Human–gorilla Last Common Ancestor
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Homininae (the hominines) is a subfamily of the family
Hominidae The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic Family (biology), family of primates that includes eight Neontology#Extant taxa versus extinct taxa, extant species in four Genus, genera: ''Orangutan ...
(hominids). (The Homininae——encompass humans, and are also called "African hominids" or "African apes".) This subfamily includes two tribes,
Hominini The Hominini (hominins) form a Tribe (biology), taxonomic tribe of the subfamily Homininae (hominines). They comprise two extant genera: ''Homo'' (humans) and ''Pan (genus), Pan'' (chimpanzees and bonobos), and in standard usage exclude the gen ...
and
Gorillini Gorillini is a taxonomic tribe containing three genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomencla ...
, both having
extant Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Exta ...
(or living) species as well as
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
species. Tribe
Hominini The Hominini (hominins) form a Tribe (biology), taxonomic tribe of the subfamily Homininae (hominines). They comprise two extant genera: ''Homo'' (humans) and ''Pan (genus), Pan'' (chimpanzees and bonobos), and in standard usage exclude the gen ...
includes: the extant genus ''
Homo ''Homo'' () is a genus of great ape (family Hominidae) that emerged from the genus ''Australopithecus'' and encompasses only a single extant species, ''Homo sapiens'' (modern humans), along with a number of extinct species (collectively called ...
'', which comprises only one extant species—the modern
human Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
s (''Homo'' sapiens), and numerous extinct human species; and the extant genus '' Pan'', which includes two extant species,
chimpanzee The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of Hominidae, great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close rel ...
s and
bonobo The bonobo (; ''Pan paniscus''), also historically called the pygmy chimpanzee (less often the dwarf chimpanzee or gracile chimpanzee), is an endangered great ape and one of the two species making up the genus ''Pan (genus), Pan'' (the other bei ...
s. Tribe
Gorillini Gorillini is a taxonomic tribe containing three genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomencla ...
(
gorilla Gorillas are primarily herbivorous, terrestrial great apes that inhabit the tropical forests of equatorial Africa. The genus ''Gorilla'' is divided into two species: the eastern gorilla and the western gorilla, and either four or five su ...
s) contains one extant genus, Gorilla, with two extant species, with variants, and one known extinct genus. Alternatively, the genus ''Pan'' is considered by some to belong, instead of to a subtribe
Panina Panina is a subtribe of tribe Hominini; it comprises all descendants of the human-chimpanzee last common ancestor (LCA) that are ''not'' of the branch of human lineage—that is, all those ancestors of the type genus '' Pan'' (chimpanzees and ...
, to its own separate tribe, (''so-called'') "Panini"—which would be a third tribe for Homininae. Some classification schemes provide a more comprehensive account of extinct groups—(see section "Taxonomic Classification", below). For example, tribe Hominini shows two subtribes: subtribe
Hominina The australopithecines (), formally Australopithecina or Hominina, are generally any species in the related genera of ''Australopithecus'' and ''Paranthropus''. It may also include members of ''Kenyanthropus'', ''Ardipithecus'', and '' Praeanth ...
, which contains at least two extinct
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
; and subtribe Panina, which presents only the extant genus, ''Pan'' (chimpanzees/bonobos), as fossils of extinct chimpanzees/bonobos are very rarely found. The Homininae comprise all
hominid The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
s that arose after the subfamily
Ponginae Ponginae , also known as the Asian hominids, is a subfamily in the family (biology), family Hominidae. Once a diverse lineage of Eurasian apes, the subfamily has only one Neontology, extant genus, ''Pongo (genus), Pongo'' (orangutans), which con ...
(
orangutan Orangutans are great apes native to the rainforests of Indonesia and Malaysia. They are now found only in parts of Borneo and Sumatra, but during the Pleistocene they ranged throughout Southeast Asia and South China. Classified in the genus ...
s) split from the line of the
great ape The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
s. The Homininae
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
has three main branches leading: to gorillas (via the tribe Gorillini); to humans and to chimpanzees (via the tribe Hominini and subtribes Hominina and Panina―see graphic "Evolutionary tree", below). There are two living species of Panina, chimpanzees and bonobos, and two living species of gorillas and one that is extinct. Traces of extinct ''Homo'' species, including ''
Homo floresiensis ''Homo floresiensis'' , also known as "Flores Man" or "Hobbit" (after Hobbit, the fictional species), is an Extinction, extinct species of small archaic humans that inhabited the island of Flores, Indonesia, until the arrival of Homo sapiens, ...
'', have been found with dates as recent as 40,000 years ago. Individual members of this subfamily are called hominine or hominines—not to be confused with the terms
hominin The Hominini (hominins) form a taxonomic tribe of the subfamily Homininae (hominines). They comprise two extant genera: ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos), and in standard usage exclude the genus '' Gorilla'' ( gorillas) ...
s or
Hominini The Hominini (hominins) form a Tribe (biology), taxonomic tribe of the subfamily Homininae (hominines). They comprise two extant genera: ''Homo'' (humans) and ''Pan (genus), Pan'' (chimpanzees and bonobos), and in standard usage exclude the gen ...
.A ''hominin'' is a member of the tribe Hominini, a ''hominine'' is a member of the subfamily Homininae, a ''hominid'' is a member of the family Hominidae, and a ''hominoid'' is a member of the superfamily Hominoidea.


