A stem is one of two main structural axes of a
vascular plant
Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They al ...
, the other being the
root
In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the sur ...
. It supports
leaves
A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
,
flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s and
fruit
In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering.
Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
s, transports water and dissolved substances between the roots and the shoots in the
xylem
Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word ''xylem'' is derived from ...
and
phloem
Phloem (, ) is the living biological tissue, tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as ''photosynthates'', in particular the sugar sucrose, to the rest of the plant. This tran ...
, stores nutrients, and produces new living tissue. The stem can also be called halm or haulm.
The stem is normally divided into nodes and internodes:
* The nodes hold one or more leaves, as well as
bud
In botany, a bud is an undeveloped or embryonic shoot and normally occurs in the axil of a leaf or at the tip of a stem. Once formed, a bud may remain for some time in a dormant condition, or it may form a shoot immediately. Buds may be spec ...
s which can grow into branches (with
leaves
A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
,
conifer cone
A conifer cone (in formal botany, botanical usage: strobilus, plural strobili) is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in con ...
s, or
flowers
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
).
Adventitious roots Important structures in plant development are buds, shoots, roots, leaves, and flowers; plants produce these tissues and structures throughout their life from meristems located at the tips of organs, or between mature tissues. Thus, a living plant ...
may also be produced from the nodes.
* The internodes distance one node from another.
The term "
shoot
In botany, a plant shoot consists of any plant stem together with its appendages, leaves and lateral buds, flowering stems, and flower buds. The new growth from seed germination that grows upward is a shoot where leaves will develop. In the spri ...
s" is often confused with "stems"; "shoots" generally refers to new fresh plant growth including both stems and other structures like leaves or flowers. In most plants stems are located above the soil surface but some plants have
underground stem Underground stems are modified plant parts that derive from stem tissue but exist under the soil surface. They function as storage tissues for food and nutrients, in propagation of new clones, and in perennation (survival from one growing season to ...
s.
Stems have four main functions which are:
* Support for and the elevation of leaves,
flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s, and
fruit
In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering.
Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
s. The stems keep the leaves in the light and provide a place for the plant to keep its flowers and fruits.
* Transport of fluids between the roots and the shoots in the xylem and phloem (see below).
* Storage of nutrients.
* Production of new living tissue. The normal lifespan of plant cells is one to three years. Stems have cells called
meristem
The meristem is a type of tissue found in plants. It consists of undifferentiated cells (meristematic cells) capable of cell division. Cells in the meristem can develop into all the other tissues and organs that occur in plants. These cells conti ...
s that annually generate new living tissue.
Stems have two pipe-like tissues called
xylem
Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word ''xylem'' is derived from ...
and
phloem
Phloem (, ) is the living biological tissue, tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as ''photosynthates'', in particular the sugar sucrose, to the rest of the plant. This tran ...
. The xylem tissue transports water by the action of
transpiration pull
Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word ''xylem'' is derived from ...
,
capillary action
Capillary action (sometimes called capillarity, capillary motion, capillary rise, capillary effect, or wicking) is the process of a liquid flowing in a narrow space without the assistance of, or even in opposition to, any external forces li ...
, and
root pressure
Root pressure is the transverse osmotic pressure within the cells of a root system that causes sap to rise through a plant stem to the leaves.
Root pressure occurs in the xylem of some vascular plants when the soil moisture level is high either ...
. The phloem tissue consists of sieve tubes and their companion cells. The two tissues are separated by
cambium
A cambium (plural cambia or cambiums), in plants, is a tissue layer that provides partially undifferentiated cells for plant growth. It is found in the area between xylem and phloem. A cambium can also be defined as a cellular plant tissue from w ...
, a tissue that divides to form xylem or phloem cells.
Specialized terms
Stems are often specialized for storage, asexual reproduction, protection, or
photosynthesis
Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...
, including the following:
![Senecio angulatus stems](https://upload.wikimedia.org/wikipedia/commons/2/25/Senecio_angulatus_stems.jpg)
*
Acaulescent
This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
: used to describe stems in plants that appear to be stemless. Actually these stems are just extremely short, the leaves appearing to rise directly out of the ground, e.g. some
''Viola'' species.
* Arborescent: tree with woody stems normally with a single trunk.
*
Axillary bud
The axillary bud (or lateral bud) is an embryonic or organogenic shoot located in the axil of a leaf. Each bud has the potential to form shoots, and may be specialized in producing either vegetative shoots (stems and branches) or reproductive sh ...
: a bud which grows at the point of attachment of an older leaf with the stem. It potentially gives rise to a shoot.
*
Branch
A branch, sometimes called a ramus in botany, is a woody structural member connected to the central trunk (botany), trunk of a tree (or sometimes a shrub). Large branches are known as boughs and small branches are known as twigs. The term '' ...
ed: aerial stems are described as being branched or unbranched.
*
Bud
In botany, a bud is an undeveloped or embryonic shoot and normally occurs in the axil of a leaf or at the tip of a stem. Once formed, a bud may remain for some time in a dormant condition, or it may form a shoot immediately. Buds may be spec ...
: an embryonic shoot with immature stem tip.
