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The Purus várzea (NT0156) is an
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of ...
of seasonally flooded
várzea forest A várzea forest is a seasonal floodplain forest inundated by whitewater rivers that occurs in the Amazon biome. Until the late 1970s, the definition was less clear and várzea was often used for all periodically flooded Amazonian forests. Althoug ...
in the central Amazon basin. It is part of the Amazon biome. The ecoregion is home to a vegetation adapted to floods of up to that may last for eight months. There is a great variety of fish and birds, but relatively fewer mammals. Ground-dwelling mammals must migrate to higher ground during the flood season. Threats include logging, cattle farming, over-fishing and mercury pollution from gold mining.


Location

The Purus várzea is a low-lying region of the central Amazon basin that is seasonally flooded. It covers of eastern Colombia and western Brazil. It extends along most of the Juruá, central Purus, and Caquetá ( Japurá) rivers and their tributaries. In the east it reaches the confluence of the Japurá and Solimões Rivers. Urban centers in or around the region are
Tefé Tefé, known in early accounts as Teffé, is a municipality in the state of Amazonas, northern Brazil. Location Tefé is located about 525 km by air or 595 km by river to the west of Manaus on the south bank of the Rio Solimões (th ...
,
Tabatinga Tabatinga, originally Forte de São Francisco Xavier de Tabatinga, is a municipality in the Três Fronteiras area of Western Amazonas. It is in the Brazilian state of Amazonas. Its population was 67,182 (2020) and its area is 3,225 km2. T ...
and Carauarí. To the southeast the varzea adjoins the Purus–Madeira moist forests, and to the northeast it adjoins the Japurá–Solimões–Negro moist forests. Streams that flow through the Southwest Amazon moist forests, Solimões–Japurá moist forests and Caquetá moist forests to the west, and the Juruá–Purus moist forests in the central region all contain stretches of the varzea. The Monte Alegre várzea is downstream along the Solimões.


Physical

Altitudes range from . The forests are seasonally flooded by whitewater rivers, which carry suspended sediment washed from the eastern slopes of the
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
and organic material. Water levels rise by up to in the flood period, which may last for eight months of the year. The soil is fertile, composed of sediments that have accumulated in the present
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...
epoch and that are renewed by the annual floods. The river course through the floodplain constantly shifts over time, creating oxbow lakes, levees, meander swales and bars. These landscape elements support diverse vegetation adapted to flooding, which gradually merges into the surrounding terra firme forest.


Ecology

The ecoregion is in the
Neotropical realm The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeo ...
and the
tropical and subtropical moist broadleaf forests Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature. Description TSMF is generally found in large, discon ...
biome.


Climate

The
Köppen climate classification The Köppen climate classification is one of the most widely used climate classification systems. It was first published by German-Russian climatologist Wladimir Köppen (1846–1940) in 1884, with several later modifications by Köppen, notabl ...
is "Af": equatorial, fully humid. Average temperatures range from , with a mean temperature of . Average annual precipitation is about . Rain falls throughout the year, but is heaviest in January–March and lightest in July–August.


