Cyatheales
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Cyatheales
The order Cyatheales, which includes most tree ferns, is a taxonomic order of the fern class, Polypodiopsida. No clear morphological features characterize all of the Cyatheales, but DNA sequence data indicate the order is monophyletic. Some species in the Cyatheales have tree-like growth forms from a vertical rhizome, others have shorter or horizontal expanding rhizomes. Some species have scales on the stems and leaves, while others have hairs. However, most plants in the Cyatheales are tree ferns and have trunk-like stems up to tall. It is unclear how many times the tree form has evolved and been lost in the order.Judd, W.S., C.S. Campbell, E.A. Kellogg, P.F. Stevens, and M.J. Donoghue (Eds.) 2008. Plant Systematics: A Phylogenetic Approach, Third Edition. Sinauer Associates, Sunderland, Massachusetts, USA. Description While the Cyatheales have been shown to be monophyletic through molecular analysis, no prominent morphological characteristics are common to the entire group ...
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Cyatheaceae
The Cyatheaceae are a family of ferns, the scaly tree ferns, one of eight families in the order Cyatheales in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the family may defined much more broadly (Cyatheaceae ''sensu lato'') as the only family in the Cyatheales, with the PPG I family treated as the subfamily Cyatheoideae. The narrower circumscription is used in this article. The family includes the world's tallest tree ferns, which reach heights up to 20 m. They are also very ancient plants, appearing in the fossil record in the late Jurassic, though the modern genera likely appeared in the Cenozoic. Cyatheaceae are the largest family of tree ferns, including about 640 species. Cyatheaceae and Dicksoniaceae, together with Metaxyaceae and Cibotiaceae, do not form a strongly supported monophyletic group and could be paraphyletic, but several individual subgroups are well supported as being monophyletic. Cyatheaceae are leptosporangiate ferns, the ...
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Thyrsopteridaceae
''Thyrsopteris'' is a genus of tree fern. It contains a single living species, ''Thyrsopteris elegans'', endemic to the Juan Fernandez Archipelago off the coast of Chile. ''Thyrsopteris'' is the only genus in the family Thyrsopteridaceae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the genus may be placed in the subfamily Thyrsopteridoideae of a more broadly defined family Cyatheaceae, the family placement used in ''Plants of the World Online'' . The oldest records of the genus are the species ''Thyrsopteris cretacea'' and ''Thyrsopteris cyathindusia'' which were described from the Burmese amber of Myanmar, dating to the Cenomanian of the Cretaceous period, around 99 million years ago. Other fossil species include ''Thyrsopteris antiqua'' from the Upper Cretaceous of Chile and ''Thyrsopteris shenii'' from the Paleogene of King George Island, Antarctica A thyrsopterid rachis In biology, a rachis (from the grc, ῥάχις [], "backbone, s ...
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Tree Ferns
The tree ferns are arborescent (tree-like) ferns that grow with a trunk elevating the fronds above ground level, making them trees. Many extant tree ferns are members of the order Cyatheales, to which belong the families Cyatheaceae (scaly tree ferns), Dicksoniaceae, Metaxyaceae, and Cibotiaceae. It is estimated that Cyatheales originated in the early Jurassic, and is the third group of ferns known to have given rise to tree-like forms. The others are the extinct ''Tempskya'' of uncertain position, and Osmundales where the extinct Guaireaceae and some members of Osmundaceae also grew into trees. In addition there were the Psaroniaceae and Tietea in the Marattiales, which is the sister group to most living ferns including Cyatheales. Other ferns which are also tree ferns, are ''Leptopteris'' and ''Todea'' in the family Osmundaceae, which can achieve short trunks under a metre tall. Fern species with short trunks in the genera ''Blechnum'', '' Cystodium'' and ''Sadleria'' from ...
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Polypodiopsida
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Metaxyaceae
''Metaxya'' is a neotropical genus of ferns in the order Cyatheales. It is the only genus in the family Metaxyaceae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the genus may be placed in the subfamily Metaxyoideae of a more broadly defined family Cyatheaceae, the family placement used in ''Plants of the World Online'' . The species of the genus are characterized by large fronds that approach 8 ft (2.5 m) in length. Species , ''Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants by ...'' accepted the following species: References External links * * Cyatheales Fern genera {{Cyatheales-stub ...
