''Nothofagus moorei'', commonly known as Antarctic beech, is an important

Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...

relict A relict is a surviving remnant of a natural phenomenon. Biology A relict (or relic) is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas. Geology and geomorphology In geology, a r ...

of the

rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...

s of the southern hemisphere. It occurs in wet, fire-free areas at high altitude in eastern

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

. The Antarctic beech group (

Nothofagaceae ''Nothofagus'', also known as the southern beeches, is a genus of 43 species of trees and shrubs native to the Southern Hemisphere in southern South America (Chile, Argentina) and Australasia (east and southeast Australia, New Zealand, New Gui ...

) is an ancient type of tree, of significance to southern hemisphere botanical distribution. Plants in the Nothofagaceae are currently found in southern South America (

Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

) and

Australasia Australasia is a region that comprises Australia, New Zealand and some neighbouring islands in the Pacific Ocean. The term is used in a number of different contexts, including geopolitically, physiogeographically, philologically, and ecologica ...

(east and southeast

New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...

New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...

and

New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...

Taxonomy

Ferdinand von Mueller Baron Sir Ferdinand Jacob Heinrich von Mueller, (german: Müller; 30 June 1825 – 10 October 1896) was a German-Australian physician, geographer, and most notably, a botanist. He was appointed government botanist for the then colony of Vict ...

described the Antarctic beech in 1866, from material collected near the

Bellinger River Bellinger River, an open and trained mature wave dominated, barrier estuary, is located in the Mid North Coast region of New South Wales, Australia. Course and features Bellinger River rises below Point Lookout within the Great Dividing Ra ...

by Charles Moore. Once referred to as 'negrohead beech', but now as 'Antarctic beech' (not to be confused with its South American relative, ''

Nothofagus antarctica ''Nothofagus antarctica'' (''Antarctic beech''; in Spanish ''Ñire'' or ''Ñirre'') is a deciduous tree or shrub native to southern Chile and Argentina from about 36°S to Tierra del Fuego (56° S), where it grows mainly in the diminishing tempe ...

'') is an

evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...

tree native to the eastern highlands of

. ''N. moorei'' proposed to be renamed ''Lophozonia moorei'' in 2013. The change in name from ''Nothofagus'' to ''Lophozonia'' is controversial. Within the genus, it is part of a lineage of three evergreen species, the other two being silver beech ('' N. menziesii'') of New Zealand and myrtle beech ('' N. cunninghamii'') of Tasmania and Victoria.

Description

These trees typically grow to 25 m (80 ft) tall and have large trunks to 1 m in diameter with scaly, dark brown bark. Maximum height is about 50 m. The

leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

are simple and alternate, growing six centimeters long. The leaf color is dark green, with new growth brilliant red, or orange in spring. The tree is deciduous in its native environment, but only partially deciduous in warmer areas, dropping half its leaves in autumn. The leaves are triangular to oblong with fine teeth along the crenate edges. The plants have separate male and female

flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...

s that occur on the same tree. The flowers are small and form inconspicuous clusters near the leaves towards the end of the branches. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...

, produced from December to February, is a small woody structure of four prickly valves. Each fruit contains three small winged nuts. Complicated root structures are frequently exhibited. These

root In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the sur ...

s would once have been

soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...

-covered, but have been exposed over the ages by

erosion Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is distin ...

, and covered in

moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...

and

lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. The Antarctic beech grows in

cool temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...

s from the

Barrington Tops Barrington Tops is part of the Great Dividing Range in New South Wales, Australia, between Gloucester and Scone. In 1934, the area was difficult to access and was described as being "not traceable to any man-made feature". Part of the area has ...

plateau in

New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...

, north to the

Lamington Plateau The Lamington National Park is a national park, lying on the Lamington Plateau of the McPherson Range on the Queensland/New South Wales border in Australia. From Southport on the Gold Coast the park is to the southwest and Brisbane is north. T ...

and

Springbrook Plateau Springbrook is a rural town and locality in the City of Gold Coast, Queensland, Australia. In the , the locality of Springbrook had a population of 659 people. The locality borders New South Wales. Geography The plateau is covered in subtrop ...

, in southern

Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...

, between altitudes of 480 m and 1550 m. It occurs in temperate to cool temperatures and with occasional snowfalls. Antarctic beech achieves its finest development at

Werrikimbe National Park The Werrikimbe National Park is a protected national park located in the catchment zone of the Upper Hastings River in New South Wales, Australia. Gazetted in 1975, the park is situated approximately north of Sydney, north-west of , and eas ...

and

Mount Banda Banda Mount Banda Banda, a mountain of the Mid North Coast region of New South Wales, Australia, is situated from Sydney within the Willi Willi National Park. Banda Banda can be seen on the north western horizon from Port Macquarie. And seen on th ...