Flora

The floodplain holds aquatic vegetation where drainage is poor, successional vegetation, forest mosaics and permanent swamp vegetation. The várzea forests holds more species than várzea on the lower Amazon, but fewer species than in the surrounding terra firme forests. The forest is continuous. The rich understory includes plants of the families
Zingiberaceae Zingiberaceae () or the ginger family is a family of flowering plants made up of about 50 genera with a total of about 1600 known species of aromatic perennial herbs with creeping horizontal or tuberous rhizomes distributed throughout tropical Af ...
,
Marantaceae The Marantaceae are a family, the arrowroot family, of flowering plants consisting of 31 genera and around 530 species, defining it as one of the most species-rich families in its order.Kennedy, H. (2000). “Diversification in pollination mechan ...
and
Heliconiaceae ''Heliconia'', derived from the Greek word (), is a genus of flowering plants in the monotypic family Heliconiaceae. Most of the ca 194 known species are native to the tropical Americas, but a few are indigenous to certain islands of the ...
. Economically valuable timber trees include ''
Carapa guianensis ''Carapa guianensis'' is a species of tree in the family Meliaceae, also known by the common names andiroba or crabwood. Description Andiroba is native to the Amazon and is widely used by the indigenous populations of the northern region o ...
'', '' Iryanthera surinamensis'', ''
Ceiba pentandra ''Ceiba pentandra'' is a tropical tree of the order Malvales and the family Malvaceae (previously emplaced in the family Bombacaceae), native to Mexico, Central America and the Caribbean, northern South America, and (as the variety ''C. pentandr ...
'' and '' Calycophyllum spruceanum''. On the upper levees the most common trees are ''
Ceiba pentandra ''Ceiba pentandra'' is a tropical tree of the order Malvales and the family Malvaceae (previously emplaced in the family Bombacaceae), native to Mexico, Central America and the Caribbean, northern South America, and (as the variety ''C. pentandr ...
'', ''
Hura crepitans ''Hura crepitans'', the sandbox tree, also known as possumwood and jabillo, is an evergreen tree of the spurge family (Euphorbiaceae), native to tropical regions of North and South America including the Amazon rainforest. It is also present in p ...
'' and '' Parinari excelsa''. On the lower levees common trees include ''
Pterocarpus santalinoides ''Pterocarpus santalinoides'' is a tree species in the legume family (biology) (Fabaceae); it is locally known as ''mututi''. It has a remarkable bi-continental distribution, native to tropical western Africa (Benin, Burkina Faso, Cameroon, Gam ...
'', '' Eschweilera albiflora'', '' Piranhea trifoliata'' and '' Neoxythece elegans''. The lowest areas contain abundant bamboo (
Bambusa ''Bambusa'' is a large genus of clumping bamboos. Most species of ''Bambusa'' are rather large, with numerous branches emerging from the nodes, and one or two much larger than the rest. The branches can be as long as 11 m (35 ft). The ...
species), and pioneer trees that include
Cecropia ''Cecropia'' is a Neotropical genus consisting of 61 recognized species with a highly distinctive lineage of dioecious trees. The genus consists of pioneer trees in the more or less humid parts of the Neotropics, with the majority of the speci ...
species, '' Pseudobombax munguba'', many
Ficus ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species extendi ...
species and the '' Astrocaryum jauari'' palm. There are relatively few palms compared to the terra firme forests. Other palm species include ''
Astrocaryum murumuru ''Astrocaryum murumuru'' (Portuguese common name: murumuru) is a palm native to Amazon Rainforest vegetation in Brazil, which bears edible fruits. ''Murumuru'' butter, extracted from the seeds of the plant, may be used as a moisturizer.One re ...
'', ''
Mauritia flexuosa ''Mauritia flexuosa'', known as the moriche palm, ''ité'' palm, ''ita'', ''buriti'', ''muriti'', ''miriti'' (Brazil), ''canangucho'' (Colombia), ''acho'' (Ecuador), or ''aguaje'' (Peru), is a palm tree. It grows in and near swamps and other wet ...
'' and
Bactris ''Bactris'' is a genus of spiny palms which are native to Mexico, South and Central America and the Caribbean. Most species are small trees about tall, but some are large trees while others are shrubs with subterranean stems. They have simple o ...
species including the endemic '' Bactris tefensis''. Trees that support fruit-eating fish that enter the forest in the flood period include yellow mombim ( Spondias mombim), jauari palm ( Astrocaryum jauari), biribá (
Rollinia deliciosa ''Annona mucosa'' is a species of flowering plant in the custard-apple family, Annonaceae, that is native to tropical South America. It is cultivated for its edible fruits, commonly known as biribá, lemon meringue pie fruit, or wild sugar-apple ...
), tarumã (
Vitex cymosa ''Vitex cymosa'' is a species of tree in the family Lamiaceae. It is native to Panama and South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small por ...
), and apui (
Ficus ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species extendi ...
species). The largest tree in the várzea is ''
Ceiba pentandra ''Ceiba pentandra'' is a tropical tree of the order Malvales and the family Malvaceae (previously emplaced in the family Bombacaceae), native to Mexico, Central America and the Caribbean, northern South America, and (as the variety ''C. pentandr ...
''. Other typical várzea trees are '' Parkia inundabilis'', '' Septotheca tessmannii'', '' Coumarouna micrantha'', '' Ceiba burchellii'', '' Ochroma lagopus'' and '' Manilkara inundata''. Flooded forests hold ''
Virola surinamensis ''Virola surinamensis'', known commonly as baboonwood, ucuuba, ucuhuba and ''chalviande'',Americas Regional Workshop (Conservation & Sustainable Management of Trees, Costa Rica, November 1996). 1998''Virola surinamensis''.The IUCN Red List of Thr ...
'' and ''
Euterpe oleracea Euterpe (; el, Εὐτέρπη, lit=rejoicing well' or 'delight , from grc, εὖ, eû, well + el, τέρπειν, térpein, to please) was one of the Muses in Greek mythology, presiding over music. In late Classical times, she was named muse ...
''.


