''Pneumatoraptor'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

of small,

paravian Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosaurids, troodontids, anchiornithids, and possibly the scansoriopterygids, the group also contains the aviala ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

that lived in

Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the ...

. It is known from a single complete left shoulder girdle (

scapulocoracoid The scapulocoracoid is the unit of the pectoral girdle that contains the coracoid and scapula. The coracoid itself is a beak-shaped bone that is commonly found in most vertebrates with a few exceptions. The scapula is commonly known as the ''shoulde ...

) found in the

Csehbánya Formation The Csehbánya Formation is a geological formation in the Transdanubian Mountains of Veszprém County, Hungary. The formation dates to the Late Santonian (around 85-84 million years ago) of the Late Cretaceous. It represents a floodplain environ ...

of the Iharkút locality in the Bakony Mountains of western Hungary. This formation dates to the late

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...

period (

Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. ...

age) about 85 million years ago.Ősi, A., Apesteguía, S.M and Kowalewski, M. (2010). "Non-avian theropod dinosaurs from the early Late Cretaceous of Central Europe." ''Cretaceous Research'' 31: 304-320, The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...

is ''Pneumatoraptor fodori'', named for Géza Fodor, who provided funding for the dig. The genus name ''Pneumatoraptor'', which means "air thief", Atkinson, L. "PNEUMATORAPTOR :: from DinoChecker's dinosaur archive". ‹http://www.dinochecker.com/dinosaurs/PNEUMATORAPTOR›. Web access: 13th Nov 2020. refers to the pneumaticity of the bone, the hollow spaces that would have been filled with air sacs in life. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...

specimen is identified by the catalog number MTM V.2008.38.1. and housed at the

Hungarian Natural History Museum The Hungarian Natural History Museum ( hu, Magyar Természettudományi Múzeum) in Budapest, dating back to 1802, houses the largest natural history collections of Hungary and the region. History of the museum Foundation In 1802, Count Fere ...

Budapest Budapest (, ; ) is the capital and most populous city of Hungary. It is the ninth-largest city in the European Union by population within city limits and the second-largest city on the Danube river; the city has an estimated population ...

Description

''Pneumatoraptor'' differs from other

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

s in having a narrow shoulder blade that is nearly circular in cross section, as well as having a large opening in the bone to house a hollow air sac cavity. The bone is small, indicating that the animal was about three times smaller than ''

Velociraptor ''Velociraptor'' (; ) is a genus of small dromaeosaurid dinosaur that lived in Asia during the Late Cretaceous epoch, about 75 million to 71 million years ago. Two species are currently recognized, although others have been assigned in the p ...

''. The shoulder girdle is L-shaped, showing it to be a member of the group

Paraves Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosaurids, troodontids, anchiornithids, and possibly the scansoriopterygids, the group also contains the avial ...

, which also includes the

dromaeosaurids Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...

troodontids Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discov ...

, and birds. While the ''Pneumatoraptor'' remains are too incomplete to tell which, if any, of these specific groups it belongs to, many of the bones features are similar to dromaeosaurids. Other paravian remains from the same formation may belong to ''Pneumatoraptor''. These include isolated teeth, claws, tail vertebrae, and a partial lower leg bone (tibia).

References

Prehistoric paravians Santonian life Late Cretaceous dinosaurs of Europe Fossils of Hungary Fossil taxa described in 2010 {{Theropod-stub