History of discoveries and classification

Until 1970, the family (and term) Hominidae meant humans only; the non-human
great ape The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
s were assigned to the then-family
Pongidae Pongidae , or the pongids is an obsolete primate taxon containing chimpanzees, gorillas and orangutans. By this definition pongids were also called "great apes". This taxon is not used today but is of historical significance. The great apes are ...
. Later discoveries led to revised classifications, with the great
ape Apes (collectively Hominoidea ) are a superfamily of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and Europe in prehistory, and counting humans are found global ...
s then united with humans (now in subfamily Homininae) as members of family Hominidae By 1990, it was recognized that
gorilla Gorillas are primarily herbivorous, terrestrial great apes that inhabit the tropical forests of equatorial Africa. The genus ''Gorilla'' is divided into two species: the eastern gorilla and the western gorilla, and either four or five su ...
s and
chimpanzee The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of Hominidae, great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close rel ...
s are more closely related to
human Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
s than they are to
orangutan Orangutans are great apes native to the rainforests of Indonesia and Malaysia. They are now found only in parts of Borneo and Sumatra, but during the Pleistocene they ranged throughout Southeast Asia and South China. Classified in the genus ...
s, leading to their (gorillas' and chimpanzees') placement in subfamily Homininae as well. The subfamily Homininae can be further subdivided into three branches, the tribe
Gorillini Gorillini is a taxonomic tribe containing three genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomencla ...
(gorillas), the tribe
Hominini The Hominini (hominins) form a Tribe (biology), taxonomic tribe of the subfamily Homininae (hominines). They comprise two extant genera: ''Homo'' (humans) and ''Pan (genus), Pan'' (chimpanzees and bonobos), and in standard usage exclude the gen ...
with subtribes
Panina Panina is a subtribe of tribe Hominini; it comprises all descendants of the human-chimpanzee last common ancestor (LCA) that are ''not'' of the branch of human lineage—that is, all those ancestors of the type genus '' Pan'' (chimpanzees and ...
(chimpanzees/bonobos) and
Hominina The australopithecines (), formally Australopithecina or Hominina, are generally any species in the related genera of ''Australopithecus'' and ''Paranthropus''. It may also include members of ''Kenyanthropus'', ''Ardipithecus'', and '' Praeanth ...
(humans and their extinct relatives), and the extinct tribe
Dryopithecini Dryopithecini is an extinct tribe of Eurasian and African great apes that are believed to be close to the ancestry of gorillas, chimpanzees and humans. Members of this tribe are known as dryopithecines. Taxonomy * Tribe Dryopithecini † ** '' ...
. The
Late Miocene The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million ye ...
fossil ''
Nakalipithecus nakayamai ''Nakalipithecus nakayamai'', sometimes referred to as the Nakali ape, is an extinct species of great ape from Nakali, Kenya from about 9.9–9.8 million years ago during the Late Miocene. It is known from a right jawbone with 3 molars and fro ...
'', described in 2007, is a basal member of this
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
, as is, perhaps, its contemporary ''
Ouranopithecus ''Ouranopithecus'' is a genus of extinct Eurasian great ape represented by two species, '' Ouranopithecus macedoniensis'', a late Miocene (9.6–8.7 mya) hominoid from Greece and '' Ouranopithecus turkae'', also from the late Miocene (8.7–7.4 ...
''; that is, they are not assignable to any of the three extant branches. Their existence suggests that the Homininae tribes diverged not earlier than about 8 million years ago (see
Human evolutionary genetics Human evolutionary genetics studies how one human genome differs from another human genome, the evolutionary past that gave rise to the human genome, and its current effects. Differences between genomes have anthropological, medical, historical and ...
). Today, chimpanzees and gorillas live in
tropical forest Tropical forests are forested ecoregions with tropical climates – that is, land areas approximately bounded by the Tropic of Cancer, tropics of Cancer and Tropic of Capricorn, Capricorn, but possibly affected by other factors such as prevailing ...
s with acid soils that rarely preserve fossils. Although no fossil gorillas have been reported, four chimpanzee teeth about 500,000 years old have been discovered in the East-African rift valley (
Kapthurin The Kapthurin Formation is a series of Middle Pleistocene sediments associated with the East African Rift Valley. Part of the East African Rift System, it is also an important archaeological site in the study of early humans who occupied ...
Formation, Kenya), where many fossils from the human lineage (hominins) have been found. This shows that some chimpanzees lived close to ''Homo'' ('' H. erectus'' or '' H. rhodesiensis'') at the time; the same is likely true for gorillas.