Fauna

199 species of mammals are found in the region, fewer than in the surrounding terra firme forests. There are no ground-dwelling species in flooded várzea forests isolated from the mainland, but areas of várzea in contact with terra firme forests house ground-dwelling mammals that migrate to higher ground in the flood periods. There are two endemic primates in the
Mamirauá Sustainable Development Reserve The Mamirauá Sustainable Development Reserve ( pt, Reserva de Desenvolvimento Sustentável Mamirauá) in the Brazilian state of Amazonas, near the city of Tefé, is a reserve near the village of Boca do Mamirauá. It includes mostly Amazonian ...
, the
bald uakari The bald uakari (''Cacajao calvus'') or bald-headed uakari is a small New World monkey characterized by a very short tail; bright, crimson face; a bald head; and long coat. The bald uakari is restricted to várzea forests and other wooded habitat ...
(''Cacajao calvus'') and
black squirrel monkey The black squirrel monkey (''Saimiri vanzolinii''), also known as the blackish squirrel monkey or black-headed squirrel monkey, is a small New World monkey, New World primate, endemic (ecology), endemic to the central Amazon Rainforest, Amazon in ...
(''Saimiri vanzolinii''). The reserve is also home to
white-footed saki The white-footed saki, buffy saki or white saki (''Pithecia albicans'') is a species of saki monkey, a type of New World monkey, endemic to western Brazil south of the Amazon Amazon most often refers to: * Amazons, a tribe of female warriors ...
(''Pithecia albicans''),
emperor tamarin The emperor tamarin (''Saguinus imperator'') is a species of tamarin allegedly named for its resemblance to the German emperor Wilhelm II. It lives in the west Brazilian states of Acre and Amazonas and the southwest Amazon Basin, in east Peru, ...
(''Saguinus imperator''),
moustached tamarin The moustached tamarin (''Saguinus mystax'') is a New World monkey and a species of tamarin. The moustached tamarin is named for the lack of coloring in the facial hair surrounding their mouth, appearing similar to a moustache. As with all New Wo ...
(''Saguinus mystax''),
Nancy Ma's night monkey Nancy Ma's night monkey (''Aotus nancymaae'') is a night monkey species from South America. It is found in Brazil and Peru. It is known in medical research as a model organism A model organism (often shortened to model) is a non-human spec ...
(''Aotus nancymaae'') and Hershkovitz's titi (''Callicebus dubius''). Other mammals include
white-lipped peccary The white-lipped peccary (''Tayassu pecari'') is a species of peccary found in Central and South America and the only member of the genus ''Tayassu''. Multiple subspecies have been identified. White-lipped peccaries are similar in appearance to ...
(''Tayassu pecari''),
common agouti The agouti (, ) or common agouti is any of several rodent species of the genus ''Dasyprocta''. They are native to Middle America, northern and central South America, and the southern Lesser Antilles. Some species have also been introduced else ...
(Dasyprocta genus),
lowland paca The lowland paca (''Cuniculus paca''), also known as the spotted paca, is a large rodent found in tropical and sub-tropical America, from east-central Mexico to northern Argentina, and has been introduced to Cuba and Algeria. The animal is cal ...
(''Cuniculus paca''), jaguar (''Panthera onca''), capybara (''Hydrochoerus hydrochaeris''), spiny tree-rat (genus ''Echimys'') and the Ega long-tongued bat (''Scleronycteris ega''). Aquatic mammals include the
Amazon river dolphin The Amazon river dolphin (''Inia geoffrensis''), also known as the boto, bufeo or pink river dolphin, is a species of toothed whale classified in the family Iniidae. Three subspecies are currently recognized: ''I. g. geoffrensis'' (Amazon river ...
(''Inia geoffrensis''),
tucuxi The tucuxi (''Sotalia fluviatilis''), alternatively known in Peru ''bufeo gris'' or ''bufeo negro'', is a species of freshwater dolphin found in the rivers of the Amazon basin. The word ''tucuxi'' is derived from the Tupi language word ''tuchu ...
(''Sotalia fluviatilis'') and Amazonian manatee' (''Trichechus inunguis''). Endangered mammals include the
white-bellied spider monkey The white-bellied spider monkey (''Ateles belzebuth''), also known as the white-fronted or long-haired spider monkey, is an endangered species of spider monkey, a type of New World monkey. It is found in the north-western Amazon in Colombia, Ecu ...
(''Ateles belzebuth''), Peruvian spider monkey (''Ateles chamek''), white-cheeked spider monkey (''Ateles marginatus''),
Marinkelle's sword-nosed bat Marinkelle's sword-nosed bat (''Lonchorhina marinkellei'') is a bat species found in Colombia. In 2013, Bat Conservation International listed this species as one of the 35 species of its worldwide priority list of conservation. Its species name ...
(''Lonchorhina marinkellei'') and
giant otter The giant otter or giant river otter (''Pteronura brasiliensis'') is a South American carnivorous mammal. It is the longest member of the weasel family, Mustelidae, a globally successful group of predators, reaching up to . Atypical of muste ...