*
Bulb
In botany, a bulb is structurally a short stem with fleshy leaves or leaf basesBell, A.D. 1997. ''Plant form: an illustrated guide to flowering plant morphology''. Oxford University Press, Oxford, U.K. that function as food storage organs duri ...
: a short vertical underground stem with fleshy storage leaves attached, e.g.
onion
An onion (''Allium cepa'' L., from Latin ''cepa'' meaning "onion"), also known as the bulb onion or common onion, is a vegetable that is the most widely cultivated species of the genus ''Allium''. The shallot is a botanical variety of the onion ...
,
daffodil
''Narcissus'' is a genus of predominantly spring flowering perennial plants of the amaryllis family, Amaryllidaceae. Various common names including daffodil,The word "daffodil" is also applied to related genera such as '' Sternbergia'', ''Is ...
, and
tulip
Tulips (''Tulipa'') are a genus of spring-blooming perennial herbaceous bulbiferous geophytes (having bulbs as storage organs). The flowers are usually large, showy and brightly coloured, generally red, pink, yellow, or white (usually in warm ...
. Bulbs often function in reproduction by splitting to form new bulbs or producing small new bulbs termed bulblets. Bulbs are a combination of stem and leaves so may better be considered as leaves because the leaves make up the greater part.
* Caespitose: when stems grow in a tangled mass or clump or in low growing mats.
* Cladode (including
phylloclade
Phylloclades and cladodes are flattened, photosynthetic shoots, which are usually considered to be modified branches. The two terms are used either differently or interchangeably by different authors. ''Phyllocladus'', a genus of conifer, is name ...
): a flattened stem that appears more-or-less leaf like and is specialized for photosynthesis, e.g.
cactus
A cactus (, or less commonly, cactus) is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1750 known species of the order Caryophyllales. The word ''cactus'' derives, through Latin, from the Ancient Greek ...
pads.
*
Climbing
Climbing is the activity of using one's hands, feet, or any other part of the body to ascend a steep topographical object that can range from the world's tallest mountains (e.g. the eight thousanders), to small boulders. Climbing is done fo ...
: stems that cling or wrap around other plants or structures.
*
Corm
A corm, bulbo-tuber, or bulbotuber is a short, vertical, swollen underground plant stem that serves as a storage organ that some plants use to survive winter or other adverse conditions such as summer drought and heat (perennation).
The word ' ...
: a short enlarged underground, storage stem, e.g.
taro
Taro () (''Colocasia esculenta)'' is a root vegetable. It is the most widely cultivated species of several plants in the family Araceae that are used as vegetables for their corms, leaves, and petioles. Taro corms are a food staple in Africa ...
,
crocus
''Crocus'' (; plural: crocuses or croci) is a genus of seasonal flowering plants in the family Iridaceae (iris family) comprising about 100 species of perennials growing from corms. They are low growing plants, whose flower stems remain undergro ...
,
gladiolus
''Gladiolus'' (from Latin, the diminutive of ''gladius'', a sword) is a genus of perennial cormous flowering plants in the iris family (Iridaceae).
It is sometimes called the 'sword lily', but is usually called by its generic name (plural ''g ...
.
* Decumbent: stems that lie flat on the ground and turn upwards at the ends.
* Fruticose: stems that grow shrublike with woody like habit.
*
Herbaceous
Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials.
Definitions of "herb" and "herbaceous"
The fourth edition of t ...
: non woody, they die at the end of the growing season.
* Internode: an interval between two successive nodes. It possesses the ability to elongate, either from its base or from its extremity depending on the species.
* Node: a point of attachment of a leaf or a
twig
A twig is a thin, often short, branch of a tree or bush.
The buds on the twig are an important diagnostic characteristic, as are the abscission scars where the leaves have fallen away. The color, texture, and patterning of the twig bark are ...
on the stem in seed plants. A node is a very small growth zone.
*
Pedicel
Pedicle or pedicel may refer to:
Human anatomy
*Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures
...
: stems that serve as the stalk of an individual flower in an
inflorescence
An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
or
infrutescence Infructescence (fruiting head) is defined as the ensemble of fruits derived from the ovaries of an inflorescence. It usually retains the size and structure of the inflorescence.
In some cases, infructescences are similar in appearance to simple fru ...
.
*
Peduncle: a stem that supports an
inflorescence
An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
.
*
Prickle: a sharpened extension of the stem's outer layers, e.g.
rose
A rose is either a woody perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred species and tens of thousands of cultivars. They form a group of plants that can be ...
s.
* Pseudostem: a false stem made of the rolled bases of leaves, which may be tall, as in
banana
A banana is an elongated, edible fruit – botanically a berry – produced by several kinds of large herbaceous flowering plants in the genus ''Musa''. In some countries, bananas used for cooking may be called "plantains", distinguis ...
.
*
Rhizome
In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
: a horizontal underground stem that functions mainly in reproduction but also in storage, e.g. most
fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
s,
iris
Iris most often refers to:
*Iris (anatomy), part of the eye
*Iris (mythology), a Greek goddess
* ''Iris'' (plant), a genus of flowering plants
* Iris (color), an ambiguous color term
Iris or IRIS may also refer to:
Arts and media
Fictional ent ...
.
