List Of Butterflies Of The Amazon River Basin And The Andes
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This is a list of butterflies of the Amazon River basin and the Andes. The
Amazon River basin The Amazon basin is the part of South America drained by the Amazon River and its tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries of Bolivi ...
may be the most speciose region for
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
. Nine countries have territory in the Amazon River basin or immediately adjoin this region:
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
,
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
,
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
,
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
,
Guyana Guyana ( or ), officially the Cooperative Republic of Guyana, is a country on the northern mainland of South America. Guyana is an indigenous word which means "Land of Many Waters". The capital city is Georgetown. Guyana is bordered by the ...
,
Suriname Suriname (; srn, Sranankondre or ), officially the Republic of Suriname ( nl, Republiek Suriname , srn, Ripolik fu Sranan), is a country on the northeastern Atlantic coast of South America. It is bordered by the Atlantic Ocean to the north ...
, and
French Guiana French Guiana ( or ; french: link=no, Guyane ; gcr, label=French Guianese Creole, Lagwiyann ) is an overseas departments and regions of France, overseas department/region and single territorial collectivity of France on the northern Atlantic ...
. The
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
extend from north to south through seven South American countries: Venezuela, Colombia, Ecuador, Peru, Bolivia,
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
, and
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. The
fauna of the Andes The fauna of the Andes, a mountain range in South America, is large and diverse. As well as a huge variety of flora, the Andes contain many different animal species. With almost 1,000 species, of which roughly 2/3 are endemic to the region, the ...
is also diverse. Both regions have many
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
species. South America as a whole constitutes the
Neotropical realm The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
. Habitats in these two regions are very various and include
Amazon rainforest The Amazon rainforest, Amazon jungle or ; es, Selva amazónica, , or usually ; french: Forêt amazonienne; nl, Amazoneregenwoud. In English, the names are sometimes capitalized further, as Amazon Rainforest, Amazon Forest, or Amazon Jungle. ...
,
Atlantic forest The Atlantic Forest ( pt, Mata Atlântica) is a South American forest that extends along the Atlantic coast of Brazil from Rio Grande do Norte state in the northeast to Rio Grande do Sul state in the south and inland as far as Paraguay and th ...
, Llanos grasslands,
Puna grassland The puna grassland ecoregion, of the montane grasslands and shrublands biome, is found in the central Andes Mountains of South America. It is considered one of the eight Natural Regions in Peru,Pulgar Vidal, Javier: Geografía del Perú; Las Ocho ...
and
Valdivian temperate forests The Valdivian temperate forests (NT0404) is an ecoregion on the west coast of southern South America, in Chile and Argentina. It is part of the Neotropical realm. The forests are named after the city of Valdivia. The Valdivian temperate rainforest ...
. Peru east of the Andes is regarded as the most important
biodiversity hotspot A biodiversity hotspot is a biogeographic region with significant levels of biodiversity that is threatened by human habitation. Norman Myers wrote about the concept in two articles in ''The Environmentalist'' in 1988 and 1990, after which the co ...
in the world. The two regions (Amazon and Andes) are South America proper excluding the
pampas The Pampas (from the qu, pampa, meaning "plain") are fertile South American low grasslands that cover more than and include the Argentine provinces of Buenos Aires, La Pampa, Santa Fe, Entre Ríos, and Córdoba; all of Uruguay; and Brazil ...
plains of
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
and
Paraguay Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...
which have a distinct butterfly fauna. Isolation has led to the evolution of endemic higher taxa. Instances are
