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A piranha or piraña (, , or ; or , ) is one of a number of freshwater fish in the family
Serrasalmidae The Serrasalmidae (serrasalmids) are a family of characiform fishes, recently elevated to family status. It includes more than 90 species. The name means "serrated salmon family", which refers to the serrated keel running along the belly of these ...
, or the subfamily Serrasalminae within the tetra family,
Characidae Characidae, the characids or characins is a family of freshwater subtropical and tropical fish, belonging to the order Characiformes. The name "characins" is the historical one, but scientists today tend to prefer "characids" to reflect their st ...
in
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
Characiformes. These fish inhabit South American rivers, floodplains, lakes and reservoirs. Although often described as extremely predatory and mainly feeding on fish, their dietary habits vary extensively, and they will also take plant material, leading to their classification as omnivorous.


Etymology

The name originates from the indigenous
Tupi people A subdivision of the Tupi-Guarani linguistic families, the Tupi people were one of the largest groups of indigenous Brazilians before its colonization. Scholars believe that while they first settled in the Amazon rainforest, from about 2,900 ...
and their respective Tupi language. It is formed from two words, meaning fish and meaning tooth; the same word is used by Indians to describe a pair of scissors. Another possible derivation is from , probably literally "biting-fish". In the mid 18th century the Portuguese merged the word into . Finally, the word may also come from the combination of meaning fish and meaning cut (which also meant "bad" or "devil" in Tupi-Guarani).


Taxonomy and evolution

Piranhas belong to the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Serrasalminae The Serrasalmidae (serrasalmids) are a family of characiform fishes, recently elevated to family status. It includes more than 90 species. The name means "serrated salmon family", which refers to the serrated keel running along the belly of these ...
, which includes closely related omnivorous fish such as
pacu Pacu () is a common name used to refer to several species of omnivorous South American freshwater serrasalmid fish that are related to the piranha. Pacu and piranha do not have similar teeth, the main difference being jaw alignment; piranha hav ...
s. Traditionally, only the four
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
''
Pristobrycon ''Pristobrycon'' is a genus of piranhas from the Orinoco and Amazon Basins, as well as rivers in the Guianas. ''Pristobrycon'' is not monophyletic. No single morphological feature has been found that completely diagnoses this genus. ''P. striola ...
'', ''
Pygocentrus ''Pygocentrus'' is a genus of the piranha family Serrasalmidae. All species are native to tropical and subtropical South America. All the species are predatory, scavengers and may form large schools. The famous red-bellied piranha, ''Pygocentrus ...
'', ''
Pygopristis ''Pygopristis denticulata'', or the Lobetoothed Piranha is a species of piranha. These fish are part of the group Ostariophysi, a large group of freshwater fish that includes minnows and catfishes. It is a rare South American fish found in the O ...
'', and ''
Serrasalmus ''Serrasalmus'' is a genus of piranhas. They are collectively known as pirambebas; the "typical" piranhas like the piraya piranha are nowadays placed in ''Pygocentrus''. Like all piranhas, ''Serrasalmus'' are native to South America. These fish ...
'' are considered to be true piranhas, due to their specialized teeth. However, a recent analysis showed, if the piranha group is to be
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
, it should be restricted to ''Serrasalmus'', ''Pygocentrus'', and part of ''Pristobrycon'', or expanded to include these taxa plus ''Pygopristis'', ''
Catoprion ''Catoprion'' is a genus of serrasalmids from tropical South America, including the basins of the Amazon, Essequibo, Orinoco and Paraguay rivers. It was believed to be a monotypic In biology, a monotypic taxon is a taxonomic group (taxon) t ...
'', and ''
Pristobrycon striolatus ''Pristobrycon striolatus'' is a species of serrasalmid fish. Range and habitat ''Pristobrycon striolatus'' inhabits mainly black or acidic waters in tributaries in the Orinoco and Amazon River Basins. Description This small and beautiful f ...
''. ''Pygopristis'' was found to be more closely related to ''Catoprion'' than the other three piranha genera. The total number of piranha species is unknown and contested, and new species continue to be described. Estimates range from fewer than 30 to more than 60.


