Orbiniida
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Orbiniida
Orbiniida is an order of small polychaete worms in the phylum Annelida. It is the earliest diverging clade in Sedentaria. Along with Protodriliformia (in Errantia), this order is composed of meiofaunal marine life, marine worms formerly known as "Archiannelida, archiannelids". These worms inhabit the Marine life, marine interstitial ecosystem, the space between sand grains. Classification First mentioned in 1977, Orbiniida was an order of the polyphyletic infraclass "Scolecida" of the class "Polychaeta". It was composed of the families Orbiniidae, Paraonidae and Questidae, all of which share in common the lack of antennae and palps, and the presence of an eversible pharynx and biramous parapodia with simple chaetae. However, through phylogenetic analysis it was found that Questidae and Paraonidae are more closely related to Clitellata and Cirratulida respectively. Because of this, the taxonomic composition of Orbiniida was changed to exclude these two families and include several i ...
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Orbiniidae
Orbiniidae is a family of polychaete worms. Orbiniids are mostly unselective deposit feeders on marine detritus. They can be found from the neritic zone to abyssal depths. The family was revised in 1957 by Hartman and some further revisions were made by Blake in 2000.Blake, J.A. 2000. A new genus and species of polychaete worm (Family Orbiniidae) from methane seeps in the Gulf of Mexico, with a review of the systematics and phylogenetic interrelationships of the genera of Orbiniidae. Cah. Biol. Mar. 41: 435-449. Genera :*'' Alcandra'' :*'' Aricia'' Savigny, 1820 :*'' Berkeleyia'' :*'' Branchethus'' :*''Califia'' :*''Clytie'' :*'' Falklandiella'' :*''Labotas'' :*''Lacydes'' :*'' Leitoscoloplos'' Day, 1977 :*'' Leodamas'' :*'' Microrbinia'' Hartman, 1965 :*'' Naidonereis'' :*'' Nainereis'' Blainville, 1828 :*'' Naineris'' Blainville, 1828 :*'' Orbinella'' :*'' Orbinia'' Quatrefages, 1865 :*'' Orbiniella'' Day, 1954 :*'' Paraorbiniella'' Rullier, 1974 :*'' Pararicia'' :*'' Petti ...
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Questidae
Orbiniidae is a family of polychaete worms. Orbiniids are mostly unselective deposit feeders on marine detritus. They can be found from the neritic zone to abyssal depths. The family was revised in 1957 by Hartman and some further revisions were made by Blake in 2000.Blake, J.A. 2000. A new genus and species of polychaete worm (Family Orbiniidae) from methane seeps in the Gulf of Mexico, with a review of the systematics and phylogenetic interrelationships of the genera of Orbiniidae. Cah. Biol. Mar. 41: 435-449. Genera :*'' Alcandra'' :*'' Aricia'' Savigny, 1820 :*'' Berkeleyia'' :*'' Branchethus'' :*''Califia'' :*''Clytie'' :*'' Falklandiella'' :*''Labotas'' :*''Lacydes'' :*'' Leitoscoloplos'' Day, 1977 :*'' Leodamas'' :*'' Microrbinia'' Hartman, 1965 :*'' Naidonereis'' :*'' Nainereis'' Blainville, 1828 :*'' Naineris'' Blainville, 1828 :*'' Orbinella'' :*'' Orbinia'' Quatrefages, 1865 :*'' Orbiniella'' Day, 1954 :*'' Paraorbiniella'' Rullier, 1974 :*'' Pararicia'' :*'' Petti ...
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Sedentaria
Sedentaria is a diverse clade of annelid worms. It is traditionally treated as a subclass of the paraphyletic class Polychaeta, but it is also a monophyletic group uniting several polychaetes and the monophyletic class Clitellata. It is the sister group of Errantia. Phylogeny The phylogeny of polychaetes is slowly being resolved. Sedentaria and Errantia are the two biggest clades of polychaetes, and together they compose clade Pleistoannelida. Sedentaria's most basal clade is Orbiniida. Other groups that are nested within Sedentaria are: Clitellata, the Sabellida/Spionida clade, Opheliida, Echiura, Cirratuliformia, Terebelliformia, Maldanomorpha and the families Siboglinidae and Capitellidae. Some taxa, such as Spintheridae and Myzostomida, are still difficult to place due to their long branching, but they likely belong to either Errantia Errantia is a diverse group of marine polychaete worms in the phylum Annelida. Traditionally a subclass of the paraphyletic class "Polycha ...
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Scolecida
Scolecida is an Infraclass (biology), infraclass of polychaete worms. Scolecids are mostly unselective Detritivore, deposit feeders on marine detritus. Characteristics Scolecids have Parapodium, parapodia with rami that are all alike. The prostomium is distinct. The head has no appendages or palps and is usually conical, though in the Scalibregmatidae, it has a "T"-shaped tip, and in Paraonidae, there is a single, central antenna. In some families there are sometimes some tiny eyespots. The oesophagus is evertable forming a sac-like proboscis which may have several finger-like lobes. The anterior segments and their appendages are all similar. The notopodia and neuropodia consist of unbranched capillary chaetae, sometimes with hooks. There is a single central gill in Cossuridae on an anterior segment and simple segmental gills are present in some other families. Systematics The families Arenicolidae, Capitellidae and Maldanidae were formerly part of the order Capitellida. They a ...
