Hadrosauroidea
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Hadrosauroidea
Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or hadrosaurids, and all dinosaurs more closely related to them than to ''Iguanodon''. Their remains have been recovered in Asia, Europe, Africa and the Americas. Many primitive hadrosauroids, such as the Asian ''Probactrosaurus'' and ''Altirhinus'', have traditionally been included in a paraphyly, paraphyletic (unnatural grouping) "Iguanodontidae". With cladistics, cladistic analysis, the traditional Iguanodontidae has been largely disbanded, and probably includes only ''Iguanodon'' and perhaps its closest relatives. Classification The cladogram below follows an analysis by Andrew McDonald, 2012, and shows the position of Hadrosauroidea within Styracosterna. The cladogram below follows an analysis by Wu Wenhao and Pascal Godefroit (2012). Cladogram after Prieto-Marquez and Norell (2010). A phylogenetic analysis performed by Ramírez-Velasco ''et al.'' (2012) foun ...
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Tethyshadros Insularis
''Tethyshadros'' ("Tethys Ocean, Tethyan hadrosauroid") is a genus of hadrosauroidea, hadrosauroid dinosaur from Trieste, Italy. The type and only species is ''T. insularis''. Discovery and naming Sometime in the 1980s, Alceo Tarlao and Giorgio Rimoli reported finding fragments of dinosaur bone while prospecting for rare bones. The abandoned quarry these were found in was only 100m inland, at Villagio del Pescatore, Trieste Province, Italy. It was from this quarry that a nearly complete hadrosaur skeleton was discovered in 1994. Lying on a vertical rockface, the specimen required a difficult excavation process, involving the removal of over 300 cubic metres of mineral and use of large equipment. Palaeontologist Fabio Della Vacchia among others served as scientific director for the excavation. Many other fossils, including various other hadrosaur specimens, were uncovered in the process. The main skeleton was not extracted until 1999. The significance of the find was immediately ...
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Datonglong
''Datonglong tianzhenensis'' is an herbivorous ornithischian dinosaur belonging to the Hadrosauroidea, which lived in the Late Cretaceous period in present-day China. It is the type species of the genus ''Datonglong''. Discovery and naming In 2008, a team from the Shanxi Museum of Geological and Mineral Science and Technology in the Kangdailiang quarry in Shanxi discovered the jaw of a euornithopod. In 2016, the species ''Datonglong tianzhenensis'' was named and described by Xu Shicha, You Hailu, Wang Jiawei, Wang Suozhu, Yi Jian, and Jia Lei. The generic name refers to the city of Datong and the Chinese word ''long'' (龍), which means “dragon”. The specific name refers to its origins in Tianzhen county. ''Datonglong'' was one of eighteen dinosaur taxa from 2015 to be described in open access or free-to-read journals. Description The holotype, SXMG V 00005, consists of a portion of the lower jaw with some preserved teeth. The fossil was found in a layer of the Huiqu ...
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Probactrosaurus
''Probactrosaurus'' (meaning "before ''Bactrosaurus''") is an early herbivorous hadrosauroid iguanodont dinosaur. It lived in China during the Late Cretaceous period. Discovery and species In 1959 and 1960 a Soviet-Chinese expedition uncovered the remains of a euornithopod in Inner Mongolia near Maortu. The type species is ''Probactrosaurus gobiensis'', described and named by A. K. Rozhdestvensky in 1966. The generic name refers to Rozhdestvensky's hypothesis that ''Probactrosaurus'' would be the direct ancestor of ''Bactrosaurus'', a notion now discarded. The specific name refers to the Gobi desert. The holotype specimen, PIN 2232/1, a partial skeleton with skull, was found in layers of the Dashuigou Formation. Another partial skeleton, PIN 2232-10, was found along with numerous other fragments. In 1966 Rozhdestvensky also named a second species, ''Probactrosaurus alashanicus'', based on fragmentary material. Its specific name refers to the Alxa League. In 2002 David B. ...
