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''Lophorhothon'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
hadrosauroid Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or hadrosaurids, and all dinosaurs more closely related to them than to ''Iguanodon''. Their remains have been recovered in Asia, Eu ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
, the first genus of dinosaur discovered in
Alabama (We dare defend our rights) , anthem = "Alabama (state song), Alabama" , image_map = Alabama in United States.svg , seat = Montgomery, Alabama, Montgomery , LargestCity = Huntsville, Alabama, Huntsville , LargestCounty = Baldwin County, Al ...
, in the United States.


Discovery and naming

Remains of this small, poorly known perhaps
saurolophine Saurolophinae is a subfamily (biology), subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. How ...
dinosaur were first discovered during the 1940s, from extensive erosional outcrops of the lower unnamed member of the
Mooreville Chalk Formation The Mooreville Chalk is a geological formation in North America, within the U.S. states of Alabama and Mississippi, which were part of the subcontinent of Appalachia. The strata date back to the early Santonian to the early Campanian stage of t ...
(
Selma Group The Selma Group is a geological formation in North America, within the U.S. states of Alabama, Mississippi, and Tennessee. The strata date from the Santonian to the Maastrichtian stages of the Late Cretaceous. The group is composed of, in ascend ...
; lower and middle
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
) in
Dallas County Dallas County may refer to: Places in the USA: * Dallas County, Alabama, founded in 1818, the first county in the United States by that name * Dallas County, Arkansas * Dallas County, Iowa * Dallas County, Missouri * Dallas County, Texas, the nin ...
, west of the town of
Selma, Alabama Selma is a city in and the county seat of Dallas County, in the Black Belt region of south central Alabama and extending to the west. Located on the banks of the Alabama River, the city has a population of 17,971 as of the 2020 census. About ...
. The taxon has since also been reported from
Black Creek Formation The Black Creek Group is a geologic group in North Carolina. It preserves fossils dating back to the Late Cretaceous period. Paleofauna * cf. '' Deinosuchus rugosus'' * cf. '' Coelosaurus antiquus'' * cf. '' Dryptosaurus sp.'' * cf. '' Lophorh ...
(Campanian) of
North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia and So ...
. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, which is housed in the collections of the Field Museum in Chicago, consists of a fragmentary and disarticulated skull and incomplete postcranial skeleton. The length on the holotype specimen has been estimated as . The genus was named by
Wann Langston Wann Langston Jr. (1921 – April 7, 2013) was an American paleontologist and professor at the University of Texas at Austin. Langston worked on a number of different reptiles and amphibians in his long career, beginning with the 1950 description ...
in 1960. It was thought to be the only species of hadrosaur from that fossil formation, until 2016 with the discovery of the primitive hadrosaur, ''
Eotrachodon orientalis ''Eotrachodon orientalis'' (meaning "dawn ''Trachodon'' from the east") is a species of hadrosaurid that was described in 2016. The holotype was found in the Mooreville Chalk Formation (Upper Santonian) in Alabama in 2007 and includes a well-pr ...
''. The name ''Lophorhothon'' means "crested nose" (
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
''lophos'' meaning 'crested' and ''rhothon'' meaning 'nose'). The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Lophorhothon atopus''. The specific name is derived from Greek ''atopos'', "uncommon" or "strange". The specimen which Langston designated as the holotype was discovered by Rainier Zangerl, Bill Turnbull and Charles Barber on a Field Museum expedition in 1946 and was given catalogue number FMNH P 27383. It consists of less than one half of the skull, a number of
vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic i ...
, and significant portions of the fore- and hindlimbs. Preserved cranial material includes a partial quadrate, left maxilla, teeth, jugal, lacrimal, nasal (with the namesake crest), postorbital, frontal, prefrontal, parietal, squamosal, and paroccipital process and a portion of the predentary bone. The specimen was likely washed out to sea by a river, where it eventually sank and was buried in the silty
carbonate A carbonate is a salt of carbonic acid (H2CO3), characterized by the presence of the carbonate ion, a polyatomic ion with the formula . The word ''carbonate'' may also refer to a carbonate ester, an organic compound containing the carbonate g ...
sediments of the
Mississippi embayment The Mississippi embayment is a physiographic feature in the south-central United States, part of the Mississippi Alluvial Plain. It is essentially a northward continuation of the fluvial sediments of the Mississippi River Delta to its conflu ...
. In 2021, a more complete specimen was unearthed in Alabama.


Classification

Since the publication of Langston's description of ''Lophorhothon'' a number of workers have questioned the validity of this
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
. It has been suggested, for example, that the material may actually represent a juvenile ''
Prosaurolophus ''Prosaurolophus'' (; meaning "before ''Saurolophus''", in comparison to the later dinosaur with a similar head crest) is a genus of hadrosaurid (or duck-billed) dinosaur from the Late Cretaceous of North America. It is known from the remains of ...
''. James Lamb in 1998 has suggested the genus may actually represent a basal
iguanodont Iguanodontia (the iguanodonts) is a clade of herbivorous dinosaurs that lived from the Middle Jurassic to Late Cretaceous. Some members include ''Camptosaurus'', ''Dryosaurus'', ''Iguanodon'', '' Tenontosaurus'', and the hadrosaurids or "duck-bil ...
, an idea that has failed to find widespread acceptance. More recent workers (Horner, Weishampel, and Forster, 2004) have classified ''Lophorhothon'' as a basal hadrosaurine and a sister taxon to all other hadrosaurines. An analysis published in 2010 indicated it was a basal member of the
Hadrosauroidea Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or hadrosaurids, and all dinosaurs more closely related to them than to ''Iguanodon''. Their remains have been recovered in Asia, Europ ...
. In 2021, new material from Alabama was unearthed not too far from where the original holotype was unearthed. This find cemented its status as a valid genus of
hadrosauromorph Hadrosauromorpha is a clade of iguanodontian ornithopods, defined in 2014 by David B. Norman to divide Hadrosauroidea into the basal taxa with compressed manual bones and a pollex, and the derived taxa that lack them. The clade is defined as a ...
.


See also

*
Timeline of hadrosaur research A timeline is a display of a list of events in chronological order. It is typically a graphic design showing a long bar labelled with dates paralleling it, and usually contemporaneous events. Timelines can use any suitable scale representin ...


References


Sources

*Horner, J. R., Weishampel, D. B., and Forster, C. A. 2004. Chapter Twenty: Hadrosauridae. in The Dinosauria (2nd edition), Weishampel, D. B., Dodson, P., and Osmólska, H., editors. University of California Press. *Lamb, J. P. 1998. ''Lophorothon'', an iguanodontian, not a hadrosaur. Journal of Vertebrate Paleontology, 18 (3 Abstracts): 59A. *Langston, W. 1960. The vertebrate fauna of the Selma Formation of Alabama, part VI: the dinosaurs. Fieldiana: Geology Memoirs 3(5): 315–359. *Schwimmer, D. R. 1997. Late Cretaceous dinosaurs in eastern USA: a taphonomic and biogeographic model of occurrences, p. 203–211. In D. L. Wolberg, E. Stump, and G. D. Rosenberg (eds.), Dinofest International. The Academy of Natural Sciences, Philadelphia. *Thurmond, J. T. and Jones, D. E. 1981. Fossil vertebrates of Alabama. University of Alabama Press. {{Taxonbar, from=Q643185 Late Cretaceous dinosaurs of North America Hadrosaurs Fossil taxa described in 1960 Mooreville Chalk Campanian genus first appearances Campanian genus extinctions Ornithischian genera