Ithomiinae Ithomiini is a butterfly tribe in the nymphalid subfamily Danainae. It is sometimes referred to as the tribe of clearwing butterflies or glasswing butterflies. Some authors consider the group to be a subfamily (Ithomiinae). These butterflies are ...
,
Dismorphiinae Dismorphiinae, the mimic sulphurs, is a subfamily of butterflies from the family Pieridae. It consists of about 100 species in seven genera, distributed mainly in the Neotropical region, of which only one species occurs in North America and one ge ...
, Phyciodina, Pyrrhopygini,
Eumaeini The Eumaeini are a tribe of gossamer-winged butterflies (family Lycaenidae). They are typically placed in the subfamily Theclinae, but sometimes considered a separate subfamily Eumaeinae. Over 1,000 species are found in the Neotropical realm Sys ...
(over 1,000 species),
Pronophilina Pronophilina is a Neotropical subtribe of butterflies of the subfamily Satyrinae. They are a species-rich group with highest diversity in the tropical and subtropical mountains, especially the Andes. Before 1970, they were poorly studied, but re ...
and
Eudaminae The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synonym of ''Urbanus''. They are largely found in the Neotropics, with some extending into temperate North Ameri ...
. Endemic genera (among very many) include ''
Morpho The morpho butterflies comprise many species of Neotropical butterfly under the genus ''Morpho''. This genus includes more than 29 accepted species and 147 accepted subspecies, found mostly in South America, Mexico, and Central America. ''Morph ...
'', ''
Agrias ''Agrias'' is a genus of Neotropical charaxine nymphalid butterflies found in South and Central America. The German lepidopterist Hans Fruhstorfer wrote: "In this magnificent tropical genus, upon which nature seems to have showered all her abu ...
'', ''
Prepona ''Prepona'' is a genus of Neotropical charaxine butterflies in the family Nymphalidae. They are strong fliers in tropical forests where they feed on fermenting fruits and animal dung. The underside of the wings is pale greyish or brownish, while ...
'' '' Caligo'', ''
Cithaerias ''Cithaerias'' is a Neotropical butterfly genus from the subfamily Satyrinae in the family Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to ...
'', '' Catagramma'', ''
Parides ''Parides'', commonly called cattlehearts, is a genus of swallowtail butterflies in the family Papilionidae. They are found in the Americas (Neotropical realm). Species Listed alphabetically within groups according to Möhn ''et al.'', with anno ...
'', '' Hamadryas'', ''
Nessaea ''Nessaea'' is a genus of nymphalid butterflies found in the Neotropical realm. Unlike virtually all other butterflies with blue coloration, the blue colors in this genus are due to pigmentation terobilin (biliverdin IXγ)ref>Doug Taron"It's all ...
'', ''
Marpesia In ancient Greek and Roman legendary history, Marpesia (Greek: Μαρπησία "Snatcher"; sometimes wrongly spelled Marthesia) was Queen of the Amazons with Lampedo ("burning torch"), her sister, as a co-ruler. They ruled with Hippo ("horse") ...
'', '' Melanis'' '' Mesosemia'', '' Symmachia'', '' Evenus'', ''
Memphis Memphis most commonly refers to: * Memphis, Egypt, a former capital of ancient Egypt * Memphis, Tennessee, a major American city Memphis may also refer to: Places United States * Memphis, Alabama * Memphis, Florida * Memphis, Indiana * Memp ...
'', ''
Pierella ''Pierella'' is a butterfly genus from the subfamily Satyrinae in the family Nymphalidae found from Mexico through Central America to South America. The species of ''Pierella'' have larger hindwings than forewings, unique among butterflies. The o ...
'', and '' Astraptes''. Other higher taxa are most speciose in the Neotropics, for instance
Riodininae __NOTOC__ Riodininae is the largest of the three subfamilies within the metalmark butterfly family, Riodinidae. Classification Riodininae contains the following tribes: * Befrostiini Grishin, 2019 * Calydnini Seraphim, Freitas & Kaminski, 201 ...
. Many species, notably ''
Heliconius ''Heliconius'' comprises a colorful and widespread genus of brush-footed butterflies commonly known as the longwings or heliconians. This genus is distributed throughout the tropical and subtropical regions of the New World, from South America a ...
'', are members of complex mimicry rings.
Adaptive radiation In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic int ...
has led to many species being geographically diverse. Examples are ''
