Myiagra
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''Myiagra'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of passerine birds in the family
Monarchidae The monarchs (family Monarchidae) comprise a family of over 100 passerine birds which includes shrikebills, paradise flycatchers, and magpie-larks. Monarchids are small insectivorous songbirds with long tails. They inhabit forest or woodland a ...
, the monarch flycatchers, native to Australasia, sometimes referred to as the broad-billed flycatchers or simply broadbills.


Taxonomy

The genus ''Myiagra'' was introduced in 1827 by the naturalists
Nicholas Vigors Nicholas Aylward Vigors (1785 – 26 October 1840) was an Irish zoologist and politician. He popularized the classification of birds on the basis of the quinarian system. Early life Vigors was born at Old Leighlin, County Carlow on 1785 as fi ...
and
Thomas Horsfield Thomas Horsfield (May 12, 1773 – July 24, 1859) was an American physician and natural history, naturalist who worked extensively in Indonesia, describing numerous species of plants and animals from the region. He was later a curator of the Eas ...
. The name combines the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''muia'' meaning "a fly" and ''agreō'' meaning "to seize".
Myiagros In ancient Greek religion, Myiagros ("He Who Chases the Flies") or Myacoris was a cult title for a divine figure who warded off flies. It could be used as an epithet for either a divinity or a hero. Pausanias characterizes Myiagros as a diviniz ...
was also the name of a Greek god. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
was designated by
George Robert Gray George Robert Gray FRS (8 July 1808 – 6 May 1872) was an English zoologist and author, and head of the ornithological section of the British Museum, now the Natural History Museum, in London for forty-one years. He was the younger brother o ...
in 1840 as the
leaden flycatcher The leaden flycatcher (''Myiagra rubecula'') is a species of passerine bird in the family Monarchidae. Around 15 cm (6 in) in length, the male is lustrous azure with white underparts, while the female possesses leaden head, mantle and ...
.


