Pleistoannelida is a group of annelid worms that comprises the vast majority of the diversity in

phylum In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclatu ...

Annelida. Discovered through phylogenetic analyses, it is the largest clade of

annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...

s, comprised by the last common ancestor of the highly diverse

sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

s Errantia and

Sedentaria Sedentaria is a diverse clade of annelid worms. It is traditionally treated as a subclass of the paraphyletic class Polychaeta, but it is also a monophyletic group uniting several polychaetes and the monophyletic class Clitellata. It is the sist ...

(

Clitellata The Clitellata are a class of annelid worms, characterized by having a clitellum - the 'collar' that forms a reproductive cocoon during part of their life cycles. The clitellates comprise around 8,000 species. Unlike the class of Polychaeta, t ...

and related polychaetes) and all the descendants of that ancestor.

Phylogeny

Pleistoannelida is composed by two big clades: Errantia ( eunicids, siboglinids and related

polychaete Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are ...

s) and

( spoon worms,

tubeworms A tubeworm is any worm-like sessile invertebrate that anchors its tail to an underwater surface and secretes around its body a mineral tube, into which it can withdraw its entire body. Tubeworms are found among the following taxa: * Annelida, the ...

clitellate The Clitellata are a class of annelid worms, characterized by having a clitellum - the 'collar' that forms a reproductive cocoon during part of their life cycles. The clitellates comprise around 8,000 species. Unlike the class of Polychaeta, th ...

s and related polychaetes). There are also smaller groups of difficult placement, such as

Myzostomida The Myzostomida or Myzostomatida are an order of small marine worms, which are parasitic on echinoderms, mostly crinoids. These highly unusual and diverse annelids were first discovered by Friedrich Sigismund Leuckart in 1827. Morphology A ...

and Spintheridae, that belong to either one of them. Closely related to Pleistoannelida is a grade of basal annelids: the

Amphinomida Amphinomida is an order of marine polychaetes. The order contains two families: * '' Amphinomidae'' Lamarck, 1818 * '' Euphrosinidae'' Williams, 1852 References Polychaetes {{annelid-stub ...

Sipunculida The Sipuncula or Sipunculida (common names sipunculid worms or peanut worms) is a class containing about 162 species of unsegmented marine annelid worms. The name ''Sipuncula'' is from the genus name '' Sipunculus'', and comes from the Latin ...

/'' Lobatocerebrum'' clade, Chaetopterida and Palaeoannelida. A possible, closer sister group to Pleistoannelida could be Dinophiliformia, a clade containing interstitial (

meiofauna Meiobenthos, also called meiofauna, are small benthic invertebrates that live in both marine and fresh water environments. The term ''meiofauna'' loosely defines a group of organisms by their size, larger than microfauna but smaller than macrof ...

l) genera previously found in Orbiniida (

). A 2019 analysis recovered the

mesozoa The Mesozoa are minuscule, worm-like parasites of marine invertebrates. Generally, these tiny, elusive creatures consist of a somatoderm (outer layer) of ciliated cells surrounding one or more reproductive cells. A recent study recovered Mesoz ...

n group

Orthonectida Orthonectida () is a small phylum of poorly known parasites of marine invertebrates that are among the simplest of multi-cellular organisms. Members of this phylum are known as orthonectids. Biology The adults, which are the sexual stage, are mi ...

as the sister group to Pleistoannelida. However, a more recent 2022 analysis found a

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...

Dicyemida Dicyemida, also known as Rhombozoa, is a phylum of tiny parasites that live in the renal appendages of cephalopods. Taxonomy Classification is controversial. Traditionally, dicyemids have been grouped with the Orthonectida in the Mesozoa, ...

clade localized outside of the annelids, between Gnathifera and

Platyhelminthes The flatworms, flat worms, Platyhelminthes, or platyhelminths (from the Greek πλατύ, ''platy'', meaning "flat" and ἕλμινς (root: ἑλμινθ-), ''helminth-'', meaning "worm") are a phylum of relatively simple bilaterian, unsegment ...

References

{{Taxonbar, from=Q115868988 Annelid unranked clades Polychaetes