*
Runner
Running is a method of terrestrial locomotion allowing humans and other animals to move rapidly on foot. Running is a type of gait characterized by an aerial phase in which all feet are above the ground (though there are exceptions). This is ...
: a type of stolon, horizontally growing on top of the ground and rooting at the nodes, aids in reproduction. e.g.
garden strawberry
The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid species of the genus ''Fragaria'', collectively known as the strawberries, which are cultivated worldwide for their fruit. The fruit is widely ap ...
, ''
Chlorophytum comosum''.
*
Scape: a stem that holds flowers that comes out of the ground and has no normal leaves.
Hosta
''Hosta'' (, syn. ''Funkia'') is a genus of plants commonly known as hostas, plantain lilies and occasionally by the Japanese name gibōshi. Hostas are widely cultivated as shade-tolerant foliage plants. The genus is currently placed in the fa ...
,
lily
''Lilium'' () is a genus of Herbaceous plant, herbaceous flowering plants growing from bulbs, all with large prominent flowers. They are the true lilies. Lilies are a group of flowering plants which are important in culture and literature in mu ...
,
iris
Iris most often refers to:
*Iris (anatomy), part of the eye
*Iris (mythology), a Greek goddess
* ''Iris'' (plant), a genus of flowering plants
* Iris (color), an ambiguous color term
Iris or IRIS may also refer to:
Arts and media
Fictional ent ...
,
garlic
Garlic (''Allium sativum'') is a species of bulbous flowering plant in the genus ''Allium''. Its close relatives include the onion, shallot, leek, chive, Allium fistulosum, Welsh onion and Allium chinense, Chinese onion. It is native to South A ...
.
*
Stolon
In biology, stolons (from Latin '' stolō'', genitive ''stolōnis'' – "branch"), also known as runners, are horizontal connections between organisms. They may be part of the organism, or of its skeleton; typically, animal stolons are external s ...
: a horizontal stem that produces rooted plantlets at its nodes and ends, forming near the surface of the ground.
*
Thorn
Thorn(s) or The Thorn(s) may refer to:
Botany
* Thorns, spines, and prickles, sharp structures on plants
* ''Crataegus monogyna'', or common hawthorn, a plant species
Comics and literature
* Rose and Thorn, the two personalities of two DC Com ...
: a
modified stem with a sharpened point.
*
Tuber
Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing ...
: a swollen, underground storage stem adapted for storage and reproduction, e.g.
potato
The potato is a starchy food, a tuber of the plant ''Solanum tuberosum'' and is a root vegetable native to the Americas. The plant is a perennial in the nightshade family Solanaceae.
Wild potato species can be found from the southern Unit ...
.
*
Woody: hard textured stems with secondary xylem.
Stem structure
Stem usually consist of three tissues,
dermal tissue,
ground tissue
The ground tissue of plants includes all tissues that are neither Epidermis (botany), dermal nor Vascular tissue, vascular. It can be divided into three types based on the nature of the cell walls.
# Parenchyma cells have thin primary walls and ...
and
vascular tissue
Vascular tissue is a complex conducting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue are the xylem and phloem. These two tissues transport fluid and nutrients internally. There ...
. The dermal tissue covers the outer surface of the stem and usually functions to waterproof, protect and control gas exchange. The ground tissue usually consists mainly of parenchyma cells and fills in around the vascular tissue. It sometimes functions in photosynthesis. Vascular tissue provides long distance transport and structural support. Most or all ground tissue may be lost in woody stems. The dermal tissue of aquatic plants stems may lack the waterproofing found in aerial stems. The arrangement of the vascular tissues varies widely among plant species.
Dicot stems
Dicot
The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, t ...
stems with primary growth have
pith
Pith, or medulla, is a tissue in the stem
Stem or STEM may refer to:
Plant structures
* Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang
* Stipe (botany), a stalk to support some other ...
in the center, with vascular bundles forming a distinct ring visible when the stem is viewed in cross section. The outside of the stem is covered with an epidermis, which is covered by a waterproof cuticle. The epidermis also may contain
stomata
In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bor ...
for gas exchange and multicellular stem hairs called
trichomes
Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant ...
. A cortex consisting of
hypodermis
The subcutaneous tissue (), also called the hypodermis, hypoderm (), subcutis, superficial fascia, is the lowermost layer of the integumentary system in vertebrates. The types of cells found in the layer are fibroblasts, adipose cells, and macro ...
(collenchyma cells) and
endodermis
The endodermis is the central, innermost layer of cortex in land plants. It is a cylinder of compact living cells, the radial walls of which are impregnated with hydrophobic substances (Casparian strip) to restrict apoplastic flow of water to th ...
(starch containing cells) is present above the
pericycle
The pericycle is a cylinder of parenchyma or sclerenchyma cells that lies just inside the endodermis and is the outer most part of the stele of plants.
Although it is composed of non-vascular parenchyma cells, it's still considered part of the vas ...
and vascular bundles.