Comboyne

There are four known populations of the Antarctic Beech in the

Comboyne Comboyne is a village on the Mid North Coast region of New South Wales. It is situated on the Comboyne Plateau, some 60 km south-west of Port Macquarie, 35 km west of Kew, New South Wales, Kew and 54 km north-west of Taree, New Sou ...

area of

. In 1925, the botanist E.C. Chisholm wrote that the Antarctic Beech at Comboyne was "extremely rare, although many trees were undoubtedly destroyed during clearing." The Comboyne Plateau was mostly cleared between 1900 and 1925. The Comboyne plateau is a scarp-bounded paleoplain located between the

Mid North Coast The Mid North Coast is a country region in the north-east of the state of New South Wales, Australia. The region covers the mid northern coast of the state, beginning from Port Stephens north of Sydney, and extending as far north as Woolgoolg ...

of New South Wales and the

Great Dividing Range The Great Dividing Range, also known as the East Australian Cordillera or the Eastern Highlands, is a cordillera system in eastern Australia consisting of an expansive collection of mountain ranges, plateaus and rolling hills, that runs rough ...

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

basalt Basalt (; ) is an aphanite, aphanitic (fine-grained) extrusive igneous rock formed from the rapid cooling of low-viscosity lava rich in magnesium and iron (mafic lava) exposed at or very near the planetary surface, surface of a terrestrial ...

s overlie much of the plateau,

creating Creation may refer to: Religion *''Creatio ex nihilo'', the concept that matter was created by God out of nothing *Creation myth, a religious story of the origin of the world and how people first came to inhabit it *Creationism, the belief that ...

relatively fertile red/brown

soils Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former term ...

. In the southern third of the plateau are underlying

Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...

sediments of the Lorne basin. The plateau has a wet, sub tropical climate, though subject to frost and occasional snow. The population at Comboyne was considered likely to be extinct by the scientific community, until published in 1994 by the botanists Bale and Williams. This community of trees regenerates well from seed and is notably vagile, with many young plants. It is the only other lowland (as low as 570 metres) population known, with those found near Dorrigo, to the north. The rainforest botanist

Alexander Floyd Alexander Geoffrey Floyd (1 April 1926 – 12 December 2022) was an Australian botanist with an expert knowledge of rainforest plants, particularly the rainforest trees of New South Wales. He has worked with the New South Wales Forestry Commiss ...

considers the Comboyne examples of the Antarctic Beech, as part of the cool temperate sub type 49, of the rainforests of New South Wales.

History

At one time it was believed that the Eastern Australian populations could not reproduce in present-day conditions, except by

sucker Sucker may refer to: General use * Lollipop or sucker, a type of confection * Sucker (slang), a slang term for a very gullible person * Hard candy ** Cough drop ** Mint (candy) Biology * Sucker (botany), a term for a shoot that arises undergro ...

ing (

asexual reproduction Asexual reproduction is a type of reproduction that does not involve the fusion of gametes or change in the number of chromosomes. The offspring that arise by asexual reproduction from either unicellular or multicellular organisms inherit the fu ...

), being remnant forest from a cooler time. It has since been shown that

sexual reproduction Sexual reproduction is a type of reproduction that involves a complex life cycle in which a gamete ( haploid reproductive cells, such as a sperm or egg cell) with a single set of chromosomes combines with another gamete to produce a zygote tha ...

may occur, but distribution in cool, isolated high-altitude environments at

temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...

and

tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...

latitude In geography, latitude is a coordinate that specifies the north– south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from –90° at the south pole to 90° at the north pol ...

s is consistent with the theory that the species was more prolific in a cooler age. The pattern of distribution around the southern Pacific Ocean rim has fed speculation that the dissemination of the genus dates to the time when

Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...

, Australia and

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...

were connected, the theoretical common land-mass referred to as

Nothofagus moorei in Lamington National Park Australia

It is an ornamental tree and cultivated specimens tolerate −7 °C (19 °F), though wild plants growing on

have withstood record low temperatures of −17 °C (1 °F), no source provenance have been selected from there and other mountains, highlands or plateaus for cultivation.Zoete, T. (2000) Vegetation survey of the Barrington Tops and Mount Royal National Parks for use in fire management. Cunninghamia 6, 511-578.

References

{{Taxonbar, from1=Q2709082, from2=Q42747110 Nothofagaceae Fagales of Australia Trees of Australia Flora of Queensland Flora of New South Wales Endemic flora of Australia Trees of mild maritime climate Garden plants of Australia Ornamental trees Plants described in 1866 Taxa named by Ferdinand von Mueller