Taxonomic classification

Homininae * Tribe
Dryopithecini Dryopithecini is an extinct tribe of Eurasian and African great apes that are believed to be close to the ancestry of gorillas, chimpanzees and humans. Members of this tribe are known as dryopithecines. Taxonomy * Tribe Dryopithecini † ** '' ...
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
** ''
Kenyapithecus ''Kenyapithecus wickeri'' is a fossil ape discovered by Louis Leakey in 1961 at a site called Fort Ternan in Kenya. The upper jaw and teeth were dated to 14 million years ago. One theory states that ''Kenyapithecus'' may be the common ancest ...
(?)'' *** ''Kenyapitheus wickeri'' ** ''
Ouranopithecus ''Ouranopithecus'' is a genus of extinct Eurasian great ape represented by two species, '' Ouranopithecus macedoniensis'', a late Miocene (9.6–8.7 mya) hominoid from Greece and '' Ouranopithecus turkae'', also from the late Miocene (8.7–7.4 ...
'' *** ''Ouranopithecus macedoniensis'' ** '' Otavipithecus'' *** ''Otavipithecus namibiensis'' ** ''
Oreopithecus ''Oreopithecus'' (from the Greek , and , , meaning "hill-ape") is an extinct genus of ape from the Miocene epoch whose fossils have been found in today's Tuscany and Sardinia in Italy. It existed 9–7 million years ago in the Tusco-Sardinian a ...
(?)'' *** ''Oreopithecus bambolii'' ** ''
Nakalipithecus ''Nakalipithecus nakayamai'', sometimes referred to as the Nakali ape, is an extinct species of great ape from Nakali, Kenya from about 9.9–9.8 million years ago during the Late Miocene. It is known from a right jawbone with 3 molars and fro ...
'' *** ''Nakalipithecus nakayamai'' ** ''
Anoiapithecus ''Anoiapithecus'' is an extinct ape genus thought to be closely related to ''Dryopithecus''. Both genera lived during the Miocene, approximately 12 million years ago. Fossil specimens named by Salvador Moyà-Solà are known from the deposits f ...
'' *** ''Anoiapithecus brevirostris'' ** ''
Dryopithecus ''Dryopithecus'' is a genus of extinct great apes from the middle–late Miocene boundary of Europe 12.5 to 11.1 million years ago (mya). Since its discovery in 1856, the genus has been subject to taxonomic turmoil, with numerous new species b ...
'' *** ''Dryopithecus fontani'' ** ''
Hispanopithecus ''Hispanopithecus'' is a genus of apes that inhabited Europe during the Miocene epoch. It was first identified in a 1944 paper by J. F. Villalta and M. Crusafont in . Anthropologists disagree as to whether ''Hispanopithecus'' belongs to the subfa ...
(?)'' *** ''Hispanopithecus laietanus'' *** ''Hispanopithecus crusafonti'' ** '' Pierolapithecus'' *** ''Pierolapithecus catalaunicus'' ** ''
Rudapithecus ''Rudapithecus'' is a chimpanzee-likeLászló Kordos: 50 years of Rudapithecus
(in Hu ...
(?)'' *** ''Rudapithecus hungaricus'' ** ''
Samburupithecus ''Samburupithecus'' is an extinct primate that lived in Kenya during the middle to late Miocene. The one species in this genus, ''Samburupithecus kiptalami'', is known only from a maxilla fragment dated to 9.5 million years ago discovered in 198 ...
'' *** ''Samburupithecus kiptalami'' ** '' Danuvius'' *** ''Danuvius guggenmosi'' * Tribe
Gorillini Gorillini is a taxonomic tribe containing three genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomencla ...
** ''
Chororapithecus ''Chororapithecus'' is an extinct great ape from the Afar region of Ethiopia roughly 8 million years ago during the Late Miocene, comprising one species, ''C. abyssinicus''. It is known from 9 isolated teeth discovered in a 2005–2007 survey of ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
*** ''Chororapithecus abyssinicus'' ** Genus ''
Gorilla Gorillas are primarily herbivorous, terrestrial great apes that inhabit the tropical forests of equatorial Africa. The genus ''Gorilla'' is divided into two species: the eastern gorilla and the western gorilla, and either four or five su ...
'' ***
Western gorilla The western gorilla (''Gorilla gorilla'') is a great ape found in Africa, one of two species of the hominid genus ''Gorilla''. Large and robust with males weighing around , the species is found in a region of midwest Africa, geographically iso ...
, ''Gorilla gorilla'' ****
Western lowland gorilla The western lowland gorilla (''Gorilla gorilla gorilla'') is one of two Critically Endangered subspecies of the western gorilla (''Gorilla gorilla'') that lives in Montane ecosystems#Montane forests, montane, Old-growth forest, primary and sec ...
, ''Gorilla gorilla gorilla'' ****
Cross River gorilla The Cross River gorilla (''Gorilla gorilla diehli'') is a critically endangered subspecies of the western gorilla (''Gorilla gorilla''). It was named a new species in 1904 by Paul Matschie, a mammalian taxonomist working at the Humboldt Univers ...
