Redondasuchus
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''Redondasuchus'' is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
aetosaur Aetosaurs () are heavily armored reptiles belonging to the extinct order Aetosauria (; from Greek, (aetos, "eagle") and (, "lizard")). They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs ...
. It may be a
junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...
of '' Typothorax coccinarum'', another aetosaur. ''Redondasuchus'' is a member of the
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
Typothoracisinae within the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Aetosaurinae Aetosaurinae is one of the two main clades of aetosaurs, the other being Desmatosuchia. It is a stem-based taxon defined as all aetosaurs more closely related to ''Aetosaurus'' than ''Desmatosuchus''. Aetosaurinae currently comprises ''Aetosaurus ...
, and lived during the middle
Norian The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic defi ...
stage of the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
. Material belonging to the genus has been found from the
Redonda Formation The Redonda Formation is a geologic formation exposed in eastern New Mexico.Dobrovolny and Summerson 1947 It contains vertebrate fossils of the late Triassic Period.Griggs and Read 1959 Fossil theropod tracks have been reported from the formation ...
in east-central
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, ''R. reseri'', was named in 1991 after having been referred to as a species of ''Typothorax'' since 1985. A second species, ''R. rineharti'', was described in 2006.


Description and species

''Redondasuchus'' was first named with the description of the type species, ''R. reseri'', in 1991. ''R. reseri'' was named on the basis of isolated scutes found at Apache Canyon and Shark Tooth Hill in
Quay County, New Mexico Quay County () is a county in the state of New Mexico. As of the 2010 census, the population was 9,041. Its county seat is Tucumcari. The county was named for Pennsylvania senator Matthew Quay, who supported statehood for New Mexico. Its easter ...
. ''R. rineharti'' was described in 2006 from several scutes and part of a right
femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with ...
found from Apache Canyon. While other aetosaurs have
scute A scute or scutum (Latin: ''scutum''; plural: ''scuta'' "shield") is a bony external plate or scale overlaid with horn, as on the shell of a turtle, the skin of crocodilians, and the feet of birds. The term is also used to describe the anterior po ...
s covered in pits and grooves that often form radial patterns, the scutes of ''Redondasuchus'' are densely covered in pits and lack any patterning. The dorsal paramedian scutes (those that line the back) are unique in that they are angled approximately 45°. In other aetosaurs, the scutes arc smoothly around the back from the midline to the lateral scutes that lie below them and run along the side of the animal. The dorsal paramedians of ''R. reseri'' are smaller than those of ''R. rineharti''. In both species, each paramedian possesses a keel on the underside that extends from the medial edge (the edge near the vertebrae) to the flexed area of the scute.


Plagiarism allegations


Background

The orientation of the dorsal paramedian osteoderms of ''Redondasuchus'' has been disputed. Originally, Hunt & Lucas (1991) argued that the angled part of the osteoderm was two thirds the way down from the medial edge. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
osteoderm of ''R. reseri'' was identified as a left paramedian based on the position of the anterior bar, an unornamented strip of bone that is known to lie on the front edge of the osteoderm in other aetosaurs. This interpretation was repeated in subsequent papers by Heckert ''et al''. (1996) and Heckert & Lucas (2000). In 2002, an unpublished
Texas Tech University Texas Tech University (Texas Tech, Tech, or TTU) is a public research university in Lubbock, Texas. Established on , and called Texas Technological College until 1969, it is the main institution of the five-institution Texas Tech University Sys ...
thesis A thesis ( : theses), or dissertation (abbreviated diss.), is a document submitted in support of candidature for an academic degree or professional qualification presenting the author's research and findings.International Standard ISO 7144: ...
by Jeffrey Martz claimed that the holotype of ''R. reseri'' was actually a right paramedian. This was justified by comparison to the structure and ossification pattern of ''Typothorax coccinarum'', a closely related species of aetosaur. Martz also suggested that ''R. reseri'' was
synonymous A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
with ''Typothorax coccinarum.'' This thesis, though widely disseminated among aetosaur workers, was not published in a peer-reviewed journal. An equivalent reinterpretation of ''Redondasuchus''' osteoderm was first described in a peer-reviewed journal by Spielmann ''et al.'' (2006), a review of the taxon published in the
New Mexico Museum of Natural History and Science The New Mexico Museum of Natural History and Science is a natural history and science museum in Albuquerque, New Mexico near Old Town Albuquerque. The Museum was founded in 1986. It operates as a public revenue facility of the New Mexico Departmen ...
Bulletin. Spielmann, Hunt, and Lucas, who all contributed to the article, were also members of the editorial board for the Bulletin, while Hunt was additionally the museum director at the time. They claimed to provide a novel interpretation of the holotype osteoderm as a right paramedian, noting that the orientation of the osteoderm in the diagrams of previous published papers may have introduced confusion. One diagram, from Heckert ''et al''. (1996), depicted the anterior margin as placed near the bottom of the page with the posterior margin near the top, which may have contributed to its misidentification. Martz's 2002 thesis was widely cited, but only in criticism of his argument that ''Redondasuchus'' and ''Typothorax'' were synonymous. His arguments on osteoderm orientation were not mentioned.


