Mochlodon
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''Mochlodon'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
rhabdodontid Rhabdodontidae is a family of herbivorous iguanodontian ornithopod dinosaurs whose earliest stem members appeared in the middle of the Lower Cretaceous. The oldest dated fossils of these stem members were found in the Barremian Castrillo de la R ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
of
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
and
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
. It lived during the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
(85-80 Ma) and two species are known: ''M. suessi'' and ''M. vorosi''.


Discovery and species

In 1859 coal mine administrator Pawlowitsch notified the
University of Vienna The University of Vienna (german: Universität Wien) is a public research university located in Vienna, Austria. It was founded by Duke Rudolph IV in 1365 and is the oldest university in the German-speaking world. With its long and rich histor ...
that some fossils had been found in the ''Gute Hoffnung'' mine at Muthmannsdorf in
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
. A team headed by geologists
Eduard Suess Eduard Suess (; 20 August 1831 - 26 April 1914) was an Austrian geologist and an expert on the geography of the Alps. He is responsible for hypothesising two major former geographical features, the supercontinent Gondwana (proposed in 1861) and t ...
and
Ferdinand Stoliczka Ferdinand Stoliczka (Czech written Stolička, 7 June 1838 – 19 June 1874) was a Moravian palaeontologist who worked in India on paleontology, geology and various aspects of zoology, including ornithology, malacology, and herpetology. He died of ...
subsequently uncovered numerous bones of several species, among them those of a euornithopod dinosaur. Stored at the university museum, the finds remained undescribed until they were studied by
Emanuel Bunzel Emanuel Bunzel (1828–1895), was an Austrian paleontologist. Biography Emanuel Bunzel was born in Prague, Kingdom of Bohemia (today the Czech Republic), in 1828. In 1871, he described a skull fragment found in an Austrian coal mine in 1859 by ...
from 1870 onwards. Bunzel in 1871 named the euornithopod a new species of ''
Iguanodon ''Iguanodon'' ( ; meaning 'iguana-tooth'), named in 1825, is a genus of iguanodontian dinosaur. While many species have been classified in the genus ''Iguanodon'', dating from the late Jurassic Period to the early Cretaceous Period of Asia, Eu ...
'': ''Iguanodon suessii''. The specific name honours Suess and is today more often spelled ''suessi''. In 1881
Harry Govier Seeley Harry Govier Seeley (18 February 1839 – 8 January 1909) was a British paleontologist. Early life Seeley was born in London on 18 February 1839, the second son of Richard Hovill Seeley, a goldsmith, and his second wife Mary Govier. When his fat ...
named a separate genus: ''Mochlodon''. The generic name is derived from Greek ''mokhlos'', "bar", and ''odon'', "tooth", a reference to the bar-like median ridge on the teeth. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Mochlodon suessi''. ''Mochlodon'' and ''
Struthiosaurus ''Struthiosaurus'' (Latin ''struthio'' = ostrich + Greek ''sauros'' = lizard) is a genus of nodosaurid dinosaurs, from the Late Cretaceous period (Santonian-Maastrichtian) of Austria, Romania, France and Hungary in Europe.
'', the latter found at the same site, are the only dinosaur genera named from Austrian finds. The
type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...
PIUW 2349 was found in the
Grünbach Formation The Grünbach Formation is an Austrian geological formation that dates to the lower Campanian age of the Late Cretaceous. it forms part of the Gosau Group, and represents a marine regression event, representing a coastal/brackish environment, bein ...
of the
Gosau Group The Gosau Group (German: ''Gosau-Gruppe'') is a geological stratigraphic group in Austria, Germany and western Slovakia whose strata date back to the Late Cretaceous to Eocene.Höfling, 1985McCann, 2008 It is exposed in numerous sporadic isolated ...
dating from the Lower
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
, about 80 million years old. It consists of a
dentary In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...
, two
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
e (presently lost), a parietal, a scapula, an ulna, a manual
ungual An ungual (from Latin ''unguis'', i.e. ''nail'') is a highly modified distal toe bone which ends in a hoof, claw, or nail. Elephants and ungulates have ungual phalanges, as did the sauropods and horned dinosaurs. A claw is a highly modified ungual ...
, a femur and a tibia. Bunzel did not assign a
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
. In 2005 the dentary was chosen as the
lectotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...
. At the end of the nineteenth century Baron
Franz Nopcsa Franz may refer to: People * Franz (given name) * Franz (surname) Places * Franz (crater), a lunar crater * Franz, Ontario, a railway junction and unorganized town in Canada * Franz Lake, in the state of Washington, United States – see ...
noted the similarity of fossils found in
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...
to both the French ''
Rhabdodon ''Rhabdodon'' (meaning "fluted tooth") is a genus of ornithopod dinosaur that lived in Europe approximately 70-66 million years ago in the Late Cretaceous. It is similar in build to a very robust "hypsilophodont" (non-iguanodont ornithopod), thou ...
'' and the Austrian ''Mochlodon''. In 1899 he named some of these ''Mochlodon inkeyi'', the specific name honouring Béla Inkey, but the same year changed the name into ''Rhabdodon inkeyi''. In 1900 Nopcsa named some Romanian remains ''Mochlodon robustum'' (emended to ''M. robustus'' in 1990 by George Olshevsky). In 1915 however, he concluded that all this material could be referred to ''Rhabdodon'', the Austrian remains to ''Rhabdodon priscus''. In later years, ''Mochlodon'' was often considered a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
''. In 2003, when ''M. robustus'' was renamed ''
Zalmoxes ''Zalmoxes'' is an extinct genus of rhabdodontid ornithopod dinosaur from the Maastrichtian of Romania. The genus is known from specimens first named as the species '' Mochlodon robustum'' in 1899 by Franz Nopcsa before being reclassified as ...
'', ''Mochlodon'' was tentatively reinstated as a separate genus for the species ''Mochlodon suessi''. In 2005 a study concluded that no unique derived features,
autapomorphies In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...
, could be established for ''Mochlodon'' in relation to ''Zalmoxes'', assigning the Austrian remains provisionally to a ''Zalmoxes'' sp.; a definite identification would give ''Mochlodon'' nomenclatural priority. A second species, ''M. vorosi'', was described from the
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. The ...
aged
Csehbánya Formation The Csehbánya Formation is a Formation (geology), geological formation in the Transdanubian Mountains of Veszprém County, Hungary. The formation dates to the Late Santonian (around 85-84 million years ago) of the Late Cretaceous. It represents ...
of
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
in 2012.


References

{{Taxonbar, from=Q18578509 Iguanodonts Campanian life Santonian life Late Cretaceous dinosaurs of Europe Cretaceous Austria Fossils of Austria Fossils of Hungary Fossil taxa described in 1881 Taxa named by Harry Seeley Ornithischian genera