, ''Gorilla gorilla diehli'' ***
Eastern gorilla The eastern gorilla (''Gorilla beringei'') is a critically endangered species of the genus ''Gorilla'' and the largest living primate. At present, the species is subdivided into two subspecies. There are 6,800 eastern lowland gorillas or Grauer ...
, ''Gorilla beringei'' ****
Mountain gorilla The mountain gorilla (''Gorilla beringei beringei'') is one of the two subspecies of the eastern gorilla. It is listed as endangered by the IUCN . There are two populations: One is found in the Virunga Mountains, Virunga volcanic mountains of C ...
, ''Gorilla beringei beringei'' ****
Eastern lowland gorilla The eastern lowland gorilla (''Gorilla beringei graueri'') or Grauer's gorilla is a Critically Endangered subspecies of eastern gorilla endemic to the mountainous forests of eastern Democratic Republic of the Congo. Important populations of th ...
, ''Gorilla beringei graueri'' * Tribe
Hominini The Hominini (hominins) form a Tribe (biology), taxonomic tribe of the subfamily Homininae (hominines). They comprise two extant genera: ''Homo'' (humans) and ''Pan (genus), Pan'' (chimpanzees and bonobos), and in standard usage exclude the gen ...
** Subtribe
Panina Panina is a subtribe of tribe Hominini; it comprises all descendants of the human-chimpanzee last common ancestor (LCA) that are ''not'' of the branch of human lineage—that is, all those ancestors of the type genus '' Pan'' (chimpanzees and ...
*** Genus '' Pan'' ****
Chimpanzee The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of Hominidae, great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close rel ...
(common chimpanzee), ''Pan troglodytes'' ***** Central chimpanzee, ''Pan troglodytes troglodytes'' *****
Western chimpanzee The western chimpanzee or West African chimpanzee (''Pan troglodytes verus'') is a Critically Endangered subspecies of the common chimpanzee. It inhabits western Africa, specifically Côte d'Ivoire, Guinea, Liberia, Mali, Senegal, Ghana, Guinea-B ...
, ''Pan troglodytes verus'' *****
Nigeria-Cameroon chimpanzee The Nigeria-Cameroon chimpanzee (''Pan troglodytes ellioti'') is a subspecies of the common chimpanzee which inhabits the rainforest along the border of Nigeria and Cameroon. Male Nigeria-Cameroon chimpanzees can weigh up to 70 kilos with a body ...
, ''Pan troglodytes ellioti'' *****
Eastern chimpanzee The eastern chimpanzee (''Pan troglodytes schweinfurthii'') is a subspecies of the common chimpanzee. It is native to the Central African Republic, South Sudan, the Democratic Republic of the Congo, Uganda, Rwanda, Burundi, and Tanzania. Taxono ...
, ''Pan troglodytes schweinfurthii'' ****
Bonobo The bonobo (; ''Pan paniscus''), also historically called the pygmy chimpanzee (less often the dwarf chimpanzee or gracile chimpanzee), is an endangered great ape and one of the two species making up the genus ''Pan (genus), Pan'' (the other bei ...
(pygmy chimpanzee), ''Pan paniscus'' ** Subtribe
Hominina The australopithecines (), formally Australopithecina or Hominina, are generally any species in the related genera of ''Australopithecus'' and ''Paranthropus''. It may also include members of ''Kenyanthropus'', ''Ardipithecus'', and '' Praeanth ...
*** ''
Graecopithecus ''Graecopithecus'' is an extinct genus of hominid that lived in southeast Europe during the late Miocene around 7.2 million years ago. Originally identified by a single lower jawbone bearing teeth found in Pyrgos Vasilissis, Athens, Greece, in ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''Graecopithecus freybergi''. Note: ''Graecopithecus'' has also been subsumed by other authors into ''Dryopithecus''. The placement of ''Graecopithecus'' within the Hominina, as shown here, represents a hypothesis, but not scientific consensus. *** ''
Sahelanthropus ''Sahelanthropus'' is an extinct genus of hominid dated to about during the Late Miocene. The type species, ''Sahelanthropus tchadensis'', was first announced in 2002, based mainly on a partial cranium, nicknamed ''Toumaï'', discovered in north ...
(?)''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''Sahelanthropus tchadensis'' *** ''
Orrorin ''Orrorin'' is an extinct genus of primate within Homininae from the Miocene Lukeino Formation and Pliocene Mabaget Formation, both of Kenya. The type species is ''O. tugenenesis'', named in 2001, and a second species, ''O. praegens'', assigne ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''Orrorin tugenensis'' **** ''Orrorin praegens'' *** ''
