''Nanyangosaurus'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of herbivorous

ornithischian Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek st ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

belonging to Hadrosauroidea that lived in the Late

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

of present-day

Henan Province Henan (; or ; ; alternatively Honan) is a landlocked province of China, in the central part of the country. Henan is often referred to as Zhongyuan or Zhongzhou (), which literally means "central plain" or "midland", although the name is al ...

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

. In 1994 the remains of ''Nanyangosaurus'' were found near the village of Houzhuang, in

Neixiang Neixiang County () is a county under the jurisdiction of Nanyang City, in the southwest of Henan province, China, has an area of and a population of 610,000 as of 2002. Neixiang is best known because it has China's best preserved Yamen, or count ...

county in

Henan Henan (; or ; ; alternatively Honan) is a landlocked province of China, in the central part of the country. Henan is often referred to as Zhongyuan or Zhongzhou (), which literally means "central plain" or "midland", although the name is al ...

. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

, ''Nanyangosaurus zhugeii'', was described by Xu Xing,

Zhao Xijin Zhao Xijin (赵喜进; born c. 1935 died July 21, 2012) was a Chinese paleontologist notable for having named numerous dinosaurs. He was a professor at Beijing's Institute of Vertebrate Paleontology and Paleoanthropology. Biography Zhao Xijin was ...

Lü Junchang Lü Junchang (; 1965 – 9 October 2018) was a Chinese palaeontologist and professor at the Institute of Geology, Chinese Academy of Geological Sciences. An expert on Mesozoic reptiles, he described and named dozens of dinosaur and pterosaur taxa ...

, Huang Wanbo, Li Zhanyang and

Dong Zhiming Dong Zhiming (Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; born January 1937) is a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) in Beijing. He began working at the ...

in 2000. The generic name is derived from the city of

Nanyang Nanyang is the romanization of two common Chinese place names. It may refer to: Written as 南洋 (Southern Ocean) * Nanyang (region), a Chinese term denoting the Southeast Asian lands surrounding the South China Sea ;China * Nanyang Fleet, Qing ...

. The specific name honours one of the most famous historic inhabitants of that city, the legendary strategist

Zhuge Liang Zhuge Liang ( zh, t=諸葛亮 / 诸葛亮) (181 – September 234), courtesy name Kongming, was a Chinese statesman and military strategist. He was chancellor and later regent of the state of Shu Han during the Three Kingdoms period. He is r ...

The specimen,

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

IVPP V 11821, was excavated in the Xiaguan Formation dating to the

Turonian The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by t ...

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...

stages. It consists of a partial skeleton lacking the skull. Eight posterior dorsal vertebrae, a

sacrum The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part ...

of six vertebrae and a tail of thirty-six vertebrae have been preserved, together with a partial

ischium The ischium () form ...

, a forelimb and a hindlimb. The describers considered ''Nanyangosaurus'' to be of Albian age because of its primitiveness, but the type horizon is now believed to be Turonian-Campanian in age based on plant and invertebrate fossils.Wang, D. et al., 2013. Discovery of invertebrate zoolite in the Xiaguan Formation of Xiaguan-Gaoqiu Basin, Henan, China, and its importance for stratigraphic subdivision comparison. ''Acta Geologica Sinica'' 87: 1049–1058. ''Nanyangosaurus'' was a rather small euornithpod with an estimated length of four to five metres. The length of the

femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with ...

is 517 millimetres. The forelimbs were relatively long with a long hand. The first digit of the hand was completely absent including the first

metacarpal In human anatomy, the metacarpal bones or metacarpus form the intermediate part of the skeletal hand located between the phalanges of the fingers and the carpal bones of the wrist, which forms the connection to the forearm. The metacarpal bones ...

; according to the describers this was not an accident of preservation but the actual condition of the living animal. The species would then not have possessed the thumb spike typical of its relatives. According to a

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...

analysis performed by the describers, ''Nanyangosaurus'' was a basal member of the

Iguanodontia Iguanodontia (the iguanodonts) is a clade of herbivorous dinosaurs that lived from the Middle Jurassic to Late Cretaceous. Some members include ''Camptosaurus'', ''Dryosaurus'', ''Iguanodon'', ''Tenontosaurus'', and the hadrosaurids or "duck-bi ...

, more derived than '' Probactrosaurus'' and closely related to the

Hadrosauroidea Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or hadrosaurids, and all dinosaurs more closely related to them than to ''Iguanodon''. Their remains have been recovered in Asia, Europ ...

Notes

{{Taxonbar, from=Q134000 Late Cretaceous dinosaurs of Asia Iguanodonts Fossil taxa described in 2000 Paleontology in Henan Taxa named by Lü Junchang Taxa named by Xu Xing Taxa named by Dong Zhiming Nanyang, Henan Ornithischian genera