Species

The genus contains 22 species include one that is now extinct: *
Oceanic flycatcher The oceanic flycatcher (''Myiagra oceanica'') is a species of bird in the family Monarchidae. It is endemic to Micronesia and can be found on the Caroline Islands. The diversity of traits in the Oceanic flycatchers resulted from three waves of c ...
(''Myiagra oceanica'') *
Palau flycatcher The Palau flycatcher (''Myiagra erythrops'') is a species of bird in the family Monarchidae. It is endemic to Palau Palau,, officially the Republic of Palau and historically ''Belau'', ''Palaos'' or ''Pelew'', is an island country a ...
(''Myiagra erythrops'') * †
Guam flycatcher The Guam flycatcher (''Myiagra freycineti''), or Guam broadbill, is an extinct species of bird in the family Monarchidae formerly endemic to Guam. Taxonomy and systematics Some authorities consider the Guam flycatcher to have been a subspecies o ...
(''Myiagra freycineti'') (extinct) *
Pohnpei flycatcher The Pohnpei flycatcher (''Myiagra pluto''), known as Koikoi in Pohnpeian, is a species of bird in the family Monarchidae. It is endemic to Micronesia and can be found on the Caroline Islands The Caroline Islands (or the Carolines) are ...
(''Myiagra pluto'') * Moluccan flycatcher (''Myiagra galeata'') * Biak black flycatcher (''Myiagra atra'') *
Leaden flycatcher The leaden flycatcher (''Myiagra rubecula'') is a species of passerine bird in the family Monarchidae. Around 15 cm (6 in) in length, the male is lustrous azure with white underparts, while the female possesses leaden head, mantle and ...
(''Myiagra rubecula'') *
Steel-blue flycatcher The steel-blue flycatcher (''Myiagra ferrocyanea'') is a species of bird in the family Monarchidae. It is found in Papua New Guinea and Solomon Islands. Taxonomy and systematics Alternate names for the steel-blue flycatcher include Solomons fly ...
(''Myiagra ferrocyanea'') * Makira flycatcher (''Myiagra cervinicauda'') *
Melanesian flycatcher The Melanesian flycatcher (''Myiagra caledonica'') is a species of bird in the monarch-flycatcher family Monarchidae. The species is found on islands in Melanesia. Taxonomy and systematics It is closely related to the Vanikoro flycatcher. Alte ...
(''Myiagra caledonica'') * Vanikoro flycatcher (''Myiagra vanikorensis'') * Samoan flycatcher (''Myiagra albiventris'') *
Azure-crested flycatcher The azure-crested flycatcher (''Myiagra azureocapilla'') or the blue-crested flycatcher, is a species of bird in the monarch flycatcher family Monarchidae. It is endemic to Fiji, where it is found on Taveuni. Its natural habitats are subtropica ...
(''Myiagra azureocapilla'') * Chestnut-throated flycatcher (''Myiagra castaneigularis'') *
Broad-billed flycatcher The broad-billed flycatcher (''Myiagra ruficollis'') is a species of bird in the family Monarchidae. It is found in northern Australia, the Lesser Sunda Islands and southern New Guinea. Its natural habitats are subtropical or tropical moist ...
(''Myiagra ruficollis'') *
Satin flycatcher The satin flycatcher (''Myiagra cyanoleuca'') is a species of bird in the family Monarchidae. Males stand out with their blue-black feathers contrasting their white bellies, and the females with their bright orange throats. It breeds mostly in s ...
(''Myiagra cyanoleuca'') *
Shining flycatcher The shining flycatcher (''Myiagra alecto'') is a species of bird in the family Monarchidae. It is found in northern Australia, and from the Moluccas to the Bismarck Archipelago. Its natural habitats are subtropical or tropical moist lowland f ...
(''Myiagra alecto'') * Mussau flycatcher (''Myiagra hebetior'') * Velvet flycatcher (''Myiagra eichorni'') * Dyaul flycatcher (''Myiagra cervinicolor'') *
Paperbark flycatcher The paperbark flycatcher (''Myiagra nana''), also known as the little restless flycatcher, is a passerine bird in the family Monarchidae. It occurs in tropical woodland and riverine habitats of northern Australia and southern New Guinea. Previou ...
(''Myiagra nana'') *
Restless flycatcher The restless flycatcher (''Myiagra inquieta'') or restless myiagra is a passerine bird in the family Monarchidae; it is also known as the razor grinder or scissors grinder because of its distinctive call. It is a native of eastern and southern A ...
(''Myiagra inquieta'')


Former species

Formerly, some authorities also considered the following species (or subspecies) as species within the genus ''Myiagra'': * Andamanen black-naped blue monarch (as ''Myiagra tytleri'') *
Pale-blue monarch The pale-blue monarch (''Hypothymis puella'') is a small passerine bird in the family Monarchidae endemic to eastern Indonesia. Taxonomy and systematics The pale-blue monarch was originally described in the genus '' Myiagra'' and then classified ...
(as ''Myiagra puella'') * White-collared monarch (as ''Piezorhynchus vidua'') * White-collared monarch (squamulatus) (as ''Piezorhynchus squamulatus'') *
White-capped monarch The white-capped monarch (''Monarcha richardsii''), or Richards' monarch, is a species of bird in the family Monarchidae. It is endemic to the Western Province in the Solomon Islands Solomon Islands is an island country consisting of ...
(as ''Piezorhynchus Richardsii'')