Ardipithecus ''Ardipithecus'' is a genus of an extinct hominine that lived during the Late Miocene and Early Pliocene epochs in the Afar Depression, Ethiopia. Originally described as one of the earliest ancestors of humans after they diverged from the chim ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Ardipithecus ramidus ''Ardipithecus ramidus'' is a species of australopithecine from the Afar region of Early Pliocene Ethiopia 4.4 million years ago (mya). The species ''A. ramidus'' is the type species for the genus ''Ardipithecus''. There is an older species in t ...
'' **** ''
Ardipithecus kadabba ''Ardipithecus kadabba'' is the scientific classification given to fossil remains "known only from teeth and bits and pieces of skeletal bones", originally estimated to be 5.8 to 5.2 million years old, and later revised to 5.77 to 5.54 million ye ...
'' *** ''
Kenyanthropus ''Kenyanthropus'' is a genus of extinct hominin identified from the Lomekwi site by Lake Turkana, Kenya, dated to 3.3 to 3.2 million years ago during the Middle Pliocene. It contains one species, ''K. platyops'', but may also include the 2 mi ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''Kenyanthropus platyops'' *** ''
Australopithecus ''Australopithecus'' (, ; or (, ) is a genus of early hominins that existed in Africa during the Pliocene and Early Pleistocene. The genera ''Homo'' (which includes modern humans), ''Paranthropus'', and ''Kenyanthropus'' evolved from some ''Aus ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Australopithecus bahrelghazali ''Australopithecus bahrelghazali'' is an extinct species of australopithecine discovered in 1995 at Koro Toro, Bahr el Gazel, Chad, existing around 3.5 million years ago in the Pliocene. It is the first and only australopithecine known from Ce ...
'' **** ''
Australopithecus anamensis ''Australopithecus anamensis'' is a hominin species that lived roughly between 4.3 and 3.8 million years ago, and is the oldest known ''Australopithecus'' species, Nearly 100 fossil specimens of ''A. anamensis'' are known from Kenya and Ethiopia ...
'' **** ''
Australopithecus afarensis ''Australopithecus afarensis'' is an extinct species of australopithecine which lived from about 3.9–2.9 million years ago (mya) in the Pliocene of East Africa. The first fossils were discovered in the 1930s, but major fossil finds would not ta ...
'' **** ''
Australopithecus africanus ''Australopithecus africanus'' is an extinct species of australopithecine which lived between about 3.3 and 2.1 million years ago in the Late Pliocene to Early Pleistocene of South Africa. The species has been recovered from Taung, Sterkfontei ...
'' **** ''
Australopithecus garhi ''Australopithecus garhi'' is a species of australopithecine from the Bouri Formation in the Afar Region of Ethiopia 2.6–2.5 million years ago (mya) during the Early Pleistocene. The first remains were described in 1999 based on several skele ...
'' **** ''
Australopithecus sediba ''Australopithecus sediba'' is an extinct species of australopithecine recovered from Malapa Fossil Site, Cradle of Humankind, Malapa Cave, Cradle of Humankind, South Africa. It is known from a partial juvenile skeleton, the holotype MH1, and a ...
'' *** ''
Paranthropus ''Paranthropus'' is a genus of extinct hominin which contains two widely accepted species: ''Paranthropus robustus, P. robustus'' and ''P. boisei''. However, the validity of ''Paranthropus'' is contested, and it is sometimes considered to be sy ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Paranthropus aethiopicus ''Paranthropus aethiopicus'' is an Extinction, extinct species of robust australopithecine from the Late Pliocene to Early Pleistocene of East Africa about 2.7–2.3 million years ago. However, it is much debated whether or not ''Paranthropus'' ...
'' **** ''
Paranthropus robustus ''Paranthropus robustus'' is a species of robust australopithecine from the Early and possibly Middle Pleistocene of the Cradle of Humankind, South Africa, about 2.27 to 0.87 (or, more conservatively, 2 to 1) million years ago. It has been ide ...
'' **** ''
Paranthropus boisei ''Paranthropus boisei'' is a species of australopithecine from the Early Pleistocene of East Africa about 2.5 to 1.15 million years ago. The holotype specimen, OH 5, was discovered by palaeoanthropologist Mary Leakey in 1959 at Olduvai Gorge, Ta ...
'' *** ''
Homo ''Homo'' () is a genus of great ape (family Hominidae) that emerged from the genus ''Australopithecus'' and encompasses only a single extant species, ''Homo sapiens'' (modern humans), along with a number of extinct species (collectively called ...
'' – immediate ancestors of modern
human Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
s **** '' Homo gautengensis''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
(probable ''H. habilis'' specimens) **** ''