Consul fabius ''Consul fabius'', the tiger leafwing, is the most common and well known species of the genus ''Consul'' of subfamily Charaxinae in the brush-footed butterfly family (Nymphalidae). It is found all over the Neotropics. Description The length ...
'' and '' Mechanitis lysimnia''. Notable entomologists associated with Neotropical butterflies are
Jean-Baptiste Godart Jean-Baptiste Godart (25 November 1775 – 27 July 1825) was a French entomologist. Born at Origny, Godart became impassioned by butterflies in his youth. He was charged by Pierre André Latreille (1762-1833) with writing the article on th ...
,
Henry Walter Bates Henry Walter Bates (8 February 1825, in Leicester – 16 February 1892, in London) was an English naturalist and explorer who gave the first scientific account of mimicry in animals. He was most famous for his expedition to the rainforests of ...
,
William Chapman Hewitson William Chapman Hewitson (9 January 1806, in Newcastle upon Tyne – 28 May 1878, in Oatlands Park, Surrey) was a British naturalist. A wealthy collector, Hewitson was particularly devoted to Coleoptera (beetles) and Lepidoptera (butterflies ...
,
Hans Fruhstorfer Hans Fruhstorfer (7 March 1866, in Passau, Germany – 9 April 1922, in Munich) was a German explorer, insect trader and entomologist who specialised in Lepidoptera. He collected and described new species of exotic butterflies, especially in Ada ...
,
Otto Staudinger Otto Staudinger (2 May 1830 – 13 October 1900) was a German entomologist and a natural history dealer considered one of the largest in the world specialising in the collection and sale of insects to museums, scientific institutions, and indi ...
, Karl Jordan and
Walter Rothschild Lionel Walter Rothschild, 2nd Baron Rothschild, Baron de Rothschild, (8 February 1868 – 27 August 1937) was a British banker, politician, zoologist and soldier, who was a member of the Rothschild family. As a Zionist leader, he was present ...
,
Anton Hermann Fassl Anton Heinrich Hermann Fassl (1876, Komotau - 1922, Manaos) was a German entomologist. Fassl collected Lepidoptera and Coleoptera in Colombia (1907-1908), Brazil and Ecuador. He was sometime in Berlin, sometime at a dealership Naturhistorisches-I ...
,
Hermann Burmeister Karl Hermann Konrad Burmeister (also known as Carlos Germán Conrado Burmeister) (15 January 1807 – 2 May 1892) was a German Argentine zoologist, entomologist, herpetologist, botany, botanist, and coleopterologist. He served as a professor at ...
,
William Schaus William Schaus (January 11, 1858 in New York City – June 20, 1942) was an American Entomology, entomologist who became known for his major contribution to the knowledge and description of new species of the Neotropical realm, Neotropical Lepidop ...
,
Eugène Le Moult Eugène Le Moult (31 December 1882, Quimper – 26 January 1967, Paris) was a French naturalist and entomologist specialised in butterflies; hunter, businessman and collector. Le Moult grew up in the tropical prison colony of French Guiana, whe ...
,
Richard Haensch Richard Haensch (active from 1890) was a German entomologist and insect dealer in Berlin. Haensch collected in Bahia (1893–1894), Minas Gerais (1896–1897) and Ecuador (1899–1900). He wrote the section "''Familie Danaidae''" in ''Die Gross-S ...
,
Gustav Weymer Gustav Weymer (1833–1914) was a German entomologist. He described many new taxa of butterflies from specimens collected by Alphons Stübel in South America. His own collections and those he worked on are conserved in Museum für Naturkund ...
,
Ferdinand Heinrich Hermann Strecker Ferdinand Heinrich Herman Strecker (24 March 1836, in Philadelphia – 30 November 1901, in Reading, Pennsylvania) was an American entomologist specialising in butterflies and moths (Lepidoptera). Strecker was born in Philadelphia to Ferdinand and ...
,
Andrey Avinoff Andrey Avinoff (14 February 1884 – 16 July 1949); was an internationally-known artist, lepidopterist, museum director, professor, bibliophile and iconographer, who served as the director of the Carnegie Museum of Natural History in Pittsburgh ...
,
Carlos Berg Carlos Berg ( lv, Kārlis Bergs, german: Karl Berg) or Friedrich Wilhelm Karl Berg ( lv, Frīdrihs Vilhelms Kārlis Bergs, german: Friedrich Wilhelm Karl Berg) (21 March 1843, Courland – 19 January 1902 Buenos Aires) was an Argentinian natu ...
, and
Vladimir Nabokov Vladimir Vladimirovich Nabokov (russian: link=no, Владимир Владимирович Набоков ; 2 July 1977), also known by the pen name Vladimir Sirin (), was a Russian-American novelist, poet, translator, and entomologist. Bo ...
.