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Loxsomataceae
The Loxsomataceae are a family of ferns in the order Cyatheales with two extant genera in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the family may be treated as the subfamily Loxsomatoideae of a very broadly defined family Cyatheaceae, the system used in ''Plants of the World Online'' . It has leaves that can be as long as long. Extant species in this family are found in New Zealand, Costa Rica, and South America. Fossil species in this family, dating from as early as the Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ..., have been found in North America, India and Japan. Genera Two genera are placed in the family in the Pteridophyte Phylogeny Group classification of 2016 (PPG I), each with only one species: *'' Loxsoma'' R.Br. *'' ...
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Fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Culcitaceae
''Culcita'' is a genus of ferns, native to the Americas, Macaronesia and Iberian Peninsula. It is the only genus in the family Culcitaceae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the family may be treated as the subfamily Culcitoideae of a very broadly defined family Cyatheaceae, the placement used for the genus in ''Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants by ...'' . Species Only two species are known: References Cyatheales Fern genera {{Cyatheales-stub ...
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Plagiogyriaceae
''Plagiogyria'' is a genus of ferns, the only genus in family Plagiogyriaceae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the family may be treated as the subfamily Plagiogyrioideae of a very broadly defined family Cyatheaceae, the placement used for the genus in ''Plants of the World Online'' . Ferns of this genus present two kind of fronds, the fertile ones longer than the sterile. These ferns are found on forest soils in mountainous areas of tropical and subtropical regions. Most are native to Asia; one is found in the Americas.Plagiogyriaceae.
Flora of China.


Species

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Dicksonia
''Dicksonia'' is a genus of tree ferns in the order Cyatheales. It is regarded as related to ''Cyathea'', but is considered to retain more primitive traits, dating back at least to the Jurassic and Cretaceous periods. The fossil record includes stems, pinnules, and spores. The genus contains 20–25 species, distributed from Mexico to Argentina, Brazil, Uruguay and Chile, St. Helena, New Zealand, New Caledonia, Australia, Indonesia, New Guinea, and the Philippines. New Guinea has the greatest diversity, with five species. Species of ''Dicksonia'' found in cultivation include: *''D. antarctica'', soft tree fern *''D. fibrosa'', woolly tree fern *''D. squarrosa'', rough or slender tree fern The genus was first described by Charles Louis L'Héritier de Brutelle in 1788. The name honors James Dickson, a prominent nurseryman and botanist.''Dicksonia antarctica'' – the soft tree fernAustralian National Botanic Gardens 2008 Species * ''Dicksonia antarctica'', soft tree fern, Ta ...
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Dicksonia Antarctica 20 - Kew Gardens
''Dicksonia'' is a genus of tree ferns in the order Cyatheales. It is regarded as related to ''Cyathea'', but is considered to retain more primitive traits, dating back at least to the Jurassic and Cretaceous periods. The fossil record includes stems, pinnules, and spores. The genus contains 20–25 species, distributed from Mexico to Argentina, Brazil, Uruguay and Chile, St. Helena, New Zealand, New Caledonia, Australia, Indonesia, New Guinea, and the Philippines. New Guinea has the greatest diversity, with five species. Species of ''Dicksonia'' found in cultivation include: *''D. antarctica'', soft tree fern *''D. fibrosa'', woolly tree fern *''D. squarrosa'', rough or slender tree fern The genus was first described by Charles Louis L'Héritier de Brutelle in 1788. The name honors James Dickson, a prominent nurseryman and botanist.''Dicksonia antarctica'' – the soft tree fernAustralian National Botanic Gardens 2008 Species * ''Dicksonia antarctica'', soft tree fern, Ta ...
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Osmundaceae
Osmundaceae (royal fern family) is a family of ferns containing four to six extant genera and 18–25 known species. It is the only living family of the order Osmundales in the class Polypodiopsida (ferns) or in some classifications the only order in the class Osmundopsida. This is an ancient (known from the Upper Permian) and fairly isolated group that is often known as the "flowering ferns" because of the striking aspect of the ripe sporangia in ''Claytosmunda'', ''Osmunda'', ''Osmundastrum'', and ''Plensium'' (subtribe Osmundinae). In these genera the sporangia are borne naked on non-laminar pinnules, while ''Todea'' and ''Leptopteris'' (subtribe Todinae) bear sporangia naked on laminar pinnules. Ferns in this family are larger than most other ferns. Description The stems of Osmundaceae contain vascular tissue arranged as an ectophloic siphonostele; that is, a ring of phloem occurs on the outside only of a ring of xylem, which surrounds pith (and no other vascular tissue). S ...
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