Woody dicots and many nonwoody dicots have
secondary growth
In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips of ...
originating from their lateral or secondary meristems: the
vascular cambium
The vascular cambium is the main growth tissue in the stems and roots of many plants, specifically in dicots such as buttercups and oak trees, gymnosperms such as pine trees, as well as in certain other vascular plants. It produces secondary xyle ...
and the
cork cambium
Cork cambium (pl. cambia or cambiums) is a tissue found in many vascular plants as a part of the epidermis. It is one of the many layers of bark, between the cork and primary phloem. The cork cambium is a lateral meristem and is responsible fo ...
or phellogen. The vascular cambium forms between the xylem and phloem in the vascular bundles and connects to form a continuous cylinder. The vascular cambium cells divide to produce
secondary xylem
Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word ''xylem'' is derived from ...
to the inside and
secondary phloem
Phloem (, ) is the living tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as ''photosynthates'', in particular the sugar sucrose, to the rest of the plant. This transport process is c ...
to the outside. As the stem increases in diameter due to production of secondary xylem and secondary phloem, the cortex and epidermis are eventually destroyed. Before the cortex is destroyed, a cork cambium develops there. The cork cambium divides to produce waterproof cork cells externally and sometimes phelloderm cells internally. Those three tissues form the
periderm
Bark is the outermost layers of stems and roots of woody plants. Plants with bark include trees, woody vines, and shrubs. Bark refers to all the tissues outside the vascular cambium and is a nontechnical term. It overlays the wood and consist ...
, which replaces the epidermis in function. Areas of loosely packed cells in the periderm that function in gas exchange are called lenticels.
Secondary
xylem
Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word ''xylem'' is derived from ...
is commercially important as
wood
Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin th ...
. The seasonal variation in growth from the
vascular cambium
The vascular cambium is the main growth tissue in the stems and roots of many plants, specifically in dicots such as buttercups and oak trees, gymnosperms such as pine trees, as well as in certain other vascular plants. It produces secondary xyle ...
is what creates yearly tree rings in temperate climates. Tree rings are the basis of
dendrochronology
Dendrochronology (or tree-ring dating) is the scientific method of dating tree rings (also called growth rings) to the exact year they were formed. As well as dating them, this can give data for dendroclimatology, the study of climate and atmos ...
, which dates wooden objects and associated artifacts.
Dendroclimatology
Dendroclimatology is the science of determining past climates from trees (primarily properties of the annual tree rings). Tree rings are wider when conditions favor growth, narrower when times are difficult. Other properties of the annual rings, s ...
is the use of tree rings as a record of past climates. The aerial stem of an adult
tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
is called a
trunk. The dead, usually darker inner wood of a large diameter trunk is termed the
heartwood
Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin tha ...
and is the result of
tylosis. The outer, living wood is termed the sapwood.
Monocot stems
![Royal Palm Trunk](https://upload.wikimedia.org/wikipedia/commons/b/b5/Royal_Palm_Trunk.jpg)
Vascular bundles are present throughout the
monocot
Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one of ...
stem, although concentrated towards the outside. This differs from the dicot stem that has a ring of vascular bundles and often none in the center. The shoot apex in monocot stems is more elongated. Leaf sheathes grow up around it, protecting it. This is true to some extent of almost all monocots. Monocots rarely produce
secondary growth
In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips of ...
and are therefore seldom woody, with
palms and
bamboo
Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, bu ...
being notable exceptions. However, many monocot stems increase in diameter via anomalous secondary growth.
Gymnosperm stems
All
gymnosperm
The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, ''Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμνό ...
s are woody plants. Their stems are similar in structure to woody dicots except that most gymnosperms produce only
tracheid
A tracheid is a long and tapered lignified cell in the xylem of vascular plants. It is a type of conductive cell called a tracheary element. Angiosperms use another type of tracheary element, called vessel elements, to transport water through th ...
s in their xylem, not the vessels found in dicots. Gymnosperm wood also often contains
resin
In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on natu ...
ducts. Woody dicots are called hardwoods, e.g.
oak
An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...
,
maple
''Acer'' () is a genus of trees and shrubs commonly known as maples. The genus is placed in the family Sapindaceae.Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 9, June 2008 nd more or less continuously updated since http ...
and
walnut
A walnut is the edible seed of a drupe of any tree of the genus ''Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''.
Although culinarily considered a "nut" and used as such, it is not a true ...
. In contrast, softwoods are gymnosperms, such as
pine
A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
,
spruce
A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfami ...
and
fir.
Image:Redwood M D Vaden.jpg, The trunk of this redwood
Sequoioideae, popularly known as redwoods, is a subfamily of coniferous trees within the family
Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affini ...
tree is its stem.
Fern stems
Most
fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
s have
rhizome
In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
s with no vertical stem. The exception is
tree ferns
The tree ferns are arborescent (tree-like) ferns that grow with a trunk elevating the fronds above ground level, making them trees. Many extant tree ferns are members of the order Cyatheales, to which belong the families Cyatheaceae (scaly tre ...
, with vertical stems up to about 20 metres. The stem anatomy of ferns is more complicated than that of dicots because fern stems often have one or more leaf gaps in cross section. A leaf gap is where the vascular tissue branches off to a
frond
A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the lar ...
. In cross section, the vascular tissue does not form a complete cylinder where a leaf gap occurs. Fern stems may have
solenosteles or
dictyosteles or variations of them. Many fern stems have phloem tissue on both sides of the xylem in cross-section.