Description

Members of this genus are separated from other members of the family, particularly the ''
Monarcha ''Monarcha'' is a genus of bird in the family Monarchidae. They are found in Australia and Melanesia. Taxonomy and systematics The genus ''Monarcha'' was introduced by naturalists Nicholas Vigors and Thomas Horsfield in 1827 with the black-fac ...
'' monarchs they resemble, by their consistent
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
, egg patterning, and the crested heads that often lack facial patterns (a few ''Monarcha'' monarchs also lack facial patterns). Flycatchers in the genus ''Myiagra'' are small birds ranging in length from 13–20 cm in length. They have broad flattened
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
s adapted to catching insects. The bill is usually black, the exception being the blue-crested flycatcher of Fiji, whose bill is bright orange. The insides of their mouths are bright orange, noticeable when singing. When perched they have an upright stance. The satiny
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
of all species is sexually dichromatic, to a greater or lesser degree. Overall males tend to have dark blue or black heads, backs, wings and tails and pale bellies and rumps. Some males have red underparts, others have the dark upper plumage all over, and the slightly aberrant shining flycatchers have chestnut wings, backs and tails. The females follow a similar pattern to males but have lighter, more washed out colours. When perched the species in this genus constantly moves their closed tail, in the
restless flycatcher The restless flycatcher (''Myiagra inquieta'') or restless myiagra is a passerine bird in the family Monarchidae; it is also known as the razor grinder or scissors grinder because of its distinctive call. It is a native of eastern and southern A ...
the tail is shacked from side to side. The orange lining to their mouths is visible when they sing. Members of this genus differ from the closely related genus ''
Monarcha ''Monarcha'' is a genus of bird in the family Monarchidae. They are found in Australia and Melanesia. Taxonomy and systematics The genus ''Monarcha'' was introduced by naturalists Nicholas Vigors and Thomas Horsfield in 1827 with the black-fac ...
'' in having their crested heads often unpatterned, and the sexes being different in appearance. They have satiny plumage and perch upright, their long tails tending to move frequently. Males typically have dark blue heads and upper parts and paler underparts, and females are mostly similar in appearance to the males but their colour is more washed out and less intense. Their broad beaks are adapted to feed on insects, which they mostly catch on the wing.


Vocalizations

The calls are generally unmusical and are described as guttural, harsh and rasping.


Behaviour and ecology

The genus is adapted to feeding on insects in a manner similar to the Old World flycatchers in the family
Muscicapidae The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Norther ...
. Prey is obtained by
sallying Hawking is a feeding strategy in birds involving catching flying insects in the air. The term usually refers to a technique of sallying out from a perch to snatch an insect and then returning to the same or a different perch, though it also appli ...
from a perch to obtain flying insects or by hover-gleaning, snatching insects from the undersides of leaves while in flight.


Distribution and habitat

The genus is spread across
Australasia Australasia is a region that comprises Australia, New Zealand and some neighbouring islands in the Pacific Ocean. The term is used in a number of different contexts, including geopolitically, physiogeographically, philologically, and ecologica ...
, with some representatives in
Wallacea Wallacea is a biogeographical designation for a group of mainly Indonesian islands separated by deep-water straits from the Asian and Australian continental shelves. Wallacea includes Sulawesi, the largest island in the group, as well as Lo ...
,
New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...
,
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, the
Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capita ...
,
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
,
Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (french: link=no, République de Vanuatu; bi, Ripablik blong Vanuatu), is an island country located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of no ...
,
Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...
,
Samoa Samoa, officially the Independent State of Samoa; sm, Sāmoa, and until 1997 known as Western Samoa, is a Polynesian island country consisting of two main islands (Savai'i and Upolu); two smaller, inhabited islands (Manono Island, Manono an ...
and
Micronesia Micronesia (, ) is a subregion of Oceania, consisting of about 2,000 small islands in the western Pacific Ocean. It has a close shared cultural history with three other island regions: the Philippines to the west, Polynesia to the east, and ...
. Members of this genus occur as occasional
vagrant Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, temporar ...
s in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
and formerly occurred in
Tonga Tonga (, ; ), officially the Kingdom of Tonga ( to, Puleʻanga Fakatuʻi ʻo Tonga), is a Polynesian country and archipelago. The country has 171 islands – of which 45 are inhabited. Its total surface area is about , scattered over in ...
and
Guam Guam (; ch, Guåhan ) is an organized, unincorporated territory of the United States in the Micronesia subregion of the western Pacific Ocean. It is the westernmost point and territory of the United States (reckoned from the geographic cent ...
.


References

* del Hoyo, J.; Elliot, A. & Christie D. (editors). (2006).
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. T ...
. Volume 11: Old World Flycatchers to Old World Warblers. Lynx Edicions. . {{Taxonbar, from=Q646544 Bird genera   Taxonomy articles created by Polbot