Papilionidae Swallowtail butterflies are large, colorful butterflies in the family Papilionidae, and include over 550 species. Though the majority are tropical, members of the family inhabit every continent except Antarctica. The family includes the larges ...


Papilioninae Papilioninae is a subfamily of the butterfly family Papilionidae. Papilioninae are found worldwide, but most species are distributed in the tropics. There are roughly 480 species, of which 27 occur in North America. Tribes This subfamily consis ...


Riodinidae Riodinidae is the family of metalmark butterflies. The common name "metalmarks" refers to the small, metallic-looking spots commonly found on their wings. The 1532 species are placed in 146 genera. Although mostly Neotropical in distribution, t ...


Pieridae The Pieridae are a large family of butterflies with about 76 genera containing about 1,100 species, mostly from tropical Africa and tropical Asia with some varieties in the more northern regions of North America and Eurasia.DeVries P. J. in Levi ...


Coliadinae Coliadinae, the sulphurs or yellows, are a subfamily of butterflies with about 300 described species. There are 36 species in North America, where they range from Mexico to northern Canada. In most species, males are easily distinguished from f ...


Dismorphiinae Dismorphiinae, the mimic sulphurs, is a subfamily of butterflies from the family Pieridae. It consists of about 100 species in seven genera, distributed mainly in the Neotropical region, of which only one species occurs in North America and one ge ...


Pierinae The Pierinae are a large subfamily of pierid butterflies. The subfamily is one of several clades of butterflies often referred to as the whites. Species It includes the following species (additional species can be found under the tribes list ...


Lycaenidae Lycaenidae is the second-largest family of butterflies (behind Nymphalidae, brush-footed butterflies), with over 6,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of the known butterfl ...


Polyommatinae Polyommatinae, the blues, are a diverse subfamily of gossamer-winged butterflies (family Lycaenidae). This subfamily was long used to assign taxa of unclear relationships, and its contents and phylogeny are still in need of revision. The followi ...


Theclinae The subfamily Theclinae is a group of butterflies, often referred to as hairstreaks, with some species instead known as elfins or by other names. The group is part of the family Lycaenidae, the "gossamer-winged butterflies". There are many tropi ...


Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species have a red ...


Apaturinae The Apaturinae are a subfamily of butterflies that includes many species commonly called emperors. Description Strikingly-coloured, with cryptic underwing. A distinguishing character of the subfamily is the green proboscis. Agathina emperor ( ...


Biblidinae Biblidinae is a subfamily of nymphalid butterflies that includes the tropical brushfoots. This subfamily was sometimes merged within the Limenitidinae, but they are now recognized as quite distinct lineages. In older literature, this subfami ...


Charaxinae __NOTOC__ The Charaxinae, the leafwings, are a nymphalid subfamily of butterflies that includes about 400 species, inhabiting mainly the tropics, although some species extend into temperate regions in North America, Europe, China, and southern A ...


Cyrestinae Cyrestinae is the name of a small subfamily of nymphalid brush-footed butterflies. It is considered to include only three genera – ''Marpesia'', ''Chersonesia'', and ''Cyrestis'' – distributed in the tropics.N. Wahlberg , J. Leneveu , U. ...


Danainae Danainae is a subfamily of the family Nymphalidae, the brush-footed butterflies. It includes the Daniadae, or milkweed butterflies, who lay their eggs on various milkweeds on which their larvae (caterpillars) feed, as well as the clearwing butt ...


Heliconiinae The Heliconiinae, commonly called heliconians or longwings, are a subfamily of the brush-footed butterflies (family Nymphalidae). They can be divided into 45–50 genera and were sometimes treated as a separate family Heliconiidae within the P ...


Libytheinae The Libytheinae are a nymphalid subfamily known as snout butterflies, containing two valid genera and about ten species: six in '' Libythea'' and four in ''Libytheana''. The common name refers to the thick labial palps (pedipalps) that look like ...


Limenitidinae The Limenitidinae are a subfamily of butterflies that includes the admirals and relatives. The common names of many species and genera reference military ranks or – namely the Adoliadini – titles of nobility (e.g., count, duke, earl, an ...


Morphinae The Morphinae are a subfamily of Nymphalidae butterflies that includes the morphos, the owl butterflies (''Caligo''), and related lineages. It is either considered a sister group of the Satyrinae, or disassembled and included therein. System ...


Nymphalinae The Nymphalinae are a subfamily of brush-footed butterflies (family Nymphalidae). Sometimes, the subfamilies Limenitidinae, and Biblidinae are included here as subordinate tribe(s), while the tribe Melitaeini is occasionally regarded as a distinc ...


Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly considered a distinct family, Satyridae. This group contains nearly half of the known divers ...


Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy ...


Hesperiinae Grass skippers or banded skippers are butterflies of the subfamily Hesperiinae, part of the skipper family, Hesperiidae. The subfamily was established by Pierre André Latreille in 1809. Description and distribution With over 2,000 described sp ...