File:Ferns.jpg, Tasmanian tree fern
''Dicksonia antarctica'', the soft tree fern or man fern, is a species of evergreen tree fern native to eastern Australia, ranging from south-east Queensland, coastal New South Wales and Victoria to Tasmania.
Anatomy and biology
These ferns ...
Relation to xenobiotics
Foreign chemicals such as air pollutants,
[C. Michael Hogan. 2010]
"Abiotic factor"
''Encyclopedia of Earth''. Emily Monosson and C. Cleveland, eds. National Council for Science and the Environment . Washington, D.C. herbicides and pesticides can damage stem structures.
Economic importance
There are thousands of species whose stems have economic uses. Stems provide a few major staple crops such as
potato
The potato is a starchy food, a tuber of the plant ''Solanum tuberosum'' and is a root vegetable native to the Americas. The plant is a perennial in the nightshade family Solanaceae.
Wild potato species can be found from the southern Unit ...
and
taro
Taro () (''Colocasia esculenta)'' is a root vegetable. It is the most widely cultivated species of several plants in the family Araceae that are used as vegetables for their corms, leaves, and petioles. Taro corms are a food staple in Africa ...
.
Sugarcane
Sugarcane or sugar cane is a species of (often hybrid) tall, Perennial plant, perennial grass (in the genus ''Saccharum'', tribe Andropogoneae) that is used for sugar Sugar industry, production. The plants are 2–6 m (6–20 ft) tall with ...
stems are a major source of sugar.
Maple sugar is obtained from trunks of
maple
''Acer'' () is a genus of trees and shrubs commonly known as maples. The genus is placed in the family Sapindaceae.Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 9, June 2008 nd more or less continuously updated since http ...
trees.
Vegetable
Vegetables are parts of plants that are consumed by humans or other animals as food. The original meaning is still commonly used and is applied to plants collectively to refer to all edible plant matter, including the flowers, fruits, stems, ...
s from stems are
asparagus
Asparagus, or garden asparagus, folk name sparrow grass, scientific name ''Asparagus officinalis'', is a perennial flowering plant species in the genus ''Asparagus''. Its young shoots are used as a spring vegetable.
It was once classified in ...
,
bamboo shoot
Bamboo shoots or bamboo sprouts are the edible shoots (new bamboo culms that come out of the ground) of many bamboo species including ''Bambusa vulgaris'' and ''Phyllostachys edulis''. They are used as vegetables in numerous Asian dishes and b ...
s, cactus pads or
nopalito
Nopalitos is a dish made with diced nopales, the naturally flat stems, called pads, of prickly pear cactus (''Opuntia''). They are sold fresh, bottled, or canned and less often dried. They have a light, slightly tart flavor, and a crisp, muci ...
s,
kohlrabi
Kohlrabi (pronounced ; scientific name ''Brassica oleracea'' Gongylodes Group (horticulture), Group), also called German turnip or turnip cabbage, is a Biennial plant, biennial vegetable, a low, stout cultivar of wild cabbage. It is a cultivar o ...
, and
water chestnut Water chestnut may refer to either of two plants (both sometimes used in Chinese cuisine):
* The Chinese water chestnut ('' Eleocharis dulcis''), eaten for its crisp corm
* The water caltrop
The water caltrop is any of three extant species of th ...
. The spice,
cinnamon
Cinnamon is a spice obtained from the inner bark of several tree species from the genus ''Cinnamomum''. Cinnamon is used mainly as an aromatic condiment and flavouring additive in a wide variety of cuisines, sweet and savoury dishes, breakfa ...
is bark from a tree trunk.
Gum arabic
Gum arabic, also known as gum sudani, acacia gum, Arabic gum, gum acacia, acacia, Senegal gum, Indian gum, and by other names, is a natural gum originally consisting of the hardened sap of two species of the '' Acacia'' tree, ''Senegalia sen ...
is an important food additive obtained from the trunks of ''
Acacia senegal
''Senegalia senegal'' (also known as ''Acacia senegal'') is a small thorny deciduous tree from the genus '' Senegalia'', which is known by several common names, including gum acacia, gum arabic tree, Sudan gum and Sudan gum arabic. In parts of I ...
'' trees.
Chicle
Chicle () is a natural gum traditionally used in making chewing gum and other products. It is collected from several species of Mesoamerican trees in the genus ''Manilkara'', including '' M. zapota'', '' M. chicle'', '' M. staminodella'', and '' ...
, the main ingredient in
chewing gum
Chewing gum is a soft, cohesive substance designed to be chewed without being swallowed. Modern chewing gum is composed of gum base, sweeteners, softeners/plasticizers, flavors, colors, and, typically, a hard or powdered polyol coating. Its te ...
, is obtained from trunks of the chicle tree.
