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Hominina
Australopithecina or Hominina is a subtribe in the tribe Hominini. The members of the subtribe are generally ''Australopithecus'' (cladistically including the genera ''Homo'', ''Paranthropus'', and ''Kenyanthropus''), and it typically includes the earlier ''Ardipithecus'', ''Orrorin'', ''Sahelanthropus'', and ''Graecopithecus''. All these closely related species are now sometimes collectively termed australopiths or homininians. They are the extinct, close relatives of humans and, with the extant genus ''Homo'', comprise the human clade. Members of the human clade, i.e. the Hominini after the split from the chimpanzees, are now called Hominina (''see Hominidae; terms "hominids" and hominins''). While none of the groups normally directly assigned to this group survived, the australopiths do not appear to be literally extinct (in the sense of having no living descendants) as the genera ''Kenyanthropus'', ''Paranthropus'' and ''Homo'' probably emerged as sister of a late ''Austral ...
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Hominidae
The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the eastern and western gorilla); '' Pan'' (the chimpanzee and the bonobo); and ''Homo'', of which only modern humans remain. Several revisions in classifying the great apes have caused the use of the term ''"hominid"'' to vary over time. The original meaning of "hominid" referred only to humans (''Homo'') and their closest extinct relatives. However, by the 1990s humans, apes, and their ancestors were considered to be "hominids". The earlier restrictive meaning has now been largely assumed by the term ''"hominin"'', which comprises all members of the human clade after the split from the chimpanzees (''Pan''). The current meaning of "hominid" includes all the great apes including humans. Usage still varies, however, and some scientists and la ...
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Hominini
The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos) and in standard usage excludes the genus ''Gorilla'' (gorillas). The term was originally introduced by Camille Arambourg (1948). Arambourg combined the categories of ''Hominina'' and ''Simiina'' due to Gray (1825) into his new subtribe. Traditionally, chimpanzees, gorillas and orangutans were grouped together as pongids. Since Gray's classification, evidence has accumulated from genetic phylogeny confirming that humans, chimpanzees, and gorillas are more closely related to each other than to the orangutan. The former pongids were reassigned to the subfamily Hominidae ("great apes"), which already included humans, but the details of this reassignment remain contested; within Hominini, not every source excludes gorillas, and not every source includes chimpanzees. Humans are the only extant species in the ...
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Sahelanthropus
''Sahelanthropus tchadensis'' is an extinct species of the Homininae (African apes) dated to about , during the Miocene epoch. The species, and its genus ''Sahelanthropus'', was announced in 2002, based mainly on a partial cranium, nicknamed ''Toumaï'', discovered in northern Chad. ''Sahelanthropus tchadensis'' lived close to the time of the chimpanzee–human divergence, possibly related to ''Orrorin'', a species of Homininae that lived about one million years later. It may have been ancestral to both humans and chimpanzees (which would place it in the tribe Hominini), or alternatively an early member of the tribe Gorillini. In 2020, the femur was analyzed, and it was found that ''Sahelanthropus'' was not bipedal, casting some doubt on its position as a human ancestor, but this was refuted in 2022. Taxonomy Discovery Funded by the Mission Paléoanthropologique Franco–Tchadienne, Ahounta Djimdoumalbaye, Fanoné Gongdibé, Mahamat Adoum, and Alain Beauvilain identified the f ...
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Homo
''Homo'' () is the genus that emerged in the (otherwise extinct) genus ''Australopithecus'' that encompasses the extant species ''Homo sapiens'' ( modern humans), plus several extinct species classified as either ancestral to or closely related to modern humans (depending on the species), most notably ''Homo erectus'' and ''Homo neanderthalensis''. The genus emerged with the appearance of '' Homo habilis'' just over 2 million years ago. ''Homo'', together with the genus '' Paranthropus'', is probably sister to ''Australopithecus africanus'', which itself had previously split from the lineage of '' Pan'', the chimpanzees. ''Homo erectus'' appeared about 2 million years ago and, in several early migrations, spread throughout Africa (where it is dubbed ''Homo ergaster'') and Eurasia. It was likely that the first human species lived in a hunter-gatherer society and was able to control fire. An adaptive and successful species, ''Homo erectus'' persisted for more than a million ye ...
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Orrorin
''Orrorin tugenensis'' is a postulated early species of Homininae, estimated at and discovered in 2000. It is not confirmed how ''Orrorin'' is related to modern humans. Its discovery was used to argue against the hypothesis that australopithecines are human ancestors, although this remains the most prevalent hypothesis of human evolution as of 2012. The name of genus ''Orrorin'' (plural ''Orroriek'') means "original man" in Tugen, and the name of the only classified species, ''O. tugenensis'', derives from Tugen Hills in Kenya, where the first fossil was found in 2000. As of 2007, 20 fossils of the species have been found. Fossils The 20 specimens found as of 2007 are believed to be from at least five individuals. They include: the posterior part of a mandible in two pieces; a symphysis and several isolated teeth; three fragments of femora; a partial humerus; a proximal phalanx; and a distal thumb phalanx. ''Orrorin'' had small teeth relative to its body size. Its dentition d ...
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Orrorin Tugenensis
''Orrorin tugenensis'' is a postulated early species of Homininae, estimated at and discovered in 2000. It is not confirmed how ''Orrorin'' is related to modern humans. Its discovery was used to argue against the hypothesis that australopithecines are human ancestors, although this remains the most prevalent hypothesis of human evolution as of 2012. The name of genus ''Orrorin'' (plural ''Orroriek'') means "original man" in Tugen, and the name of the only classified species, ''O. tugenensis'', derives from Tugen Hills in Kenya, where the first fossil was found in 2000. As of 2007, 20 fossils of the species have been found. Fossils The 20 specimens found as of 2007 are believed to be from at least five individuals. They include: the posterior part of a mandible in two pieces; a symphysis and several isolated teeth; three fragments of femora; a partial humerus; a proximal phalanx; and a distal thumb phalanx. ''Orrorin'' had small teeth relative to its body size. Its dentition di ...
