''Opegraphaceae'' is a

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.lichenicolous fungi A lichenicolous fungus is a parasitic fungus that only lives on lichen as the host. A lichenicolous fungus is not the same as the fungus that is the component of the lichen, which is known as a lichenized fungus. They are most commonly specific to ...

in the order

Arthoniales The Arthoniales is the second largest order of mainly crustose lichens, but fruticose lichens are present as well. The order contains around 1500 species, while the largest order with lichenized fungi, the Lecanorales, contains more than 14000 sp ...

. It was originally proposed by German lichenologist

Ernst Stizenberger Ernst Stizenberger (14 June 1827, in Konstanz – 27 September 1895) was a German physician and lichenologist. He studied medicine at the University of Freiburg, afterwards furthering his medical training in Prague and Vienna. In 1851 he returned ...

in 1862. It fell into disuse, but was resurrected in a

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

study of the order Arthoniales published in 2010. It now includes

taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

that were previously referred to the family

Roccellaceae The Roccellaceae are a family of fungi in the order Arthoniomycetes. Most taxa are lichenized with green algae, although some are lichenicolous, growing on other lichens. Genera , Species Fungorum (in the Catalogue of Life) accepts 47 genera an ...

, its

sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...

Description

Most taxa in the family have a

crustose Crustose is a habit of some types of algae and lichens in which the organism grows tightly appressed to a substrate, forming a biological layer. ''Crustose'' adheres very closely to the substrates at all points. ''Crustose'' is found on rocks and ...

thallus, although in rare cases it is

fruticose A fruticose lichen is a form of lichen fungi that is characterized by a coral-like shrubby or bushy growth structure. It is formed from a symbiotic relationship of a photobiont such as green algae or less commonly cyanobacteria and one, two or mor ...

. The lichens typically grow on bark,

wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin th ...

, or

rocks In geology, rock (or stone) is any naturally occurring solid mass or aggregate of minerals or mineraloid matter. It is categorized by the minerals included, its chemical composition, and the way in which it is formed. Rocks form the Earth's ...

. Some members, such as those in the genus '' Fouragea'', grow on leaves. The family has a

cosmopolitan distribution In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all or most of the world in appropriate habitats. Such a taxon, usually a species, is said to exhibit cosmopolitanism or cosmopolitism. The ext ...

, but is predominant in

tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...

and semi-

arid A region is arid when it severely lacks available water, to the extent of hindering or preventing the growth and development of plant and animal life. Regions with arid climates tend to lack vegetation and are called xeric or desertic. Most ar ...

subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...

regions. The

photobiont A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.green alga The green algae (singular: green alga) are a group consisting of the Prasinodermophyta and its unnamed sister which contains the Chlorophyta and Charophyta/Streptophyta. The land plants (Embryophytes) have emerged deep in the Charophyte alga as ...

l genus '' Trentepohlia''. A variety of secondary chemicals have been identified in this family. The most common types are

depside A depside is a type of polyphenolic compound composed of two or more monocyclic aromatic units linked by an ester bond. Depsides are most often found in lichens, but have also been isolated from higher plants, including species of the Ericaceae, L ...

depsidone Depsidones (+ "depside" + "one") are chemical compounds that are sometimes found as secondary metabolites in lichens. They are esters that are both depsides and cyclic ethers. An example is norstictic acid Norstictic acid is a depsidone produced ...

dibenzofuran Dibenzofuran is a heterocyclic organic compound with the chemical structure shown at right. It is an aromatic compound that has two benzene rings fused to a central furan ring. All the numbered carbon atoms have a hydrogen atom bonded to each of th ...

s, and

anthraquinone Anthraquinone, also called anthracenedione or dioxoanthracene, is an aromatic organic compound with formula . Isomers include various quinone derivatives. The term anthraquinone however refers to the isomer, 9,10-anthraquinone (IUPAC: 9,10-dioxo ...

Genera

Species Fungorum ''Index Fungorum'' is an international project to index all formal names (scientific names) in the fungus kingdom. the project is based at the Royal Botanic Gardens, Kew, one of three partners along with Landcare Research and the Institute of Mi ...

accepts 15 genera and 167 species in the family Opegraphaceae. This is a list of the genera in the Opegraphaceae based on a 2020 review and summary of fungal

classification Classification is a process related to categorization, the process in which ideas and objects are recognized, differentiated and understood. Classification is the grouping of related facts into classes. It may also refer to: Business, organizat ...

by Wijayawardene and colleagues. Following the genus name is the

taxonomic authority In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given ...

(those who first

circumscribed In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius. Not every polyg ...

the genus; standardized author abbreviations are used), year of publication, and the number of species: *'' Combea'' – 2 spp. *'' Cresponea'' – 22 spp. *'' Dictyographa'' – 2 spp. *'' Dolichocarpus'' – 2 spp. *'' Fouragea'' – 4 spp. *'' Ingaderia'' – 3 spp. *'' Nyungwea'' – 3 *''

Opegrapha ''Opegrapha'' is a genus of lichenized fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organi ...

'' (= '' Kalaallia'' ) – ca. 100 + ca. 200 orphaned *'' Paraingaderia'' – 1 sp. *'' Paralecanographa'' – 1 sp. *'' Paraschismatomma'' – 1 sp. *'' Pentagenella'' – 5 spp. *'' Schizopelte'' – 4 spp. *'' Sclerophyton'' – ca. 15 spp. *'' Sparria'' – 2 spp.

References

{{Taxonbar, from=Q28872118 Arthoniomycetes Lichen families Lichenicolous fungi Taxa described in 1862 Taxa named by Ernst Stizenberger Ascomycota families