Medicines obtained from stems include
quinine
Quinine is a medication used to treat malaria and babesiosis. This includes the treatment of malaria due to ''Plasmodium falciparum'' that is resistant to chloroquine when artesunate is not available. While sometimes used for nocturnal leg cr ...
from the bark of
cinchona
''Cinchona'' (pronounced or ) is a genus of flowering plants in the family Rubiaceae containing at least 23 species of trees and shrubs. All are native to the Tropical Andes, tropical Andean forests of western South America. A few species are ...
trees,
camphor
Camphor () is a waxy, colorless solid with a strong aroma. It is classified as a terpenoid and a cyclic ketone. It is found in the wood of the camphor laurel ('' Cinnamomum camphora''), a large evergreen tree found in East Asia; and in the k ...
distilled from wood of a tree in the same genus that provides cinnamon, and the muscle relaxant
curare
Curare ( /kʊˈrɑːri/ or /kjʊˈrɑːri/; ''koo-rah-ree'' or ''kyoo-rah-ree'') is a common name for various alkaloid arrow poisons originating from plant extracts. Used as a paralyzing agent by indigenous peoples in Central and South ...
from the bark of tropical vines.
Wood
Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin th ...
is used in thousands of ways, e.g.
buildings,
furniture
Furniture refers to movable objects intended to support various human activities such as seating (e.g., stools, chairs, and sofas), eating (tables), storing items, eating and/or working with an item, and sleeping (e.g., beds and hammocks). Fu ...
,
boat
A boat is a watercraft of a large range of types and sizes, but generally smaller than a ship, which is distinguished by its larger size, shape, cargo or passenger capacity, or its ability to carry boats.
Small boats are typically found on inl ...
s,
airplane
An airplane or aeroplane (informally plane) is a fixed-wing aircraft that is propelled forward by thrust from a jet engine, propeller, or rocket engine. Airplanes come in a variety of sizes, shapes, and wing configurations. The broad spe ...
s,
wagon
A wagon or waggon is a heavy four-wheeled vehicle pulled by draught animals or on occasion by humans, used for transporting goods, commodities, agricultural materials, supplies and sometimes people.
Wagons are immediately distinguished from ...
s,
car
A car or automobile is a motor vehicle with wheels. Most definitions of ''cars'' say that they run primarily on roads, seat one to eight people, have four wheels, and mainly transport people instead of goods.
The year 1886 is regarded as ...
parts,
musical instrument
A musical instrument is a device created or adapted to make musical sounds. In principle, any object that produces sound can be considered a musical instrument—it is through purpose that the object becomes a musical instrument. A person who pl ...
s,
sports equipment
Sports equipment, sporting equipment, also called sporting goods, are the tools, materials, apparel, and gear used to compete in a sport and varies depending on the sport. The equipment ranges from balls, nets, and protective gear like helmets. ...
,
railroad tie
A railroad tie, crosstie (American English), railway tie (Canadian English) or railway sleeper (Australian and British English) is a rectangular support for the rails in railroad tracks. Generally laid perpendicular to the rails, ties transfer ...
s,
utility pole
A utility pole is a column or post typically made out of wood used to support overhead power lines and various other public utilities, such as electrical cable, optical fiber, fiber optic cable, and related equipment such as Distribution transfor ...
s, fence posts,
pilings
A deep foundation is a type of foundation (architecture), foundation that transfers building loads to the earth farther down from the surface than a shallow foundation does to a subsurface layer or a range of depths. A pile or piling is a ...
,
toothpick
A toothpick is a small thin stick of wood, plastic, bamboo, metal, bone or other substance with at least one and sometimes two pointed ends to insert between teeth to remove detritus, usually after a meal. Toothpicks are also used for festive ...
s,
match
A match is a tool for starting a fire. Typically, matches are made of small wooden sticks or stiff paper. One end is coated with a material that can be ignited by friction generated by striking the match against a suitable surface. Wooden matc ...
es,
plywood
Plywood is a material manufactured from thin layers or "plies" of wood veneer that are glued together with adjacent layers having their wood grain rotated up to 90 degrees to one another. It is an engineered wood from the family of manufactured ...
,
coffin
A coffin is a funerary box used for viewing or keeping a corpse, either for burial or cremation.
Sometimes referred to as a casket, any box in which the dead are buried is a coffin, and while a casket was originally regarded as a box for jewel ...
s,
shingles
Shingles, also known as zoster or herpes zoster, is a viral disease characterized by a painful skin rash with blisters in a localized area. Typically the rash occurs in a single, wide mark either on the left or right side of the body or face. ...
,
barrel
A barrel or cask is a hollow cylindrical container with a bulging center, longer than it is wide. They are traditionally made of wooden staves and bound by wooden or metal hoops. The word vat is often used for large containers for liquids, ...
staves,
toy
A toy or plaything is an object that is used primarily to provide entertainment. Simple examples include toy blocks, board games, and dolls. Toys are often designed for use by children, although many are designed specifically for adults and pet ...
s,
tool
A tool is an object that can extend an individual's ability to modify features of the surrounding environment or help them accomplish a particular task. Although many animals use simple tools, only human beings, whose use of stone tools dates ba ...
handles,
picture frame
A picture frame is a protective and decorative edging for a picture, such as a painting or photograph. It makes displaying the work safer and easier and both sets the picture apart from its surroundings and aesthetically integrates it with them. ...
s,
veneer
Veneer may refer to:
Materials
* Veneer (dentistry), a cosmetic treatment for teeth
* Masonry veneer, a thin facing layer of brick
* Stone veneer, a thin facing layer of stone
* Wood veneer, a thin facing layer of wood
Arts and entertainment
* ' ...