Homo rudolfensis ''Homo rudolfensis'' is an extinct species of archaic human from the Early Pleistocene of East Africa about 2 million years ago (mya). Because ''H. rudolfensis'' coexisted with several other hominins, it is debated what specimens can be confiden ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Homo habilis ''Homo habilis'' ( 'handy man') is an extinct species of archaic human from the Early Pleistocene of East and South Africa about 2.4 million years ago to 1.65 million years ago ( mya). Upon species description in 1964, ''H. habilis'' was highly ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Homo floresiensis ''Homo floresiensis'' , also known as "Flores Man" or "Hobbit" (after Hobbit, the fictional species), is an Extinction, extinct species of small archaic humans that inhabited the island of Flores, Indonesia, until the arrival of Homo sapiens, ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Homo erectus ''Homo erectus'' ( ) is an extinction, extinct species of Homo, archaic human from the Pleistocene, spanning nearly 2 million years. It is the first human species to evolve a humanlike body plan and human gait, gait, to early expansions of h ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Homo ergaster ''Homo ergaster'' is an extinct species or subspecies of archaic humans who lived in Africa in the Early Pleistocene. Whether ''H. ergaster'' constitutes a species of its own or should be subsumed into '' H. erectus'' is an ongoing and unresol ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Homo antecessor ''Homo antecessor'' (Latin "pioneer man") is an extinct species of archaic human recorded in the Spanish Archaeological Site of Atapuerca, Sierra de Atapuerca, a productive archaeological site, from 1.2 to 0.8 million years ago during the Early ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Homo heidelbergensis ''Homo heidelbergensis'' is a species of archaic human from the Middle Pleistocene of Europe and Africa, as well as potentially Asia depending on the taxonomic convention used. The species-level classification of ''Homo'' during the Middle Pleis ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Homo cepranensis ''Homo'' () is a genus of great ape (family Hominidae) that emerged from the genus ''Australopithecus'' and encompasses only a single extant species, ''Homo sapiens'' (modern humans), along with a number of extinct species (collectively called ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
(probable early ''H. sapiens'' specimens) ****
Denisovans The Denisovans or Denisova hominins ( ) are an extinct species or subspecies of archaic human that ranged across Asia during the Lower Paleolithic, Lower and Middle Paleolithic, and lived, based on current evidence, from 285 thousand to 25 thou ...
(scientific name has not yet been assigned)
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Homo neanderthalensis Neanderthals ( ; ''Homo neanderthalensis'' or sometimes ''H. sapiens neanderthalensis'') are an extinct group of archaic humans who inhabited Europe and Western and Central Asia during the Middle to Late Pleistocene. Neanderthal extinctio ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
**** ''
Homo rhodesiensis ''Homo rhodesiensis'' is the species name proposed by Arthur Smith Woodward (1921) to classify Kabwe 1 (the "Kabwe skull" or "Broken Hill skull", also "Rhodesian Man"), a Middle Stone Age fossil recovered from Broken Hill mine in Kabwe, Northe ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
(probable late ''H. heidelbergensis'' specimens) **** ''
Homo sapiens Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
'' *****
Anatomically modern human Anatomy () is the branch of morphology concerned with the study of the internal structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science ...
, ''Homo sapiens sapiens'' ****** '' Archaic Homo sapiens'' (
Cro-Magnon Cro-Magnons or European early modern humans (EEMH) were the first early modern humans (''Homo sapiens'') to settle in Europe, migrating from western Asia, continuously occupying the continent possibly from as early as 56,800 years ago. They in ...
)
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
****** Red Deer Cave people
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
(scientific name has not yet been assigned) ***** ''
Homo sapiens idaltu Herto Man refers to human remains (''Homo sapiens'') discovered in 1997 from the Upper Herto member (geology), member of the Bouri Formation in the Afar Triangle, Ethiopia. The remains have been dated as between 154,000 and 160,000 years old. The ...
''
† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...
(classification not widely accepted)


