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Mesozoa
The Mesozoa are minuscule, worm-like parasites of marine invertebrates. Generally, these tiny, elusive creatures consist of a somatoderm (outer layer) of ciliated cells surrounding one or more reproductive cells. A recent study recovered Mesozoa as a monophyletic group that emerged in the Lophotrochozoa as sister of the Rouphozoa. Some workers previously classified Mesozoa as the sole phylum of the lonely subkingdom Agnotozoa. Cavalier-Smith argued that at least some of the mesozoans are in fact protistans, not animals. In the 19th century, the Mesozoa were a wastebasket taxon for multicellular organisms which lacked the invaginating gastrula which was thought to define the Metazoa. Evolution Mesozoa were once thought to be evolutionary intermediate forms between Protozoans and Metazoans, but now they are thought to be degenerate or simplified metazoa. Their ciliated larvae are similar to the miracidium of trematodes, and their internal multiplication is similar to what h ...
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Dicyema
''Dicyema'' is a genus of worms belonging to the family Dicyemidae. Species: * '' Dicyema acciaccatum'' McConnaughey, 1949 * '' Dicyema acheroni'' McConnaughey, 1949 * '' Dicyema acuticephalum'' Nouvel, 1947 - parasitizes ''Octopus vulgaris'' * '' Dicyema aegira'' McConnaughey & Kritzler, 1952 * '' Dicyema akashiense'' Furuya, 2005 * '' Dicyema apalachiensis'' Short, 1962 * '' Dicyema apollyoni'' Nouvel, 1947 * '' Dicyema australis'' Penchaszadeh, 1969 * '' Dicyema awajiense'' Furuya, 2005 * '' Dicyema balanocephalum'' Furuya, 2006 * '' Dicyema banyulensis'' Furuya & Hochberg, 1999 * '' Dicyema benedeni'' Furuya & Hochberg, 1999 * ''Dicyema benthoctopi'' Hochberg & Short, 1970 * '' Dicyema bilobum'' Couch & Short, 1964 * ''Dicyema briarei'' Short, 1961 * ''Dicyema caudatum'' Bogolepova-Dobrokhotova, 1960 * ''Dicyema clavatum'' Furuya & Koshida, 1992 - parasitizes ''Octopus minor'' * ''Dicyema colurum'' Furuya, 1999 - parasitizes ''Octopus fangsiao'' * ''Dicyema dolichocephalum' ...
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Mesozoan
The Mesozoa are minuscule, worm-like parasites of marine invertebrates. Generally, these tiny, elusive creatures consist of a somatoderm (outer layer) of ciliated cells surrounding one or more reproductive cells. A recent study recovered Mesozoa as a monophyletic group that emerged in the Lophotrochozoa as sister of the Rouphozoa. Some workers previously classified Mesozoa as the sole phylum of the lonely subkingdom Agnotozoa. Cavalier-Smith argued that at least some of the mesozoans are in fact protistans, not animals. In the 19th century, the Mesozoa were a wastebasket taxon for multicellular organisms which lacked the invaginating gastrula which was thought to define the Metazoa. Evolution Mesozoa were once thought to be evolutionary intermediate forms between Protozoans and Metazoans, but now they are thought to be degenerate or simplified metazoa. Their ciliated larvae are similar to the miracidium of trematodes, and their internal multiplication is similar to what h ...
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Orthonectida
Orthonectida () is a small phylum of poorly known parasites of marine invertebrates that are among the simplest of multi-cellular organisms. Members of this phylum are known as orthonectids. Biology The adults, which are the sexual stage, are microscopic wormlike animals, consisting of a single layer of ciliated outer cells surrounding a mass of sex cells. They swim freely within the bodies of their hosts, which include flatworms, polychaete worms, bivalve molluscs, and echinoderms. Most are gonochoristic, with separate male and female individuals, but a few species are hermaphroditic. When they are ready to reproduce, adults leave the host, and sperm from the males penetrate the bodies of the females to achieve internal fertilisation. The resulting zygote develops into a ciliated larva that escapes from the mother to seek out new hosts. Once it finds a host, the larva loses its cilia and develops into a syncytial plasmodium larva. This, in turn, breaks up into numerous individua ...
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Dicyemida
Dicyemida, also known as Rhombozoa, is a phylum of tiny parasites that live in the renal appendages of cephalopods. Taxonomy Classification is controversial. Traditionally, dicyemids have been grouped with the Orthonectida in the Mesozoa, and, as of 2017, molecular evidence appears to confirm this. However, other molecular phylogenies have placed the dicyemids more closely related to the roundworms. Additional molecular evidence suggests that this phylum is derived from the ''Lophotrochozoa. The phylum is not divided in classes or orders, but contains three families, Conocyemidae, Dicyemidae, and Kantharellidae. Anatomy Adult dicyemids range in length from , and they can be easily viewed through a light microscope. They display eutely, a condition in which each adult individual of a given species has the same number of cells, making cell number a useful identifying character. Dicyemida lack respiratory, circulatory, excretory, digestive, and nervous systems. The organism' ...
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Subkingdom (biology)
In biology, a kingdom is the second highest taxonomic rank, just below domain. Kingdoms are divided into smaller groups called phyla. Traditionally, some textbooks from the United States and Canada used a system of six kingdoms (Animalia, Plantae, Fungi, Protista, Archaea/Archaebacteria, and Bacteria/Eubacteria) while textbooks in Great Britain, India, Greece, Brazil and other countries use five kingdoms only (Animalia, Plantae, Fungi, Protista and Monera). Some recent classifications based on modern cladistics have explicitly abandoned the term ''kingdom'', noting that some traditional kingdoms are not monophyletic, meaning that they do not consist of all the descendants of a common ancestor. The terms ''flora'' (for plants), ''fauna'' (for animals), and, in the 21st century, '' funga'' (for fungi) are also used for life present in a particular region or time. Definition and associated terms When Carl Linnaeus introduced the rank-based system of nomenclature into biolo ...