Allegations

In 2007, Martz, along with fellow paleontologists
Darren Naish Darren William Naish is a British vertebrate palaeontologist, author and science communicator. As a researcher, he is best known for his work describing and reevaluating dinosaurs and other Mesozoic reptiles, including ''Eotyrannus'', ''Xenopos ...
, Mike Taylor, and Matt Wedel, publicly accused Spielmann ''et al.'' of committing plagiarism. They asserted that the authors of Spielmann ''et al''. (2006) could not have come up with the same interpretation as Martz independently. They pointed to how the authors were in possession of the 2002 thesis for three years, their extensive citation of the thesis, and similarities between certain figures in Spielmann ''et al''. (2006) and the thesis. This controversy was connected to a similar allegation of ethical misconduct and editorial mismanagement related to an aetosaur paper published by NMMNH staff in December 2006. The paper in question provided the genus name '' Rioarribasuchus'' for a species of aetosaur which Bill Parker was in the process of publishing under the name ''Heliocanthus'', although Parker's paper was not fully published until January 2007. These coinciding controversies, termed " Aetogate", were publicized both in the scientific community and in local Albuquerque news. After requests for a
New Mexico Department of Cultural Affairs The New Mexico Department of Cultural Affairs is a state agency of the New Mexico government. Created as the Office of Cultural Affairs (OCA) in 1980, the New Mexico Department of Cultural Affairs was elevated to a state Cabinet-level agency in 200 ...
inquiry, Martz ''et al''.'s allegations, as well as those of Parker, were reviewed and rejected in a 2008 NMMNH meeting which was subsequently reported by Lucas. Lucas's report phrased the omittance of Martz's identification as an "oversight" by Spielmann ''et al.'' (2006), and also argued that pre-2002 papers on the topic presented "inconsistencies" rather than a specific argument on orientation. The report's arguments and conclusions were heavily criticized by the accusing parties. A separate independent investigation by the
Society of Vertebrate Paleontology The Society of Vertebrate Paleontology (SVP) is a professional organization that was founded in the United States in 1940 to advance the science of vertebrate paleontology around the world. Mission and Activities SVP has about 2,300 members inter ...
did not find evidence for explicit, intentional plagiarism in Spielmann ''et al.'' (2006). The SVP report did not "absolve either party from responsibility", and also criticize the editorial and regulatory practices of the NMMNH and its staff, the lack of communication or collaboration experienced during the situation, and the difficulty in proving plagiarism or conflicts of interest in paleontological work. A revised code of ethics was published by the SVP alongside the results of their investigation.


Phylogeny

Heckert ''et al.'' (1996) considered ''Redondasuchus'' to be part of an advanced grade of aetosaurs that also included ''
Neoaetosauroides ''Neoaetosauroides'' is an extinct genus of primitive aetosaur. Its type and only species is ''N. engaeus''. Fossils have been found in Los Colorados Formation outcropping along the Sierra Morada River in the Ischigualasto-Villa Unión Basin in ...
'', ''
Longosuchus ''Longosuchus'' (meaning "Long's crocodile") is an extinct genus of aetosaur from the Late Triassic of North America and Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of Nort ...
'', ''
Desmatosuchus ''Desmatosuchus'' (, from Greek δεσμός ''desmos'' 'link' + σοῦχος ''soûkhos'' 'crocodile') is an extinct genus of archosaur belonging to the Order Aetosauria. It lived during the Late Triassic. Description ''Desmatosuchus'' was a ...
'', '' Paratypothorax'', and ''
Typothorax ''Typothorax'' is an extinct genus of typothoracine aetosaur that lived in the Late Triassic. Its remains have been found in North America. Two species are known: ''T. coccinarum'', the type species, and ''T. antiquum''. Description ''Typothor ...
''.


References

{{Taxonbar, from=Q2908646 Aetosaurs of North America Prehistoric pseudosuchian genera