Evolution

The age of the subfamily Homininae (of the Homininae–Ponginae last common ancestor) is estimated at some 14 to 12.5 million years (''
Sivapithecus ''Sivapithecus'' () (syn: ''Ramapithecus)'' is a genus of extinct apes. Fossil remains of animals now assigned to this genus, dated from 12.2 million years old in the Miocene, have been found since the 19th century in the Sivalik Hills of the Ind ...
''). Its separation into Gorillini and Hominini (the "gorilla–human last common ancestor", GHLCA) is estimated to have occurred at about (TGHLCA) during the late
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
, close to the age of ''
Nakalipithecus nakayamai ''Nakalipithecus nakayamai'', sometimes referred to as the Nakali ape, is an extinct species of great ape from Nakali, Kenya from about 9.9–9.8 million years ago during the Late Miocene. It is known from a right jawbone with 3 molars and fro ...
''. There is evidence there was interbreeding of Gorillas and the Pan–Homo ancestors until right up to the Pan–Homo split.


Evolution of bipedalism

Recent studies of ''
Ardipithecus ramidus ''Ardipithecus ramidus'' is a species of australopithecine from the Afar region of Early Pliocene Ethiopia 4.4 million years ago (mya). The species ''A. ramidus'' is the type species for the genus ''Ardipithecus''. There is an older species in t ...
'' (4.4 million years old) and ''
Orrorin tugenensis ''Orrorin'' is an extinct genus of primate within Homininae from the Miocene Lukeino Formation and Pliocene Mabaget Formation, both of Kenya. The type species is ''O. tugenenesis'', named in 2001, and a second species, ''O. praegens'', assigne ...
'' (6 million years old) suggest some degree of bipedalism. ''
Australopithecus ''Australopithecus'' (, ; or (, ) is a genus of early hominins that existed in Africa during the Pliocene and Early Pleistocene. The genera ''Homo'' (which includes modern humans), ''Paranthropus'', and ''Kenyanthropus'' evolved from some ''Aus ...
'' and early ''
Paranthropus ''Paranthropus'' is a genus of extinct hominin which contains two widely accepted species: ''Paranthropus robustus, P. robustus'' and ''P. boisei''. However, the validity of ''Paranthropus'' is contested, and it is sometimes considered to be sy ...
'' may have been
bipedal Bipedalism is a form of terrestrial locomotion where an animal moves by means of its two rear (or lower) limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' ...
. Very early hominins such as ''
Ardipithecus ramidus ''Ardipithecus ramidus'' is a species of australopithecine from the Afar region of Early Pliocene Ethiopia 4.4 million years ago (mya). The species ''A. ramidus'' is the type species for the genus ''Ardipithecus''. There is an older species in t ...
'' may have possessed an arboreal type of bipedalism. The evolution of
bipedalism Bipedalism is a form of terrestrial locomotion where an animal moves by means of its two rear (or lower) Limb (anatomy), limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from ...
encouraged multiple changes among hominins especially when it came to
bipedalism Bipedalism is a form of terrestrial locomotion where an animal moves by means of its two rear (or lower) Limb (anatomy), limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from ...
in humans as they were now able to do many other things as they began to walk with their feet. These changes included the ability to now use their hands to create tools or carry things with their hands, the ability to travel longer distances at a faster speed, and the ability to hunt for food. According to researchers, humans were able to be bipedalists due to Darwin's Principle of
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the Heredity, heritable traits characteristic of a population over generation ...
. Darwin himself believed that larger brains in humans made an upright gait necessary, but had no hypothesis for how the mechanism evolved. The first major theory attempting to directly explanation the origins of
bipedalism Bipedalism is a form of terrestrial locomotion where an animal moves by means of its two rear (or lower) Limb (anatomy), limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from ...
was the
Savannah hypothesis The savannah hypothesis (or savanna hypothesis) is a hypothesis that human bipedalism evolved as a direct result of human ancestors' transition from an arboreal lifestyle to one on the savannas. According to the hypothesis, hominins left the woo ...
(Dart 1925.) This theory hypothesized that hominins became bipedalists due to the environment of the Savanna such as the tall grass and dry climate. This was later proven to be incorrect due to fossil records that showed that hominins were still climbing trees during this era. Anthropologist
Owen Lovejoy Owen Lovejoy (January 6, 1811 – March 25, 1864) was an American lawyer, Congregational minister, Abolitionism in the United States, abolitionist, and Republican United States Congress, congressman from Illinois. He was also a "conductor ...
has suggested that
bipedalism Bipedalism is a form of terrestrial locomotion where an animal moves by means of its two rear (or lower) Limb (anatomy), limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from ...
was a result of
sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
in efforts to help with the collecting of food. In his Male Provisioning Hypothesis introduced in 1981, lowered birth rates in early hominids increased pressure on males to provide for females and offspring. While females groomed and cared for their children with the family group, males ranged to seek food and returned bipadally with full arms. Males who could better provide for females in this model were more likely to mate and produce offspring. Anthropologist Yohannes Haile-Selassie, an expert on ''
Australopithecus anamensis ''Australopithecus anamensis'' is a hominin species that lived roughly between 4.3 and 3.8 million years ago, and is the oldest known ''Australopithecus'' species, Nearly 100 fossil specimens of ''A. anamensis'' are known from Kenya and Ethiopia ...
'', discusses the evidence that Australopithecus were one of the first hominins to evolve into obligate bipedalists. The remains of this subfamily are very important in the field of research as it presents possible information regarding how these primates adapted from tree life to terrestrial life. This was a huge adaptation as it encouraged many evolutionary changes within hominins including the ability to use their hand to make tools and gather food, as well as a larger brain development due to their change in diet.