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Polychaete
Polychaeta () is a paraphyletic class (biology), class of generally marine invertebrate, marine annelid worms, common name, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are made of chitin. More than 10,000 species are described in this class. Common representatives include the lugworm (''Arenicola marina'') and the Alitta virens, sandworm or Alitta succinea, clam worm ''Alitta''. Polychaetes as a class are robust and widespread, with species that live in the coldest ocean temperatures of the abyssal plain, to forms which tolerate the extremely high temperatures near hydrothermal vents. Polychaetes occur throughout the Earth's oceans at all depths, from forms that live as plankton near the surface, to a 2- to 3-cm specimen (still unclassified) observed by the robot ocean probe Nereus (underwater vehicle), ''Nereus'' at the bottom of the Challenger Deep, the deepes ...
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Protodriliformia
Protodriliformia is a clade of small marine life, marine polychaetes, comprised by the groups of meiofaunal interstitial worms Protodrilida and Polygordiidae, formerly considered "archiannelida, archiannelids". It is the most basal clade of Errantia. Evolutionary history Phylogenetic analysis of annelids has found Protodriliformia to be the earliest diverging clade of Errantia. At the same time, the other half of "archiannelid" worms, Orbiniida, was found to be the earliest diverging clade of Sedentaria. The Convergent evolution, convergence between these two groups occurred through progenesis and miniaturization, as a way to adapt to the marine interstitial ecosystem between sand grains (i.e. interstitium). This means that the larval or juvenile stages of a larger pleistoannelida, pleistoannelid ancestor temporarily inhabiting the interstitium stopped their somatic cell, somatic growth, became Sexual maturity, sexually mature and stayed in the interstitium permanently. Reference ...
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Polychaeta
Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are made of chitin. More than 10,000 species are described in this class. Common representatives include the lugworm (''Arenicola marina'') and the sandworm or clam worm ''Alitta''. Polychaetes as a class are robust and widespread, with species that live in the coldest ocean temperatures of the abyssal plain, to forms which tolerate the extremely high temperatures near hydrothermal vents. Polychaetes occur throughout the Earth's oceans at all depths, from forms that live as plankton near the surface, to a 2- to 3-cm specimen (still unclassified) observed by the robot ocean probe ''Nereus'' at the bottom of the Challenger Deep, the deepest known spot in the Earth's oceans. Only 168 species (less than 2% of all polychaetes) are known from f ...
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Parergodrilidae
Parergodrilidae is an enigmatic family of polychaetes with only two genera, one living on the coast, the other terrestrial. They share much in common with the clitellates, but molecular data place them with Questa and Orbiniidae Orbiniidae is a family of polychaete worms. Orbiniids are mostly unselective deposit feeders on marine detritus. They can be found from the neritic zone to abyssal depths. The family was revised in 1957 by Hartman and some further revisions w .... References Polychaetes {{annelid-stub ...
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Clitellata
The Clitellata are a class of annelid worms, characterized by having a clitellum - the 'collar' that forms a reproductive cocoon during part of their life cycles. The clitellates comprise around 8,000 species. Unlike the class of Polychaeta, they do not have parapodia and their heads are less developed. Characteristics Clitellate annelids are segmented worms characterised by the clitellum or girdle which is located near the head end of mature individuals. The mouth is on the ventral surface and is overhung by the prostomium (proboscis). The brain is not located in the head but in one of the body segments. The clitellum is formed by a modification of several segments, and either includes the female gonopores or is located just behind them. During copulation, this glandular tissue secretes mucus that keeps the paired individuals together while they exchange sperm. Afterwards it secretes material that forms a cocoon that encircles the animal's body and encloses the eggs and sperm. T ...
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Chaeta
A chaeta or cheta (from Greek χαίτη “crest, mane, flowing hair"; plural: chaetae) is a chitinous bristle or seta found in annelid worms, (although the term is also frequently used to describe similar structures in other invertebrates such as arthropods). Polychaete annelids, ('polychaeta' literally meaning "many bristles") are named for their chaetae. In Polychaeta, chaetae are found as bundles on the parapodia, paired appendages on the side of the body. The chaetae are epidermal extracellular structures, and clearly visible in most polychaetes. They are probably the best studied structures in these animals. Use in taxonomy and identification The ultrastructure of chaetae is fundamentally similar for all taxa but there is vast diversity in chaetal morphology. Moreover, chaetae bear precise characters for determination of species and taxonomic assessment. The shape, absolute and relative size, number, position, ornamentation and type are important taxonomic characters a ...
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Cirratulida
Cirratulida is an order of polychaetes belonging to the class Polychaeta. Families: * Cirratulidae Cirratulidae is a family of marine polychaete worms. Members of the family are found worldwide, mostly living in mud or rock crevices. Most are deposit feeders, but some graze on algae or are suspension feeders. Description Cirratulids vary in ... Carus, 1863 * Paraonidae Cerruti, 1909 References Polychaetes {{annelid-stub ...
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Phylogenetic Analysis
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does n ...
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