Markham's storm petrel (''Hydrobates markhami'') is a species of storm petrel in the family

Hydrobatidae Northern storm petrels are seabirds in the genus ''Hydrobates'' in the family Hydrobatidae, part of the order Procellariiformes. The family was once lumped with the similar austral storm petrels in the combined storm petrels, but have been split ...

. An all-black to sooty brown

seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...

, Markham's storm petrel is difficult to differentiate from the

black petrel The black petrel (''Procellaria parkinsoni''), also called the Parkinson's petrel, is a medium-sized, black-plumaged petrel, the smallest of the ''Procellaria''. The species is an endemic breeder of New Zealand, breeding only on Great Barrier Is ...

''Procellaria parkinsoni'' in life, and was once described as

conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organ ...

with, or biologically identical to,

Tristram's storm petrel Tristram's storm petrel or ''akihikeehiale'' (''Hydrobates tristrami'') is a species of seabird in the storm petrel family Hydrobatidae. The species' common and scientific name is derived from the English clergyman Henry Baker Tristram; the specie ...

''Hydrobates tristrami''. Markham's storm petrel inhabits open seas in the Pacific Ocean around Peru, Chile, and Ecuador, but only nests in northern Chile and Peru, with ninety-five percent of all known breeding populations in 2019 found in the

Atacama Desert The Atacama Desert ( es, Desierto de Atacama) is a desert plateau in South America covering a 1,600 km (990 mi) strip of land on the Pacific coast, west of the Andes Mountains. The Atacama Desert is the driest nonpolar desert in the ...

. First described by British ornithologist

Osbert Salvin Osbert Salvin FRS (25 February 1835 – 1 June 1898) was an English naturalist, ornithologist, and herpetologist best known for co-authoring ''Biologia Centrali-Americana'' (1879–1915) with Frederick DuCane Godman. This was a 52 volume encyc ...

in 1883, the bird was named in honor of

Albert Hastings Markham Admiral Sir Albert Hastings Markham (11 November 1841 – 28 October 1918) was a British explorer, author, and officer in the Royal Navy. In 1903 he was invested as a Knight Commander of the Order of the Bath. He is also remembered for desig ...

, a naval officer who collected a specimen off Peru. Markham's storm petrel nests in natural cavities in

saltpeter Potassium nitrate is a chemical compound with the chemical formula . This alkali metal nitrate salt is also known as Indian saltpetre (large deposits of which were historically mined in India). It is an ionic salt of potassium ions K+ and nitrat ...

, and pairs produce one egg per season. After hatching, fledglings make their way to sea, and can be either attracted to or disoriented by artificial lights. The diet of Markham's storm petrel consists of fish,

cephalopod A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head ...

s, and

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

s, with about ten percent of stomach contents traceable to scavenging according to a 2002 study. Since at least 2012, Markham's storm petrel has been listed as an

endangered species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...

in Chile, and, in 2019, the

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

(IUCN) classified the

conservation status The conservation status of a group of organisms (for instance, a species) indicates whether the group still exists and how likely the group is to become extinct in the near future. Many factors are taken into account when assessing conservation ...

of Markham's storm petrel as

Near Threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify fo ...

due to

habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...

on its nesting grounds. The (SAG), a department of the

Ministry of Agriculture An agriculture ministry (also called an) agriculture department, agriculture board, agriculture council, or agriculture agency, or ministry of rural development) is a ministry charged with agriculture. The ministry is often headed by a minister f ...

of Chile, has undertaken conservation efforts in the country.

Taxonomy

The

northern storm petrel Northern storm petrels are seabirds in the genus ''Hydrobates'' in the family Hydrobatidae, part of the order Procellariiformes. The family was once lumped with the similar austral storm petrels in the combined storm petrels, but have been split ...

family, Hydrobatidae, is a group of

s characterized by long legs and a high adaption to marine environments, with its species predominately

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to the northern hemisphere. In Hydrobatidae, Markham's storm petrel is a member of the genus '' Hydrobates'', the only genus in the family, and is large compared to other members in the genus. Hydrobatidae probably diverged from other

petrel Petrels are tube-nosed seabirds in the bird order Procellariiformes. Description The common name does not indicate relationship beyond that point, as "petrels" occur in three of the four families within that group (all except the albatross f ...

s at an early stage; according to John Warham, the petrel group had "substantial radiation" by the

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

. Storm petrel fossils are rare; those which are found date from the

Upper Miocene The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene Epoch made up of two stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million years ago) to 5.333 Ma. The ev ...

in California.

Wilson's storm petrel Wilson's storm petrel (''Oceanites oceanicus''), also known as Wilson's petrel, is a small seabird of the austral storm petrel family Oceanitidae. It is one of the most abundant bird species in the world and has a circumpolar distribution mainly ...

, ''Oceanites oceanicus'', in the physically similar but only distantly related

Oceanitidae Austral storm petrels, or southern storm petrels, are seabirds in the family Oceanitidae, part of the order Procellariiformes. These smallest of seabirds feed on planktonic crustaceans and small fish picked from the surface, typically while hove ...

, may have been the first storm petrel to inhabit the Northern Hemisphere, thus possibly starting the subfamily Hydrobatinae of which Markham's storm petrel is a member. British ornithologist

first described Markham's storm petrel as ''Cymochorea markhami'' in 1883. Markham's storm petrel is named after Sir

, a British explorer and naval officer who picked up the

type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...

off

Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...

. The bird was thought by ornithologist

James L. Peters James Lee Peters (August 13, 1889 – April 19, 1952) was an American ornithologist. He was born in Boston, Massachusetts to Dr. Austin Peters and Francis Howie Lee on August 13, 1889. His early education was at the Roxbury Latin School, followe ...

in 1931 as

, or biologically identical, with

''Oceanodroma tristrami'', though the two species were later distinguished by size. Similarly, ornithologist Reginald Wagstaffe considered Tristram's storm petrel a

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

of Markham's storm petrel in 1972, though subsequent research recognized them as different species. In 2016, the

(IUCN) reclassified ''Oceanodroma markhami'' as ''Hydrobates markhami'' based on ''

HBW and BirdLife International Illustrated Checklist of the Birds of the World The ''HBW and BirdLife International Illustrated Checklist of the Birds of the World'' is a checklist of the birds of the world published by Lynx Edicions Lynx Edicions is a Spanish publishing company specializing in ornithology and natural histo ...

'', volume 1, by Josep del Hoyo and British ornithologist Nigel J. Collar. In some studies, the genus ''Oceanodroma'' was found to be

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

with respect to ''Hydrobates'', and all former ''Oceanodroma'' species were transferred to ''Hydrobates'' under some authorities. In 2021, all species in ''Oceanodroma'' were transferred to ''Hydrobates'' by the

International Ornithologists' Union The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...

. Its name in Spanish is ''Golondrina de mar negra'' or ''paíño ahumado''.

Description

Markham's storm petrel is an all-black to sooty brown storm petrel with a dull lead-gray gloss on its head, neck and mantle in fresh plumage. Its underside, from the neck down, and wing lining are blackish brown, and become almost

fuscous {{Short pages monitor