4me-9-5Kt.
Symptoms of infestation
The infestation of the pest can result in yellowing and wilting of palms, that may lead to the death of the affected plant.
The crown wilts first, and lower leaves will follow, due to damage to vascular tissue. Major symptoms such as crown loss or leaf wilt are usually only visible long after the palm has become infested. Secondary infections of opportunistic bacteria and fungi may occur within damaged tissues, accelerating decline. By the time these external symptoms are observed, the damage is usually sufficient to kill the tree, and the infestation may have been present for six months or longer. In high-density infestations, sounds of the larvae burrowing and chewing can be heard by placing one's ear to the trunk of the palm. Recent research has been conducted using electronic listening devices or dogs trained to recognize the scent of weevils or palm decay to detect infestations at low densities earlier in the process.
Image:Phoenix_canariensis_first_sign_of_infection_with_Rhynchophorus_ferrugineus.JPG, Canary Island date palm, ''Phoenix canariensis
''Phoenix canariensis'', the Canary Island date palm or pineapple palm, is a species of flowering plant in the palm family Arecaceae, native to the Canary Islands off the coast of Morocco. It is a relative of ''Phoenix dactylifera'', the true ...
'', with first obvious infestation signs
Image:Phoenix canariensis destroyed by Rhynchophorus ferrugineus 1.JPG, Destroyed crown of a Canary Island date palm
Image:Phoenix canariensis destroyed by Rhynchophorus ferrugineus 4.JPG, Bases of palm leaves fallen from the crown of the tree, with burrows and extracted pupal cases.
Image:WeevilStrickenDatePalm-in-Kfar Saba-RJP.jpg, A date palm that was killed by the pest, in Kfar Saba
Kfar Saba ( he, כְּפַר סָבָא), officially Kefar Sava, is a city in the Sharon region, of the Central District of Israel. In 2019 it had a population of 110,456, making it the 16th-largest city in Israel. The population of Kfar Saba i ...
, Israel
Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
.
Control
The main control method is through the application of a
systemic insecticide. Insecticide is usually applied through a funnel about above the infested area of the trunk. The red palm weevil can be monitored using
pheromone lures and alternative forms of control use field sanitation and mass
trapping
Animal trapping, or simply trapping or gin, is the use of a device to remotely catch an animal. Animals may be trapped for a variety of purposes, including food, the fur trade, hunting, pest control, and wildlife management.
History
Neolithic ...
with traps
baited with
pheromone
A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...
and plant derived
semiochemical
A semiochemical, from the Greek wiktionary:σημεῖον, σημεῖον (''semeion''), meaning "signal", is a chemical substance or mixture released by an organism that affects the behaviors of other individuals. Semiochemical communication c ...
s. New alternative technologies using semiochemicals and
bioinsecticide
Biological control or biocontrol is a method of controlling pests, such as insects, mites, weeds, and plant diseases, using other organisms. It relies on predation, parasitism, herbivory, or other natural mechanisms, but typically also invo ...
s are being developed to attract the weevils to a point source and kill them. Another management technique is to drench the base of palm
frond
A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the lar ...
s with the entomopathogenic fungus ''
Metarhizium robertsii
''Metarhizium robertsii'' formerly known as ''M. anisopliae'', and even earlier as ''Entomophthora anisopliae'' ( basionym) is a fungus that grows naturally in soils throughout the world and causes disease in various insects by acting as a par ...
'' (
syn. ''M. anisopliae'', ''Entomophthora anisopliae''), or ''
Beauveria bassiana
''Beauveria bassiana'' is a fungus that grows naturally in soils throughout the world and acts as a parasite on various arthropod species, causing white muscardine disease; it thus belongs to the entomopathogenic fungi. It is used as a biological ...
''.
An Italian company claims to have developed a microwave collar that can be used to sterilize individual trees. For early detection, bioacoustic analysis may be implemented by inserting a sensitive microphone into the tree and recording any produced sounds.
[
] These sounds are analyzed by digital signal processing and artificial intelligence to decide whether they are generated by palm weevils.
Palms' natural
defense
Defense or defence may refer to:
Tactical, martial, and political acts or groups
* Defense (military), forces primarily intended for warfare
* Civil defense, the organizing of civilians to deal with emergencies or enemy attacks
* Defense industr ...
against this weevil is understudied.
RNA interference
RNA interference (RNAi) is a biological process in which RNA molecules are involved in sequence-specific suppression of gene expression by double-stranded RNA, through translational or transcriptional repression. Historically, RNAi was known by o ...
(RNAi, a kind of gene silencing) is a discovered defense system in many host-pathogen systems.
RNAi shows promise as a
breeding target when breeding palm for RPW resistance.
[
]
Prevention
As the weevil prefers to lay its eggs in softer tissues, avoiding mechanical damage to plants can help to reduce infestation. Tarring wounds after pruning a plant of dead or old leaves can also reduce the probability of infestation. The movement of plant material such as husks, dead leaves, or untreated
coir
Coir (), also called coconut fibre, is a natural fibre extracted from the outer husk of coconut and used in products such as floor mats, doormats, brushes, and mattresses. Coir is the fibrous material found between the hard, internal shell ...
from infested to uninfested areas is not recommended.
Culinary uses
The larval grub is considered a delicacy in
Vietnam
Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
.
In Vietnam, the larvae are usually eaten alive with fish sauce. Other methods of cooking include toasting and steaming. They are eaten with sticky rice and salad or cooked with porridge. The larvae are known in the Vietnamese language as ''đuông dừa'' ("coconut beetle-larva").
"Sago worms" reported from other countries (e.g., East Malaysia, New Guinea) refer to different, related species of ''Rhynchophorus''.
References
Additional resources
European Commission - Directorate-General for Health & Consumers (2011) The insect killing our palm trees. EU efforts to stop the Red Palm Weevil.
Red Palm Weevil HomeRed Palm Weevil Control & TipsRed Palm Weevil Research Chair, King Saud University, RiyadhGreek webpage dedicated to management*
USDA
The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of com ...
br>
information page
{{DEFAULTSORT:Rhynchophorus Ferrugineus
Dryophthorinae
Beetles of Asia
Fauna of Pakistan
Beetles described in 1790
Pests of oil palm