,
charcoal
Charcoal is a lightweight black carbon residue produced by strongly heating wood (or other animal and plant materials) in minimal oxygen to remove all water and volatile constituents. In the traditional version of this pyrolysis process, cal ...
and
firewood
Firewood is any wooden material that is gathered and used for fuel. Generally, firewood is not highly processed and is in some sort of recognizable log or branch form, compared to other forms of wood fuel like pellets or chips. Firewood can ...
.
Wood pulp
Pulp is a lignocellulosic fibrous material prepared by chemically or mechanically separating cellulose fibers from wood, fiber crops, waste paper, or rags. Mixed with water and other chemical or plant-based additives, pulp is the major raw mate ...
is widely used to make
paper
Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, rags, grasses or other vegetable sources in water, draining the water through fine mesh leaving the fibre evenly distributed ...
,
paperboard
Paperboard is a thick paper-based material. While there is no rigid differentiation between paper and paperboard, paperboard is generally thicker (usually over 0.30 mm, 0.012 in, or 12 Inch#equivalences, points) than paper and has certain ...
,
cellulose
Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell wall ...
sponges,
cellophane
Cellophane is a thin, transparent sheet made of regenerated cellulose. Its low permeability to air, oils, greases, bacteria, and liquid water makes it useful for food packaging. Cellophane is highly permeable to water vapour, but may be coated w ...
and some important
plastic
Plastics are a wide range of synthetic or semi-synthetic materials that use polymers as a main ingredient. Their plasticity makes it possible for plastics to be moulded, extruded or pressed into solid objects of various shapes. This adaptab ...
s and
textile
Textile is an umbrella term that includes various fiber-based materials, including fibers, yarns, filaments, threads, different fabric types, etc. At first, the word "textiles" only referred to woven fabrics. However, weaving is not the ...
s, such as
cellulose acetate
In biochemistry, cellulose acetate refers to any acetate ester of cellulose, usually cellulose diacetate. It was first prepared in 1865. A bioplastic, cellulose acetate is used as a film base in photography, as a component in some coatings, and ...
and
rayon
Rayon is a semi-synthetic fiber, made from natural sources of regenerated cellulose, such as wood and related agricultural products. It has the same molecular structure as cellulose. It is also called viscose. Many types and grades of viscose f ...
.
Bamboo
Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, bu ...
stems also have hundreds of uses, including paper, buildings, furniture, boats, musical instruments,
fishing pole
A fishing rod is a long, thin rod used by anglers to catch fish by manipulating a line ending in a hook (formerly known as an ''angle'', hence the term "angling"). At its most basic form, a fishing rod is a straight rigid stick/pole with a ...
s,
water pipe
Plumbing is any system that conveys fluids for a wide range of applications. Plumbing uses pipes, valves, plumbing fixtures, tanks, and other apparatuses to convey fluids. Heating and cooling (HVAC), waste removal, and potable water delivery ...
s, plant stakes, and
scaffolding
Scaffolding, also called scaffold or staging, is a temporary structure used to support a work crew and materials to aid in the construction, maintenance and repair of buildings, bridges and all other man-made structures. Scaffolds are widely use ...
. Trunks of
palm trees
Palm most commonly refers to:
* Palm of the hand, the central region of the front of the hand
* Palm plants, of family Arecaceae
**List of Arecaceae genera
* Several other plants known as "palm"
Palm or Palms may also refer to:
Music
* Palm ( ...
and
tree fern
The tree ferns are arborescent (tree-like) ferns that grow with a trunk elevating the fronds above ground level, making them trees. Many extant tree ferns are members of the order Cyatheales, to which belong the families Cyatheaceae (scaly tree ...
s are often used for building. Stems of
reed
Reed or Reeds may refer to:
Science, technology, biology, and medicine
* Reed bird (disambiguation)
* Reed pen, writing implement in use since ancient times
* Reed (plant), one of several tall, grass-like wetland plants of the order Poales
* ...
are an important building material for use in
thatching
Thatching is the craft of building a roof with dry vegetation such as straw, water reed, sedge (''Cladium mariscus''), rushes, heather, or palm branches, layering the vegetation so as to shed water away from the inner roof. Since the bulk of ...
in some areas.
Tannin
Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids.
The term ''tannin'' (from Anglo-Norman ''tanner'', ...
s used for tanning
leather
Leather is a strong, flexible and durable material obtained from the tanning, or chemical treatment, of animal skins and hides to prevent decay. The most common leathers come from cattle, sheep, goats, equine animals, buffalo, pigs and hogs, ...
are obtained from the wood of certain trees, such as
quebracho.
Cork
Cork or CORK may refer to:
Materials
* Cork (material), an impermeable buoyant plant product
** Cork (plug), a cylindrical or conical object used to seal a container
***Wine cork
Places Ireland
* Cork (city)
** Metropolitan Cork, also known as G ...
is obtained from the bark of the
cork oak
''Quercus suber'', commonly called the cork oak, is a medium-sized, evergreen oak tree in the section ''Quercus'' sect. ''Cerris''. It is the primary source of cork for wine bottle stoppers and other uses, such as cork flooring and as the cores ...