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Zuoyunlong
''Zuoyunlong'' is an extinct genus of herbivorous ornithischian dinosaur belonging to the Hadrosauroidea, that lived during the Late Cretaceous in the area of present China. Discovery and naming The holotype, SXMG V 00 004, was found by the team of the Shanxi Regional Geological Survey in a layer of the Zhumapu Formation in the province of Shanxi which dates from the Cenomanian, roughly ninety-five million years old. It consists of two bones from the right-hand half of the pelvis, a partial right ilium with the field number ZY004-001 to which the cover sheet is missing, and the lower end of the shaft of the right ischium, field number ZY004-002. Additional specimens are unassigned. In 2017 the type species ''Zuoyunlong huangi'' was named and described by Wang Runfu, You Hailu, Wang Suozhu, Shichao Xu, Yi Jian, Xie Lijuan, Jia and Lei Xing Hai. The generic name refers to the prefecture of Zuoyun with the Chinese word ''long'', "dragon". The specific name honors the Chinese paleo ...
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Ouranosaurus
''Ouranosaurus'' is a genus of herbivorous basal hadrosauriform dinosaur that lived during the Aptian stage of the Early Cretaceous of modern-day Niger and Cameroon. ''Ouranosaurus'' measured about long. Two rather complete fossils were found in the Elrhaz Formation, Gadoufaoua deposits, Agadez, Niger, in 1965 and 1970, with a third indeterminate specimen known from the Koum Formation of Cameroon. The animal was named in 1976 by French paleontologist Philippe Taquet; the type species being ''Ouranosaurus nigeriensis''. The generic name is a combination of ''ourane,'' a word with multiple meanings ("an Arabic name that signifies valor, courage, recklessness"; the Tuareg name of the desert monitor given by the Tuareg Berbers of Niger & Algeria), and ''sauros'', the Greek word for lizard. The specific epithet ''nigeriensis'' alludes to ''Niger'', its country of discovery (in Latin, the adjectival suffix ''-iensis'' means "originating from"). And so, ''Ouranosaurus nigeriensis'' co ...
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Hadrosauromorpha
Hadrosauromorpha is a clade of iguanodontian ornithopods, defined in 2014 by David B. Norman to divide Hadrosauroidea into the basal taxa with compressed manual bones and a pollex, and the derived taxa that lack them. The clade is defined as all the taxa closer to ''Edmontosaurus regalis'' than '' Probactrosaurus gobiensis''. This results in different taxon inclusion depending on the analysis. Classification Hadrosauromorpha was first used in literature by David B. Norman in 2014 in a discussion of phylogenetics of ''Hypselospinus''. In his 2014 paper Norman references another of his publications as the authority for Hadrosauromorpha, a chapter in the book ''Hadrosaurs''. However, the book was in fact published later, in 2015. Following Article 19.4 of the PhyloCode, the authorship of the clade is thus Norman (2015), while the authorship of the definition is Norman (2014). Definition Hadrosauromorpha was defined by Norman (2014 and 2015) as hadrosauroid taxa closer to ''Edmon ...
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Jinzhousaurus
''Jinzhousaurus'' is a genus of hadrosauroid dinosaur of the Early Cretaceous of China. The type species is ''Jinzhousaurus yangi''. The generic name refers to the town Jinzhou. The specific name honours Yang Zhongjian as the founder of Chinese paleontology. It was first described by Wang Xiao-lin and Xu Xing in 2001. Discovery Its fossil, holotype IVPP V12691, was found near Baicaigou in Yixian County in the Dawangzhangzi Beds of the Yixian Formation in Liaoning, China, having an oldest determinable age of 122 million years, during the early Aptian stage of the early Cretaceous Period.Zhou, Z. (2006). "Evolutionary radiation of the Jehol Biota: chronological and ecological perspectives." ''Geological Journal'', 41: 377-393. It consists of a nearly complete skeleton, compressed on a slab. Description ''Jinzhousaurus'' has a length of about 7 metres (23 ft) and its skull is about half a metre long. Its snout was elongated with large nares and lacking an antorbi ...
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Equijubus
''Equijubus'' (; ''Mǎzōng'' meaning "horse mane" after the area Mǎzōng Mountain 马鬃山 in which it was found), is a genus of herbivorous hadrosauroid dinosaur from the Early Cretaceous (Albian stage) of northwestern China. Discovery and naming The type (and only known) specimen was found in the summer of 2000 by a Chinese-American expedition in the Mazong (= "horse mane") Shan area of China's Gansu Province.You, Luo, Shubin, Witmer, Tang and Tang (2003). "The earliest-known duck-billed dinosaur from deposits of late Early Cretaceous age in northwest China and hadrosaurid evolution." ''Cretaceous Research'', 24: 347-353. In 2002 You Hialu in a dissertation named and described the species ''Equijubus normani''. The generic name is derived from Latin ''equus'', "horse", and ''juba'', "mane". The specific epithet "normani" is in honour of British palaeontologist David B. Norman. However, such a ''nomen ex dissertatione'' does not constitute a valid name. The type ...