(''Pteronura brasiliensis''). 633 bird species have been reported. These include many aquatic species such as heron and egret (genera '' Egretta'' and '' Ardea''), whistling duck (''Dendrocygna'' species), ibis ('' Cercibis'' and ''
Theristicus ''Theristicus'' is a genus of birds in the family Threskiornithidae. They are found in open, grassy habitats in South America. All have a long, decurved dark bill, relatively short reddish legs that do not extend beyond the tail in flight (unlik ...
'' species) and
roseate spoonbill The roseate spoonbill (''Platalea ajaja'') is a gregarious wading bird of the ibis and spoonbill family, Threskiornithidae. It is a resident breeder in both South and North America. Taxonomy The roseate spoonbill is sometimes placed in its own ...
s (''Platalea ajaja''). Endangered birds include
wattled curassow The wattled curassow (''Crax globulosa'') is a threatened member of the family Cracidae, the curassows, guans, and chachalacas. It is found in remote rainforests in the western Amazon basin in South America. Males have black plumage, except ...
(''Crax globulosa'') and green-thighed parrot (''Pionites leucogaster''). The very large
green anaconda The green anaconda (''Eunectes murinus''), also known as the giant Emerald anaconda, common anaconda, common water boa or sucuri, is a boa species found in South America. It is the heaviest and one of the longest known extant snake species. Lik ...
(''Eunectes murinus'') is found in the várzea. Other reptiles include black caiman (''Melanosuchus niger'') and spectacled caiman (''Caiman crocodilus''). The ecoregion is home to the endangered Johnson's horned treefrog ('' Hemiphractus johnsoni''). Large fruit-eating fish that enter the forest during the flood period include pacu (genera ''
Metynnis ''Metynnis'' is a genus of serrasalmid fish from tropical and subtropical South America. They are herbivorous or omnivorous, and inhabit a wide range of freshwater habitats, ranging from rivers and streams (both fast- and slow-flowing), to flood ...
'' and ''
Mylossoma ''Mylossoma'' is a genus of serrasalmids from tropical and subtropical South America, including the basins of the Amazon, Orinoco, Lake Maracaibo and Paraguay- Paraná. These common fish are found both in main river sections and floodplains. T ...
''), tambaqui (''Colossoma macropomum''), pirarucu (''
Arapaima gigas ''Arapaima gigas'', also known as pirarucu, is a species of arapaima native to the basin of the Amazon River. Once believed to be the sole species in the genus, it is among the largest freshwater fish. The species is an obligate air-breather, s ...
'') and sardinha ('' Triportheus angulatus''). Other fish include piranha ('' Serrasalmus'' species) and decorative fish from whitewater rivers and their blackwater tributaries and lakes such as blue discus (''
Symphysodon aequifasciatus ''Symphysodon aequifasciatus'', the blue discus or brown discus, is a species of cichlid native to rivers of the eastern and central Amazon Basin downriver from the Púrus Arch. This discus is found in black-, clear- and whitewater, but its p ...
''), cichlids (genus ''
Cichlasoma ''Cichlasoma'' is a genus of freshwater fish in the cichlid family. The genus was previously very large (a wastebasket taxon), including cichlids from North America, including Central America, and South America. Reclassification and subsequent ...
''), characins (family
Anostomidae The Anostomidae are a family (biology), family of ray-finned fishes that belong to the order (biology), order Characiformes. Closely related to the Chilodontidae (fishes), Chilodontidae and formerly included with them, the Anostomidae contain abo ...
), tetras (genera ''
Hemigrammus ''Hemigrammus'' is a genus of freshwater fish in the family Characidae native to South America (including Trinidad) and commonly seen in the aquarium trade. These are medium-small tetras where the largest species reach up to around . Speci ...
'' and ''
Hyphessobrycon ''Hyphessobrycon'' is a genus of freshwater fish in the family Characidae. These species are among the fishes known as tetras. The genus is distributed in the Neotropical realm from southern Mexico to Río de la Plata in Argentina. Many of these ...
''), and catfish (families
Aspredinidae The Aspredinidae are a small South American family of catfishes (order Siluriformes) also known as the banjo catfishes, with about 43 species. Distribution Aspredinids are found throughout the major tropical rivers of South America (e.g., Magda ...
,
Callichthyidae Callichthyidae is a family of catfishes (order Siluriformes), called armored catfishes due to the two rows of bony plates (or scutes) along the lengths of their bodies. It contains some of the most popular freshwater aquarium fish, such as many ...
,
Doradidae The Doradidae are a family of catfishes also known as thorny catfishes, raphael catfishes or talking catfishes. These fish are native to South America, primarily the Amazon basin and the Guianas. Doradids are omnivorous. Taxonomy As of 2007, 3 ...
and
Loricariidae The Loricariidae is the largest family of catfish (order Siluriformes), with 92 genera and just over 680 species. Loricariids originate from freshwater habitats of Costa Rica, Panama, and tropical and subtropical South America. These fish are not ...
).