Pyrginae


Eudaminae The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synonym of ''Urbanus''. They are largely found in the Neotropics, with some extending into temperate North Ameri ...


Pyrrhopyginae Firetips or firetail skippers are skipper butterflies in the subfamily Pyrrhopyginae (family (biology), family Hesperiidae). The roughly 150 species are found only in the Neotropics, with the exception of one species which just reaches into the U ...


References

*
Bernard d'Abrera Bernard d'Abrera (28 August 1940 – 13 January 2017) was an Australian entomological taxonomist and philosopher of science, particularly noted for his books on true butterflies (Papilionoidea) and larger moths of the world (Saturniidae and Sphing ...
, 1981-1995 ''Butterflies of the Neotropical Region'' Hill House/Lansdowne Melbourne in association with E.W. Classey Part1 1981 Papilionidae. Pieridae ; part 2 1984 Danaidae, Ithomiidae, Heliconidae & Morphidae ; 19- part 3 Brassolidae, Acraeidae, Nymphalidae (partim) plus corrections to Parts I and II ; part 4 1987 Nymphalidae (partim) including Callicore, Asterope, Agrias and Prepona ; part 5 (Nymphalidae (concl.), Anaea (Memphis) Satyridae (complete), Libytheidae)ISBN ? ; part 6 1994 Riodinidae ; part 7 1995 Lycaenidae * Andrade-C., M. Gonzalo. Biodiversidad de las Mariposas (Lepidoptera: Rhopalocera) de Colombia. pp. 153–172. In: ''Proyecto de Red Iberoamericana de Biogeografía y Entomología Sistemática PrIBES 2002''. Costa. C., Vanin, S. A., Lobo, J. M. & Melic, A. (Eds.) Monografías Tercer Milenio, vol. 2, SEA, Zaragoza, Julio-2002. * Bollino, M., & Onore, G. (2001). ''Mariposas del Ecuador / Butterflies of Ecuador: Vol. 10a. Familia: Papilionidae''. Quito: Pontificia Universidad Católica del Ecuador. * Garwood, K. M., Lehman, Carter, W., & Carter, G. (2007). ''Butterflies of Southern Amazonia''. Mission, Texas: Neotropical Butterflies. * Lacomme, D., & Manil, L. (Eds.). (2002). ''Lépidoptères de Guyane''. Paris: Association des Lépidopteristes Parisiens. * Lamas, G. (Ed.). (2004). ''Atlas of Neotropical Lepidoptera. Checklist: Part 4A. Hesperioidea-Papilionoidea''. Gainesville: Association for Tropical Lepidoptera. 094541728 (v. 5A) * Lamas, G. ''Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea)'' Revised Electronic Edition (2007) *Mittermeier, R. A., P. R. Gil, and C. G. Mittermeier 1999. ''Megadiversity: Earth's Biologically Wealthiest Nations''. Conservation International, USA. 501 pp. * Neild, A. F. E. (1997). ''The butterflies of Venezuela. Pt. 1. Nymphalidae I (Limenitidinae, Apaturinae, Charaxinae)''. Greenwich, UK: Meridian Publ. *
Adalbert Seitz Friedrich Joseph Adalbert Seitz, (24 February 1860 in Mainz – 5 March 1938 in Darmstadt) was a German physician and entomologist who specialised in Lepidoptera. He was a director of the Frankfurt zoo from 1893 to 1908 and is best known for editi ...
''Die Großschmetterlinge der Erde'', Verlag Alfred Kernen, Stuttgart Band 5 Die exotischen Großschmetterlinge, Die Großschmetterlinge des amerikanischen Faunengebietes, 1907-192
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Biodiversity Heritage Library The Biodiversity Heritage Library (BHL) is the world’s largest open access digital library for biodiversity literature and archives. BHL operates as worldwide consortiumof natural history, botanical, research, and national libraries working toge ...
* Tyler, H. A., Brown, K. S., Jr., & Wilson, K. (1994). ''Swallowtail Butterflies of the Americas: A Study in Biological Dynamics, Ecological Diversity, Biosystematics, and Conservation''. Gainesville: Scientific Publishers.


External links


Butterflies of the Americas Willmott, K.R., & Hall, J.P.W. (2008). ''Butterflies of Ecuador''.
{{South America in topic, List of butterflies of ' ' ' °Amazon basin and Andes Lepidoptera of Brazil Lepidoptera of Colombia Lepidoptera of Venezuela