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Agnotozoa
Agnotozoa is the name of a taxon of simple animals. The name first appeared in an invertebrate paleontology book as one of the "branches" of the subkingdom Metazoa. The branch contained only one group: Mesozoa. More recently, some have used the name to refer to a sub-kingdom of three small phyla of simple animals without organs. These phyla are Placozoa, Orthonectida, and Rhombozoa. They still have differentiated tissue, but that tissue is only organized in simple ways; for example, by being layered. The Orthonectida and Rhombozoa are also grouped into the Mesozoa. Biologists today generally do not use the taxon Agnotozoa. There are a few likely reasons for this. * There is doubt that placozoans are closely related to mesozoans. * There is doubt even that the two mesozoan groups (orthonectids and rhombozoans) are related to each other. * Even if the two mesozoan groups are related, there is little need for another name in addition to mesozoa. Wikispecies places the mesozoans i ...
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Annelid
The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecologies – some in marine environments as distinct as tidal zones and hydrothermal vents, others in fresh water, and yet others in moist terrestrial environments. The Annelids are bilaterally symmetrical, triploblastic, coelomate, invertebrate organisms. They also have parapodia for locomotion. Most textbooks still use the traditional division into polychaetes (almost all marine), oligochaetes (which include earthworms) and leech-like species. Cladistic research since 1997 has radically changed this scheme, viewing leeches as a sub-group of oligochaetes and oligochaetes as a sub-group of polychaetes. In addition, the Pogonophora, Echiura and Sipuncula, previously regarded as separate phyla, are now regarded as sub-groups of polycha ...
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Lophotrochozoa
Lophotrochozoa (, "crest/wheel animals") is a clade of protostome animals within the Spiralia. The taxon was established as a monophyletic group based on molecular evidence. The clade includes animals like annelids, molluscs, bryozoans, brachiopods, and platyhelminthes. Groups Lophotrochozoa was defined in 1995 as the "last common ancestor of the three traditional lophophorate taxa (brachiopods, bryozoans, and phoronid worms), the mollusks and the annelids, and all of the descendants of that common ancestor". It is a cladistic definition (a node-based name), so the affiliation to Lophotrochozoa of spiralian groups not mentioned directly in the definition depends on the topology of the spiralian tree of life, and in some phylogenetic hypotheses, Lophotrochozoa may even be synonymous to Spiralia. Nemertea and Orthonectida (if not directly considered as part of Annelida) are probably lophotrochozoan phyla; Dicyemida, Gastrotricha, and Platyhelminthes may be lophotrochozoans ...
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Phylum
In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about 8 phyla. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguished them as a group ("a self-contained unity" ...
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Protista
A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the exclusion of other eukaryotes means that protists do not form a natural group, or clade. Therefore, some protists may be more closely related to animals, plants, or fungi than they are to other protists. However, like the groups ''algae'', ''invertebrates'', and ''protozoans'', the biological category ''protist'' is used for convenience. Others classify any unicellular eukaryotic microorganism as a protist. The study of protists is termed protistology. History The classification of a third kingdom separate from animals and plants was first proposed by John Hogg in 1860 as the kingdom Protoctista; in 1866 Ernst Haeckel also proposed a third kingdom Protista as "the kingdom of primitive forms". Originally these also included prokaryotes, but ...
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Thomas Cavalier-Smith
Thomas (Tom) Cavalier-Smith, FRS, FRSC, NERC Professorial Fellow (21 October 1942 – 19 March 2021), was a professor of evolutionary biology in the Department of Zoology, at the University of Oxford. His research has led to discovery of a number of unicellular organisms (protists) and advocated for a variety of major taxonomic groups, such as the Chromista, Chromalveolata, Opisthokonta, Rhizaria, and Excavata. He was known for his systems of classification of all organisms. Life and career Cavalier-Smith was born on 21 October 1942 in London. His parents were Mary Maude (née Bratt) and Alan Hailes Spencer Cavalier Smith. He was educated at Norwich School, Gonville and Caius College, Cambridge (MA) and King's College London (PhD). He was under the supervision of Sir John Randall for his PhD thesis between 1964 and 1967; his thesis was entitled "''Organelle Development in'' Chlamydomonas reinhardii". From 1967 to 1969, Cavalier-Smith was a guest investigato ...
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Monoblastozoa
''Salinella salve'' is a dubious species of a very simple animal that may not exist, but which some have named as the sole member of the phylum Monoblastozoa. It was discovered in 1892 by Johannes Frenzel in the salt pans of Argentina and cultivated in a laboratory by him. This animal has not been found since and its real existence is considered as doubtful. Michael Schrödl from the Zoological State Collection in Munich is involved in a project to search for ''Salinella'' in Argentina. ''Monoblastozoa'' was granted the title of phylum after the recognition that Mesozoa was too diverse to be a phylum into itself. Description According to Frenzel's description, ''S. salve'' is more organized than Protozoa, but still very primitive for a multicellular organism. They are characterised by their distinct anterior/posterior parts and being densely ciliated, especially around the "mouth" and "anus". They have only one layer of cells, and reproduce asexually by transverse fiss ...
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