Kaatedocus
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''Kaatedocus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
diplodocine Diplodocinae is an extinct subfamily of diplodocid sauropods that existed from the Late Jurassic to Early Cretaceous of North America, Europe, Africa and South America, about 161.2 to 136.4 million years ago. Genera within the subfamily include ' ...
flagellicaudata Flagellicaudata is a clade of Dinosauria. It belongs to Sauropoda and includes two families, the Dicraeosauridae and the Diplodocidae. Phylogeny The clade Flagellicaudata was erected by Harris and Dodson (2004) for the diplodocoid clade formed ...
n sauropod known from the middle
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
(
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxford ...
stage) of northern
Wyoming Wyoming () is a state in the Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to the s ...
, United States. It is known from well-preserved
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
and cervical vertebrae which were collected in the lower part of the Morrison Formation. The type and only species is ''Kaatedocus siberi'', described in
2012 File:2012 Events Collage V3.png, From left, clockwise: The passenger cruise ship Costa Concordia lies capsized after the Costa Concordia disaster; Damage to Casino Pier in Seaside Heights, New Jersey as a result of Hurricane Sandy; People gat ...
by Emanuel Tschopp and
Octávio Mateus Octávio Mateus (born 1975) is a Portuguese dinosaur paleontologist and biologist Professor of Paleontology at the Faculdade de Ciências e Tecnologia da Universidade Nova de Lisboa. He graduated in Universidade de Évora and received his PhD at U ...
.


History

In 1934, a team of the American Museum of Natural History (AMNH) headed by
Barnum Brown Barnum Brown (February 12, 1873 – February 5, 1963), commonly referred to as Mr. Bones, was an American paleontologist. Named after the circus showman P. T. Barnum, he discovered the first documented remains of ''Tyrannosaurus'' during a career ...
, financed by the Sinclair Oil Corporation, uncovered about three thousand sauropod bones on the land of rancher Barker Howe near
Shell Shell may refer to: Architecture and design * Shell (structure), a thin structure ** Concrete shell, a thin shell of concrete, usually with no interior columns or exterior buttresses ** Thin-shell structure Science Biology * Seashell, a hard o ...
, in Big Horn County. Plans for further excavations in 1935 had to be cancelled after Howe, convinced by the large publicity surrounding the find that the remains were very valuable, demanded higher payment. The bones would not be described and most of them were lost in a fire at the AMNH during the 1940s; other were thrown away in the 1960s after having rotted because of being stowed in a chicken run at Shell. Only about 10% of the fossils survived, among them a skull. They were generally interpreted as belonging to ''
Barosaurus ''Barosaurus'' ( ) was a giant, long-tailed, long-necked, plant-eating sauropod dinosaur closely related to the more familiar ''Diplodocus''. Remains have been found in the Morrison Formation from the Upper Jurassic Period of Utah and South Da ...
''. In 1989 the site was reopened by Hans-Jakob Siber, the founder of the Swiss Aathal Dinosaur Museum. His team immediately adjacent to the old How Quarry discovered another 450 bones that became part of the collection of the Swiss museum. The finds included an exceptionally complete neck. In Switzerland, this became the subject of several lines of scientific investigation. In 2005 Daniela Schwarz studied the pneumatisation of the vertebrae by
tomography Tomography is imaging by sections or sectioning that uses any kind of penetrating wave. The method is used in radiology, archaeology, biology, atmospheric science, geophysics, oceanography, plasma physics, materials science, astrophysics, ...
scanning them with
neutron The neutron is a subatomic particle, symbol or , which has a neutral (not positive or negative) charge, and a mass slightly greater than that of a proton. Protons and neutrons constitute the nuclei of atoms. Since protons and neutrons beh ...
s and
X-rays An X-ray, or, much less commonly, X-radiation, is a penetrating form of high-energy electromagnetic radiation. Most X-rays have a wavelength ranging from 10  picometers to 10 nanometers, corresponding to frequencies in the range 30&nbs ...
. In 2010 Andreas Christian used the well-preserved vertebrae to support his hypothesis that sauropod necks were held in a rather upright position, which was confirmed by Armin Schmitt studying the vestibular system of ''Kaatedocus''. In 2012 Tschopp used a scan to create a replica of the neck by means of a 3D-printer.Emanuel Tschopp & Gordon Dzemski, 2012, "3-Dimensional Reproduction Techniques to Preserve and Spread Paleontological Material – a Case Study with a Diplodocid Sauropod Neck", ''Journal of Paleontological Techniques'' 10 During the intense study of the fossils it became clear that they did not represent ''Barosaurus'' but a species new to science. In 2012 this was named ''Kaatedocus siberi'', by the Swiss palaeontologist Emanuel Tschopp, who as a boy had visited the excavations, and his Portuguese colleague Octávio Mateus. The generic name, which means "small beam", combines a reference to the related form ''
Diplodocus ''Diplodocus'' (, , or ) was a genus of diplodocid sauropod dinosaurs, whose fossils were first discovered in 1877 by S. W. Williston. The generic name, coined by Othniel Charles Marsh in 1878, is a neo-Latin term derived from Greek Î´Î¹Ï ...
'' with a
Crow Indian The Crow, whose autonym is Apsáalooke (), also spelled Absaroka, are Native Americans living primarily in southern Montana. Today, the Crow people have a federally recognized tribe, the Crow Tribe of Montana, with an Indian reservation locate ...
diminutive suffix ''~kaate''. The specific name honours Siber.


Classification

Tschopp, Mateus, and Benson (2015) found ''Kaatedocus'' to an advanced member of the Diplodocinae, closely related to ''
Barosaurus ''Barosaurus'' ( ) was a giant, long-tailed, long-necked, plant-eating sauropod dinosaur closely related to the more familiar ''Diplodocus''. Remains have been found in the Morrison Formation from the Upper Jurassic Period of Utah and South Da ...
'', as shown below. On the other hand, Whitlock and Wilson (2020) find ''Kaatedocus'' to be the most basal
dicraeosaurid Dicraeosauridae is a family of diplodocoid sauropods who are the sister group to Diplodocidae. Dicraeosaurids are a part of the Flagellicaudata, along with Diplodocidae. Dicraeosauridae includes genera such as ''Amargasaurus'', ''Suuwassea'', ' ...
, finding such a position more likely than being a diplodocid.


References

{{Taxonbar, from=Q1027296 Late Jurassic dinosaurs of North America Dinosaurs of the Morrison Formation Fossil taxa described in 2012 Diplodocids Taxa named by Octávio Mateus Paleontology in Wyoming