Mesodermochelys Undulatus
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''Mesodermochelys'' is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
sea turtle Sea turtles (superfamily Chelonioidea), sometimes called marine turtles, are reptiles of the order Testudines and of the suborder Cryptodira. The seven existing species of sea turtles are the flatback, green, hawksbill, leatherback, loggerhe ...
known from the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...
to the
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval ...
(Late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
) of what today is Japan.Hirayama R. (2007). "Cranial morphology of ''Mesodermochelys'' (Chelonioidea; Testudines) from the Late Cretaceous of Japan". ''Journal of Vertebrate Paleontology'' 27 (3): 89A
online, retrieved 28 July 2008
/ref> One species is known, the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
''M. undulatus''; it was given its binomial name by Ren Hirayama and Tsutomu Chitoku in 1996.Hirayama R, Fujii A, Takahashi K. (2006). "A dermochelyid sea turtle from the Upper Cretaceous (Late Campanian) Izumi Group of Shionoe, Takamatsu, Kagawa Prefecture, Western Japan". ''Fossils (Palaeontological Society of Japan)'' 80: 17-20
online, retrieved 28 July 2008
/ref> Studies of its skull indicate that it was a primitive member of the Dermochelyidae ( leatherback turtle family) that was closely related to the
Protostegidae Protostegidae is a family of extinct marine turtles that lived during the Cretaceous period. The family includes some of the largest sea turtles that ever existed. The largest, '' Archelon'', had a head long. Like most sea turtles, they had fl ...
. It has been described as the best representative of
Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretace ...
dermochelyids.


Description

Like other dermochelyids, ''Mesodermochelys'' had elongated front flippers. One fossil found in Japan's Kagawa Prefecture had a carapace estimated to be 1.3 metres in length. Only the neural or spinal
scute A scute or scutum (Latin: ''scutum''; plural: ''scuta'' "shield") is a bony external plate or scale overlaid with horn, as on the shell of a turtle, the skin of crocodilians, and the feet of birds. The term is also used to describe the anterior po ...
s, or individual plates, of the carapace are well grooved.


Specimens

The holotype specimen of ''Mesodermochelys undulatus'' Hobetsu Museum (HMG) 5 was collected in 1980 from the
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval ...
Hakobuchi Formation (
Yezo Group The Yezo Group is a stratigraphic group in Hokkaido, Japan and Sakhalin, Russia which is primarily Late Cretaceous in age ( Aptian to Earliest Paleocene). It is exposed as roughly north–south trending belt extending 1,500 kilometres through cen ...
) in Inasato, Hobetsu,
Hokkaido is Japan's second largest island and comprises the largest and northernmost prefecture, making up its own region. The Tsugaru Strait separates Hokkaidō from Honshu; the two islands are connected by the undersea railway Seikan Tunnel. The lar ...
, Japan. It comprises a supraoccipital and partial postcranial skeleton and was reported in 1996 along with 14 referred specimens collected between 1977 and 1994 and also reposited at HMG. In October 2005, Yoshiyuki Hattori uncovered a skull presumed to be ''Mesodermochelys'', from an unnamed
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. ...
-aged interval of the
Yezo Group The Yezo Group is a stratigraphic group in Hokkaido, Japan and Sakhalin, Russia which is primarily Late Cretaceous in age ( Aptian to Earliest Paleocene). It is exposed as roughly north–south trending belt extending 1,500 kilometres through cen ...
in the
Hokkaido Prefecture is Japan's second largest island and comprises the largest and northernmost prefecture, making up its own region. The Tsugaru Strait separates Hokkaidō from Honshu; the two islands are connected by the undersea railway Seikan Tunnel. The ...
. The skull is very well preserved but is missing most of the dermal roof bones through weathering. It was donated to the defunct Historical Museum of Hokkaido, which has since been integrated into and replaced by the
Hokkaido Museum opened in Sapporo, Hokkaidō, Japan in 2015. Located within Nopporo Shinrin Kōen Prefectural Natural Park, the permanent exhibition is dedicated to the nature, history, and culture of Hokkaido. Also known as , the museum integrates and replaces ...
.


References

{{Taxonbar, from=Q6821460 Late Cretaceous turtles of Asia Dermochelyidae Cretaceous animals of Asia Fossil taxa described in 1996 Prehistoric turtle genera Campanian genus first appearances Monotypic prehistoric reptile genera Fossils of Japan Maastrichtian genus extinctions