Status

According to the
World Wildlife Fund The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wo ...
the status of the ecoregion is Critical/Endangered. Smallholders along the rivers undertake mixed agriculture, forest exploitation, small-scale logging and livestock raising. In some areas the várzea forest is threatened by large-scale logging and cattle farming. Fish populations are threatened by large-scale fishing operations and may be threatened by excessive collection of aquarium fish such as the red discus ('' Symphysodon discus''). Gold mining contaminates the Purus and Japurá with mercury. Conservation units in the region include the
Mamirauá Sustainable Development Reserve The Mamirauá Sustainable Development Reserve ( pt, Reserva de Desenvolvimento Sustentável Mamirauá) in the Brazilian state of Amazonas, near the city of Tefé, is a reserve near the village of Boca do Mamirauá. It includes mostly Amazonian ...
,
Amanã Sustainable Development Reserve Amanã Sustainable Development Reserve ( pt, Reserva de Desenvolvimento Sustentável Amanã) is a sustainable development reserve in the state of Amazonas, Brazil. Location The Amanã Sustainable Development Reserve (RDS) covers in the north-c ...
and
Jaú National Park The Jaú National Park ( pt, Parque Nacional do Jaú) is a national park located in the state of Amazonas, Brazil. It is one of the largest forest reserve in South America, and part of a World Heritage Site. Location The name "Jaú" comes from ...
.


Notes


Sources

* * * {{DEFAULTSORT:Purus varzea Ecoregions of Brazil Amazon biome Neotropical tropical and subtropical moist broadleaf forests