.
Rubber
Rubber, also called India rubber, latex, Amazonian rubber, ''caucho'', or ''caoutchouc'', as initially produced, consists of polymers of the organic compound isoprene, with minor impurities of other organic compounds. Thailand, Malaysia, and ...
is obtained from the trunks of ''
Hevea brasiliensis
''Hevea brasiliensis'', the Pará rubber tree, ''sharinga'' tree, seringueira, or most commonly, rubber tree or rubber plant, is a flowering plant belonging to the spurge family Euphorbiaceae
Euphorbiaceae, the spurge family, is a large fami ...
.''
Rattan
Rattan, also spelled ratan, is the name for roughly 600 species of Old World climbing palms belonging to subfamily Calamoideae. The greatest diversity of rattan palm species and genera are in the closed-canopy old-growth tropical forests of ...
, used for furniture and baskets, is made from the stems of tropical vining palms.
Bast fiber
Bast fibre (also called phloem fibre or skin fibre) is plant fibre collected from the phloem (the "inner bark", sometimes called "skin") or bast surrounding the stem of certain dicotyledonous plants. It supports the conductive cells of the phlo ...
s for textiles and rope are obtained from stems include
flax
Flax, also known as common flax or linseed, is a flowering plant, ''Linum usitatissimum'', in the family Linaceae. It is cultivated as a food and fiber crop in regions of the world with temperate climates. Textiles made from flax are known in ...
,
hemp
Hemp, or industrial hemp, is a botanical class of ''Cannabis sativa'' cultivars grown specifically for industrial or medicinal use. It can be used to make a wide range of products. Along with bamboo, hemp is among the fastest growing plants o ...
,
jute
Jute is a long, soft, shiny bast fiber that can be spun into coarse, strong threads. It is produced from flowering plants in the genus ''Corchorus'', which is in the mallow family Malvaceae. The primary source of the fiber is ''Corchorus olit ...
and
ramie
Ramie (pronounced: , ; from Malay ) is a flowering plant in the nettle family Urticaceae, native to eastern Asia. It is a herbaceous perennial growing to tall; . The earliest paper was obtained from the stems of
papyrus
Papyrus ( ) is a material similar to thick paper that was used in ancient times as a writing surface. It was made from the pith of the papyrus plant, '' Cyperus papyrus'', a wetland sedge. ''Papyrus'' (plural: ''papyri'') can also refer to a ...
by the ancient Egyptians.
Amber
Amber is fossilized tree resin that has been appreciated for its color and natural beauty since Neolithic times. Much valued from antiquity to the present as a gemstone, amber is made into a variety of decorative objects."Amber" (2004). In Ma ...
is fossilized sap from tree trunks; it is used for
jewelry
Jewellery ( UK) or jewelry (U.S.) consists of decorative items worn for personal adornment, such as brooches, rings, necklaces, earrings, pendants, bracelets, and cufflinks. Jewellery may be attached to the body or the clothes. From a western ...
and may contain ancient animals. Resins from conifer wood are used to produce
turpentine
Turpentine (which is also called spirit of turpentine, oil of turpentine, terebenthene, terebinthine and (colloquially) turps) is a fluid obtained by the distillation of resin harvested from living trees, mainly pines. Mainly used as a special ...
and
rosin
Rosin (), also called colophony or Greek pitch ( la, links=no, pix graeca), is a solid form of resin obtained from pines and some other plants, mostly conifers, produced by heating fresh liquid resin to vaporize the volatile liquid terpene comp ...
. Tree bark is often used as a
mulch
A mulch is a layer of material applied to the surface of soil. Reasons for applying mulch include conservation of soil moisture, improving fertility and health of the soil, reducing weed growth and enhancing the visual appeal of the area.
A m ...
and in growing media for container plants. It also can become the natural
habitat
In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
of
lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.[paper birch
''Betula papyrifera'' (paper birch, also known as (American) white birch and canoe birch) is a short-lived species of birch native to northern North America. Paper birch is named for the tree's thin white bark, which often peels in paper like ...]
* Twisted branches of
corkscrew willow
''Salix babylonica'' (Babylon willow or weeping willow; ) is a species of willow native to dry areas of northern China, but cultivated for millennia elsewhere in Asia, being traded along the Silk Road to southwest Asia and Europe.Flora of China' ...
and
Harry Lauder's walking stick (''Corylus avellana'' 'Contorta')
* Red, peeling bark of
paperbark maple
''Acer griseum'', the paperbark maple or blood-bark maple, is a species of flowering plant in the family Sapindaceae, native to central China.Flora of China (draft)''Acer griseum''/ref> ''Acer griseum'' is found in the Chinese provinces of Gan ...
References
Further reading
*
External links
*
{{Authority control
Stem
Stem or STEM may refer to:
Plant structures
* Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang
* Stipe (botany), a stalk to support some other structure
* Stipe (mycology), the stem of a mushro ...
Stem
Stem or STEM may refer to:
Plant structures
* Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang
* Stipe (botany), a stalk to support some other structure
* Stipe (mycology), the stem of a mushro ...