Distribution

Piranhas are indigenous to the
Amazon basin The Amazon basin is the part of South America drained by the Amazon River and its tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries of Bolivi ...
, in the
Orinoco The Orinoco () is one of the longest rivers in South America at . Its drainage basin, sometimes known as the Orinoquia, covers , with 76.3 percent of it in Venezuela and the remainder in Colombia. It is the fourth largest river in the wor ...
, in rivers of the
Guianas The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * ...
, in the ParaguayParaná, and the
São Francisco River The São Francisco River (, ) is a large river in Brazil. With a length of , it is the longest river that runs entirely in Brazilian territory, and the fourth longest in South America and overall in Brazil (after the Amazon, the Paraná and t ...
systems, but there are major differences in the species richness. In a review where 38–39 piranha species were recognized, 25 were from the Amazon and 16 from Orinoco, while only three were present in Paraguay–Paraná and two in São Francisco. Most species are restricted to a single river system, but some (such as the red-bellied piranha) occur in several. Many species can occur together; for example, seven are found in Caño Maporal, a stream in Venezuela.
Aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
piranhas have been unsuccessfully introduced into parts of the United States. In many cases, however, reported captures of piranhas are misidentifications of pacu (e.g., red-bellied pacu or ''
Piaractus brachypomus ''Piaractus brachypomus'', the pirapitinga, is a large species of pacu, a close relative of piranhas and silver dollars, in the serrasalmid family.Nico, L.; P. Fuller; and M. Neilson (22 October 2013)Piaractus brachypomus.USGS Nonindigenous Aqua ...
'' is frequently misidentified as red-bellied piranha or ''
Pygocentrus nattereri The red-bellied piranha, also known as the red piranha (''Pygocentrus nattereri''), is a type of piranha native to South America, found in the Amazon, Paraguay, Paraná and Essequibo basins, as well as coastal rivers of northeastern Brazil.
''). Piranhas have also been discovered in the Kaptai Lake in southeast Bangladesh. Research is being carried out to establish how piranhas have moved to such distant corners of the world from their original habitat. Some rogue exotic fish traders are thought to have released them in the lake to avoid being caught by antipoaching forces. Piranhas were also spotted in the
Lijiang River The Li River or Li Jiang () is the name for the upper reaches of the Gui River in northwestern Guangxi, China. It is part of the Xijiang River system in the Pearl River Basin. The river flows from Xing'an County to Pingle County, where the kars ...
in
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
.


Description


Size

Depending on the exact species, most piranhas grow to between long. A few can grow larger, with the largest living species, the red-bellied, reaching up to . There are claims of
São Francisco piranha ''Pygocentrus piraya'', often called the piraya piranha or San Francisco piranha, and sometimes sold as the man-eating piranha, is a large, aggressive piranha from the São Francisco River basin in Brazil. Description It is one of the largest ...
s at up to , but the largest confirmed specimens are considerably smaller. The extinct ''
Megapiranha ''Megapiranha'' is an extinct serrasalmid characin fish from the Late Miocene (8–10 million years ago) Ituzaingó Formation of Argentina, described in 2009. The type species is ''M. paranensis''.dentition Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ...
. All piranhas have a single row of sharp teeth in both jaws. The teeth are tightly packed and interlocking (via small cusps) and are used for rapid puncture and shearing. Individual teeth are typically broadly triangular, pointed, and blade-like (flat in profile). The variation in the number of cusps is minor. In most species, the teeth are tricuspid with a larger middle cusp which makes the individual teeth appear markedly triangular. The exception is ''Pygopristis'', which has pentacuspid teeth and a middle cusp usually only slightly larger than the other cusps.


Biting abilities

Piranhas have one of the strongest bites found in bony fishes. Relative to body mass, the black piranha (''Serrasalmus rhombeus'') produces one of the most forceful bites measured in vertebrates. This extremely powerful and dangerous bite is generated by large jaw muscles (adductor mandibulae) that are attached closely to the tip of the jaw, conferring the piranha with a mechanical advantage that favors force production over bite speed. Strong jaws combined with finely serrated teeth make them adept at tearing flesh.