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Portellsaurus
''Portellsaurus'' (meaning "Portell Lizard") is a genus of hadrosauroid dinosaur from the Lower Cretaceous Margas de Mirambell Formation of Spain. The genus contains a single species, ''Portellsaurus sosbaynati'', known from a partial right dentary. Discovery and naming ''Portellsaurus'' was discovered in 1998 by Miquel Guardiola, Julián Yuste and Silvia Fabregat in a site near Mas de Curolles. The generic name, ''Portellsaurus'', comes from the Portell de Morella, the town where the holotype was recovered, and the specific name, ''sosbaynati'', refers to Vicente Sos Baynat, a Spanish geologist who was the first scientist honored by Universitat Jaume I as "Doctor Honoris Causa". Description ''Portellsaurus'' is known from an almost complete right dentary, MQ98-II-1, which is stored at the Colección Museográfica de Cinctorres in Castellón. ''Portellsaurus'' can be distinguished from other styracosternans through the possession of two unique features: the absence of a bulg ...
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Koshisaurus
''Koshisaurus'' is a monospecific genus of basal hadrosauroid from the Kitadani Formation in Japan. The discovery of the genus suggests that hadrosauroids had higher diversity along the eastern margin of Asia in the Early Cretaceous. "Koshi" means an old Japanese regional name including Fukui prefecture where fossils of the genus were discovered. Description ''Koshisaurus'' was unlike most hadrosauroids due to the fact that it possessed an antorbital fossa, as well as three subsidiary ridges on its maxillary teeth, similar to those of '' Equijubus''. These ridges are also present on the also Asian genera '' Xuwulong'', ''Jinzhousaurus'' and '' Altirhinus''; but all three lack antorbital fossae, placing them as more advanced than ''Koshisaurus''., See also * Timeline of hadrosaur research A timeline is a display of a list of events in chronological order. It is typically a graphic design showing a long bar labelled with dates paralleling it, and usually contemporane ...
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Gongpoquansaurus
''Gongpoquansaurus'' (meaning "Gongpoquan reptile") is an extinct genus of basal hadrosauroid dinosaur that was not formally named until 2014, while the name was a ''nomen nudum'' for many years previously. It is known from IVPP V.11333, a partial skull and postcranial skeleton. It was collected in 1992 at locality IVPP 9208–21, from the Albian Zhonggou Formation (Xinminpu Group), in Mazongshan, Gansu Province, China. The specimen was first described and named by Lü Junchang in 1997 as the third species of ''Probactrosaurus'', ''Probactrosaurus mazongshanensis''.Lü J., 1997, "A new Iguanodontidae (''Probactrosaurus mazongshanensis'' sp. nov.) from Mazongshan area, Gansu Province, China", In: Z. Dong (ed.), ''Sino-Japanese Silk Road Dinosaur Expedition''. China Ocean Press, Beijing pp. 27-47 Following its description, several studies found it to be less derived than the type species of ''Probactrosaurus'' in relation to Hadrosauridae Hadrosaurids (), or duck-billed dinosau ...
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Mantellisaurus
''Mantellisaurus'' is a genus of iguanodontian dinosaur that lived in the Barremian and early Aptian ages of the Early Cretaceous Period of Europe. Its remains are known from Belgium (Bernissart), England, Spain and Germany. The type and only species is ''M. atherfieldensis''. Formerly known as ''Iguanodon atherfieldensis'', the new genus ''Mantellisaurus'' was erected for the species by Gregory Paul in 2007. According to Paul, ''Mantellisaurus'' was more lightly built than ''Iguanodon'' and more closely related to ''Ouranosaurus'', making ''Iguanodon'' in its traditional sense paraphyletic. It is known from many complete and almost complete skeletons. The genus name honours Gideon Mantell, the discoverer of ''Iguanodon''. History of discovery Discovery and naming of the holotype The holotype fossil, NHMUK R5764, was originally discovered by Reginald Walter Hooley in 1914 in the upper Vectis Formation of southern England and reported upon in 1917. He posthumously named it ''Ig ...
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