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Paranthropus Robustus
''Paranthropus robustus'' is a species of robustness (morphology), robust australopithecine from the Early Pleistocene, Early and possibly Middle Pleistocene, Middle Pleistocene of the Cradle of Humankind, South Africa, about 2.27 to 0.87 (or, more conservatively, 2 to 1) million years ago. It has been identified in Kromdraai, Swartkrans, Sterkfontein, Gondolin Cave, Gondolin, Cooper's Cave, Cooper's, and Drimolen Caves. Discovered in 1938, it was among the first early hominins described, and became the type species for the genus ''Paranthropus''. However, it has been argued by some that ''Paranthropus'' is an invalid grouping and synonym (taxonomy), synonymous with ''Australopithecus'', so the species is also often classified as ''Australopithecus robustus''. Robust australopithecines—as opposed to gracility, gracile australopithecines—are characterised by heavily built skulls capable of producing high stress (physics), stresses and bite forces, as well as inflated cheek teet ...
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Paranthropus Boisei
''Paranthropus boisei'' is a species of australopithecine from the Early Pleistocene of East Africa about 2.5 to 1.15 million years ago. The holotype specimen, OH 5, was discovered by palaeoanthropologist Mary Leakey in 1959, and described by her husband Louis a month later. It was originally placed into its own genus as "''Zinjanthropus boisei''", but is now relegated to ''Paranthropus'' along with other robust australopithecines. However, it is also argued that ''Paranthropus'' is an invalid grouping and synonymous with ''Australopithecus'', so the species is also often classified as ''Australopithecus boisei''. Robust australopithecines are characterised by heavily built skulls capable of producing high stresses and bite forces, and some of the largest molars with the thickest enamel of any known ape. ''P. boisei'' is the most robust of this group. Brain size was about , similar to other australopithecines. Some skulls are markedly smaller than others, which is taken as evi ...
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Australopithecus Bahrelghazali
''Australopithecus bahrelghazali'' is an extinct species of australopithecine discovered in 1995 at Koro Toro, Bahr el Gazel, Chad, existing around 3.5 million years ago in the Pliocene. It is the first and only australopithecine known from Central Africa, and demonstrates that this group was widely distributed across Africa as opposed to being restricted to East and southern Africa as previously thought. The validity of ''A. bahrelghazali'' has not been widely accepted, in favour of classifying the specimens as ''A. afarensis'', a better known Pliocene australopithecine from East Africa, because of the anatomical similarity and the fact that ''A. bahrelghazali'' is known only from 3 partial jawbones and an isolated premolar. The specimens inhabited a lakeside grassland environment with sparse tree cover, possibly similar to the modern Okavango Delta, and similarly predominantly ate C4 savanna foods—such as grasses, sedges, storage organs, or rhizomes—and to a lesser degree ...
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Encephalization
Encephalization quotient (EQ), encephalization level (EL), or just encephalization is a relative brain size measure that is defined as the ratio between observed to predicted brain mass for an animal of a given size, based on nonlinear regression on a range of reference species. It has been used as a proxy for intelligence and thus as a possible way of comparing the intelligences of different species. For this purpose it is a more refined measurement than the raw brain-to-body mass ratio, as it takes into account allometric effects. Expressed as a formula, the relationship has been developed for mammals and may not yield relevant results when applied outside this group. Perspective on intelligence measures Encephalization quotient was developed in an attempt to provide a way of correlating an animal's physical characteristics with perceived intelligence. It improved on the previous attempt, brain-to-body mass ratio, so it has persisted. Subsequent work, notably Roth, found ...
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Australopithecus Afarensis
''Australopithecus afarensis'' is an extinct species of australopithecine which lived from about 3.9–2.9 million years ago (mya) in the Pliocene of East Africa. The first fossils were discovered in the 1930s, but major fossil finds would not take place until the 1970s. From 1972 to 1977, the International Afar Research Expedition—led by anthropologists Maurice Taieb, Donald Johanson and Yves Coppens—unearthed several hundreds of hominin specimens in Hadar, Ethiopia, the most significant being the exceedingly well-preserved skeleton AL 288-1 ("Lucy") and the site AL 333 ("the First Family"). Beginning in 1974, Mary Leakey led an expedition into Laetoli, Tanzania, and notably recovered fossil trackways. In 1978, the species was first described, but this was followed by arguments for splitting the wealth of specimens into different species given the wide range of variation which had been attributed to sexual dimorphism (normal differences between males and females). ''A. afa ...
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Paranthropus Aethiopicus
''Paranthropus aethiopicus'' is an extinct species of robust australopithecine from the Late Pliocene to Early Pleistocene of East Africa about 2.7–2.3 million years ago. However, it is much debated whether or not ''Paranthropus'' is an invalid grouping and is synonymous with ''Australopithecus'', so the species is also often classified as ''Australopithecus aethiopicus''. Whatever the case, it is considered to have been the ancestor of the much more robust '' P. boisei''. It is debated if ''P. aethiopicus'' should be subsumed under ''P. boisei'', and the terms ''P. boisei'' sensu lato ("in the broad sense") and ''P. boisei'' sensu stricto ("in the strict sense") can be used to respectively include and exclude ''P. aethiopicus'' from ''P. boisei''. Like other ''Paranthropus'', ''P. aethiopicus'' had a tall face, thick palate, and especially enlarged cheek teeth. However, likely due to its archaicness, it also diverges from other ''Paranthropus'', with some aspects resembling th ...
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