Ecology

Piranhas vary extensively in ecology and behavior depending on exact species. Piranhas, especially the red-bellied (''Pygocentrus nattereri''), have a reputation as ferocious predators that hunt their prey in schools. Recent research, however, which "started off with the premise that they school as a means of cooperative hunting", discovered they are timid fish that schooled for protection from their own predators, such as
cormorant Phalacrocoracidae is a family of approximately 40 species of aquatic birds commonly known as cormorants and shags. Several different classifications of the family have been proposed, but in 2021 the IOC adopted a consensus taxonomy of seven ge ...
s, caimans, and dolphins. Piranhas are "basically like regular fish with large teeth". A few other species may also occur in large groups, while the remaining are solitary or found in small groups. Although popularly described as highly predatory and primarily feeding on fish, piranha diets vary extensively, leading to their classification as omnivorous. In addition to fish (occasionally even their own species), documented food items for piranhas include other vertebrates (mammals, birds, reptiles), invertebrates (insects, crustaceans), fruits, seeds, leaves and
detritus In biology, detritus () is dead particulate organic material, as distinguished from dissolved organic material. Detritus typically includes the bodies or fragments of bodies of dead organisms, and fecal material. Detritus typically hosts commun ...
. The diet often shifts with age and size. Research on the species ''Serrasalmus'' aff. ''brandtii'' and ''Pygocentrus nattereri'' in Viana Lake in Maranhão, which is formed during the wet season when the
Pindaré River The Pindaré River is a river in Maranhão state of north-central Brazil. The Pindaré rises in the low hills which separate its basin from that of the Tocantins River to the south. In its lower reaches it is called the Pindaré-Mirim. It is a lef ...
(a tributary of the
Mearim River The Mearim River is a river in Maranhão state of northern Brazil. The river originates in the southern part of Maranhão, and drains north into the Baía de São Marcos, an estuary that also receives the Pindaré and Grajaú rivers, which are ...
) floods, has shown that they primarily feed on fish, but also eat vegetable matter. In another study of more than 250 ''Serrasalmus rhombeus'' at Ji-Paraná (Machado) River, 75% to 81% (depending on season) of the stomach content was fish, but about 10% was fruits or seeds. In a few species such as ''Serrasalmus serrulatus'', the dietary split may be more equal, but this is less certain as based on smaller samples: Among 24 ''S. serrulatus'' from flooded forests of Ji-Paraná (Machado) River, there were several with fish remains in their stomachs, but half contained masticated seeds and in most of these this was the dominant item. Piranhas will often scavenge, and some species such as ''Serrasalmus elongatus'' are specialized scale-eaters, feeding primarily on scales and fins of other fish. Scale- and fin-eating is more widespread among juvenile and sub-adult piranhas. Piranhas lay their eggs in pits dug during the breeding season and swim around to protect them. Newly hatched young feed on zooplankton, and eventually move on to small fish once large enough.


Relationship with humans

Piranha teeth are often used as tools themselves (such as for carving wood or cutting hair) or to modify other tools (such as sharpening of darts). This behavior has been documented among several South American tribes including the Camayura and
Shavante The Xavante (also Shavante, Chavante, Akuen, A'uwe, Akwe, Awen, or Akwen) are an indigenous peoples, indigenous people, comprising 15,315 individuals within the territory of eastern Mato Grosso state in Brazil. They speak the Xavante language, pa ...
in Brazil and the
Pacahuara Pacahuara people are an indigenous people of Bolivia. A small group live in Tujuré, a community located near the Chácobo people on the Alto Ivón River in the Beni Department. The group only consists of four people. The fifth, a 57-year-old wo ...
in Bolivia. Piranhas are also popular as food. They are often considered a nuisance by fishers since they steal bait, eat catches, damage fishing gear and may bite when accidentally caught. Piranhas can be bought as pets in some areas but they are illegal in many parts of the United States and in the Philippines where importers face six months to four years in jail and the piranhas destroyed to prevent proliferation in the latter. The most common aquarium piranha is ''Pygocentrus nattereri'', the red-bellied piranha. Piranhas can be bought fully grown or as young, often no larger than a thumbnail. It is important to keep ''Pygocentrus'' piranhas alone or in groups of four or more, not in pairs, since aggression among them is common, not allowing the weaker fish to survive, and is distributed more widely when kept in larger groups. It is not uncommon to find individual piranhas with one eye missing due to a previous attack.