Brain size evolution

There has been a gradual increase in brain volume (
brain size The size of the brain is a frequent topic of study within the fields of anatomy, biological anthropology, animal science and evolution. Measuring brain size and cranial capacity is relevant both to humans and other animals, and can be done by wei ...
) as the ancestors of modern humans progressed along the
timeline of human evolution The timeline of human evolution outlines the major events in the evolutionary lineage of the modern human species, ''Homo sapiens'', throughout the history of life, beginning some 4 billion years ago down to recent evolution within ''H. sapiens ...
, starting from about 600 cm3 in ''
Homo habilis ''Homo habilis'' ( 'handy man') is an extinct species of archaic human from the Early Pleistocene of East and South Africa about 2.4 million years ago to 1.65 million years ago ( mya). Upon species description in 1964, ''H. habilis'' was highly ...
'' up to 1500 cm3 in ''
Homo neanderthalensis Neanderthals ( ; ''Homo neanderthalensis'' or sometimes ''H. sapiens neanderthalensis'') are an extinct group of archaic humans who inhabited Europe and Western and Central Asia during the Middle to Late Pleistocene. Neanderthal extinctio ...
''. However, modern ''Homo sapiens'' have a brain volume slightly smaller (1250 cm3) than Neanderthals, women have a brain slightly smaller than men and the Flores hominids (''
Homo floresiensis ''Homo floresiensis'' , also known as "Flores Man" or "Hobbit" (after Hobbit, the fictional species), is an Extinction, extinct species of small archaic humans that inhabited the island of Flores, Indonesia, until the arrival of Homo sapiens, ...
''), nicknamed hobbits, had a cranial capacity of about 380 cm3 (considered small for a chimpanzee), about a third of the ''
Homo erectus ''Homo erectus'' ( ) is an extinction, extinct species of Homo, archaic human from the Pleistocene, spanning nearly 2 million years. It is the first human species to evolve a humanlike body plan and human gait, gait, to early expansions of h ...
'' average. It is proposed that they evolved from ''H. erectus'' as a case of insular dwarfism. In spite of their smaller brain, there is evidence that ''H. floresiensis'' used fire and made stone tools at least as sophisticated as those of their proposed ancestors ''H. erectus''. In this case, it seems that for intelligence, the structure of the brain is more important than its size. The current size of the human brain is a big distinguishing factor that separates humans from other primates. Recent examination of the human brain shows that the brain of a human is about more than four times the size of
great ape The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
s and 20 times larger than the brain size of
old world monkeys Old World monkeys are primates in the family (biology), family Cercopithecidae (). Twenty-four genus, genera and 138 species are recognized, making it the largest primate family. Old World monkey genera include baboons (genus ''Papio''), red colo ...
. A study was conducted to help determine the evolution of the brain size within the sub family Homininae that tested the genes ASPM ( abnormal spindle-like microcephaly associated) and MCHP1 (microcephalin-1) and their association with the human brain. In this study researchers discovered that the increase in brain size is correlated to the increase of both ASP and MCPH1. MCPH1 is very polymorphic in humans compared to
gibbon Gibbons () are apes in the family Hylobatidae (). The family historically contained one genus, but now is split into four extant genera and 20 species. Gibbons live in subtropical and tropical forests from eastern Bangladesh and Northeast Indi ...
s,
Old World monkey Old World monkeys are primates in the family Cercopithecidae (). Twenty-four genera and 138 species are recognized, making it the largest primate family. Old World monkey genera include baboons (genus '' Papio''), red colobus (genus '' Piliocolob ...
s. This gene helps encourage the growth of the brain. Further research indicated that the MCPH1 gene in humans could have also been an encouraging factor of population expansion. Other researchers have included that the diet was an encouraging factor to brain size as protein intake increased this helped brain development.


Evolution of family structure and sexuality

Sexuality is related to family structure and partly shapes it. The involvement of fathers in education is quite unique to humans, at least when compared to other Homininae.
Concealed ovulation Concealed ovulation or hidden estrus is the lack of any perceptible change (e.g., a change in appearance or scent) when an adult female is fertile and near ovulation. Some examples of perceptible changes are swelling and redness of the vulva in ba ...
and
menopause Menopause, also known as the climacteric, is the time when Menstruation, menstrual periods permanently stop, marking the end of the Human reproduction, reproductive stage for the female human. It typically occurs between the ages of 45 and 5 ...
in women both also occur in a few other primates however, but are uncommon in other species.
Testis A testicle or testis ( testes) is the gonad in all male bilaterians, including humans, and is Homology (biology), homologous to the ovary in females. Its primary functions are the production of sperm and the secretion of Androgen, androgens, p ...
and penis size seems to be related to family structure:
monogamy Monogamy ( ) is a social relation, relationship of Dyad (sociology), two individuals in which they form a mutual and exclusive intimate Significant other, partnership. Having only one partner at any one time, whether for life or #Serial monogamy ...
or
promiscuity Promiscuity is the practice of engaging in sexual activity frequently with different partners or being indiscriminate in the choice of sexual partners. The term can carry a moral judgment. A common example of behavior viewed as promiscuous by man ...
, or
harem A harem is a domestic space that is reserved for the women of the house in a Muslim family. A harem may house a man's wife or wives, their pre-pubescent male children, unmarried daughters, female domestic Domestic worker, servants, and other un ...
, in humans, chimpanzees or gorillas, respectively. The levels of
sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
are generally seen as a marker of
sexual selection Sexual selection is a mechanism of evolution in which members of one sex mate choice, choose mates of the other sex to mating, mate with (intersexual selection), and compete with members of the same sex for access to members of the opposite sex ...
. Studies have suggested that the earliest hominins were dimorphic and that this lessened over the course of the evolution of the genus ''Homo'', correlating with humans becoming more monogamous, whereas gorillas, who live in harems, show a large degree of sexual dimorphism. Concealed (or "hidden") ovulation means that the phase of fertility is not detectable in women, whereas chimpanzees advertise ovulation via an obvious swelling of the genitals. Women can be partly aware of their ovulation along the menstrual phases, but men are essentially unable to detect ovulation in women. Most primates have semi-concealed ovulation, thus one can think that the common ancestor had semi-concealed ovulation, that was inherited by gorillas, and that later evolved in concealed ovulation in humans and advertised ovulation in chimpanzees. Menopause also occurs in rhesus monkeys, and possibly in chimpanzees, but does not in gorillas and is quite uncommon in other primates (and other mammal groups).


See also

* * * * *


Notes


References


Further reading

* * * * * * * *


External links


Human Timeline (Interactive)
– Smithsonian,
National Museum of Natural History The National Museum of Natural History (NMNH) is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. With 4.4 ...
(August 2016). {{Taxonbar, from=Q242047 Hominidae Extant Miocene first appearances Mammal subfamilies Taxa named by John Edward Gray Taxa described in 1825