Attacks

Although often described as extremely dangerous in the media, piranhas typically do not represent a serious risk to humans. However, attacks have occurred, especially when the piranhas are in a stressed situation such as the dense groups that may occur when the water is lower during the dry season and food is relatively scarce. Swimming near fishermen may increase the risk of attacks due to the commotion caused by struggling fish and the presence of bait in the water. Splashing attracts piranhas and for this reason children are more often attacked than adults. Being in the water when already injured or otherwise incapacitated also increases the risk. There are sometimes warning signs at high-risk locations and beaches in such areas are sometimes protected by a barrier. Most piranha attacks on humans only result in minor injuries, typically to the feet or hands, but they are occasionally more serious and very rarely can be fatal. Near the city of Palmas in Brazil, 190 piranha attacks, all involving single bites to the feet, were reported in the first half of 2007 in an artificial lake which appeared after the damming of the Tocantins River. In the state of São Paulo, a series of attacks in 2009 in the Tietê River resulted in minor injuries to 15 people. In 2011, another series of attacks at
José de Freitas José de Freitas is a municipality in the state of Piauí in the Northeast region of Brazil. Tourism Located in the metropolitan region of Grande Teresina, Barragem do Bezerro, in the county of José de Freitas, is one of the most sought after ...
in the Brazilian state of
Piauí Piaui (, ) is one of the states of Brazil, located in the country's Northeast Region. The state has 1.6% of the Brazilian population and produces 0.7% of the Brazilian GDP. Piaui has the shortest coastline of any coastal Brazilian state at 66&n ...
resulted in 100 people being treated for bites to their toes or heels. On 25 December 2013, more than 70 bathers were attacked at
Rosario Rosario () is the largest city in the central provinces of Argentina, Argentine province of Santa Fe Province, Santa Fe. The city is located northwest of Buenos Aires, on the west bank of the Paraná River. Rosario is the third-most populous ci ...
in Argentina, causing injuries to their hands or feet.Mintz, Zoe (26 December 2013
Piranha Attack In Argentina Injures More Than 70, Fish Tore 'Bits Of Flesh' Off Swimmers On Christmas
ibtimes.com
In 2011, a drunk 18-year-old man was attacked and killed in Rosario del Yata,
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
. In 2012, a five-year-old Brazilian girl was attacked and killed by a shoal of ''P. nattereri''. In February 2015, a six-year-old girl died after being attacked by piranhas when her grandmother's boat capsized during a vacation in Brazil. Whereas fatal attacks on humans are rare, piranhas will readily feed on bodies of people that already have died, such as drowning victims.


Reputation

Various stories exist about piranhas, such as how they can skeletonize a
human body The human body is the structure of a Human, human being. It is composed of many different types of Cell (biology), cells that together create Tissue (biology), tissues and subsequently organ systems. They ensure homeostasis and the life, viabi ...
or cattle in seconds. These legends refer specifically to the red-bellied piranha. A common falsehood is that they can be attracted by blood and are exclusively carnivores. A Brazilian legend called "piranha cattle" states that they sweep the rivers at high speed and attack the first of the cattle entering the water allowing the rest of the group to traverse the river. These legends were dismissed through research by
Hélder Queiroz Hélder Lima de Queiroz () (born 1963) is a Brazilian conservation biologist, primatologist, and fish behaviorist. He is the Director of the Instituto de Desenvolvimento Sustentável Mamirauá (MISD) in Amazonas state, dedicated to protect ...
and Anne Magurran and published in '' Biology Letters''.


Theodore Roosevelt

When former US President Theodore Roosevelt visited Brazil in 1913, he went on a hunting expedition through the
Amazon Rainforest The Amazon rainforest, Amazon jungle or ; es, Selva amazónica, , or usually ; french: Forêt amazonienne; nl, Amazoneregenwoud. In English, the names are sometimes capitalized further, as Amazon Rainforest, Amazon Forest, or Amazon Jungle. ...
. While standing on the bank of the Amazon River, he witnessed a spectacle created by local fishermen. After blocking off part of the river and starving the piranhas for several days, they pushed a cow into the water, where it was quickly torn apart and skeletonized by a school of hungry piranhas. Roosevelt later described piranhas as vicious creatures in his 1914 book '' Through the Brazilian Wilderness''.


See also

* Animal bite * Animal attack


References


External links

* {{curlie, Recreation/Pets/Fish_and_Aquaria/Freshwater/Species/Characins/Piranhas/ * Eric J. Lyman
"Piranha meat could take a bite out of what ails you"
'' Houston Chronicle'', 17 July 1998
How to maintain healthy piranha in the home aquarium

"Piranha Feeding Frenzy"
National Geographic.com (2015-03-19) Fish common names Fish of South America Fish of the Amazon basin Fauna of the Pantanal Serrasalmidae Portuguese words and phrases