sensory nervous system
The sensory nervous system is a part of the nervous system responsible for processing sensory information. A sensory system consists of sensory neurons (including the sensory receptor cells), neural pathways, and parts of the brain involved i ...
, that reacts to
visible light
Light or visible light is electromagnetic radiation that can be perceived by the human eye. Visible light is usually defined as having wavelengths in the range of 400–700 nanometres (nm), corresponding to frequencies of 750–420 te ...
and allows humans to use visual information for various purposes including seeing things, keeping balance, and maintaining circadian rhythm.
The eye can be considered as a living
optical device
Optics is the branch of physics that studies the behaviour and properties of light, including its interactions with matter and the construction of instruments that use or detect it. Optics usually describes the behaviour of visible, ultravio ...
. It is approximately spherical in shape, with its outer layers, such as the outermost, white part of the eye (the sclera) and one of its inner layers (the pigmented choroid) keeping the eye essentially light tight except on the eye's
optic axis
An optical axis is a line along which there is some degree of rotational symmetry in an optical system such as a camera lens, microscope or telescopic sight.
The optical axis is an imaginary line that defines the path along which light propagat ...
. In order, along the optic axis, the optical components consist of a first
lens
A lens is a transmissive optical device which focuses or disperses a light beam by means of refraction. A simple lens consists of a single piece of transparent material, while a compound lens consists of several simple lenses (''elements ...
aperture
In optics, an aperture is a hole or an opening through which light travels. More specifically, the aperture and focal length of an optical system determine the cone angle of a bundle of rays that come to a focus in the image plane.
An ...
(the
pupil
The pupil is a black hole located in the center of the Iris (anatomy), iris of the Human eye, eye that allows light to strike the retina.Cassin, B. and Solomon, S. (1990) ''Dictionary of Eye Terminology''. Gainesville, Florida: Triad Publishing ...
crystalline lens
The lens, or crystalline lens, is a transparent biconvex structure in the eye that, along with the cornea, helps to refract light to be focused on the retina. By changing shape, it functions to change the focal length of the eye so that it ca ...
) that accomplishes the remaining focussing of light into
images
An image is a visual representation of something. It can be two-dimensional, three-dimensional, or somehow otherwise feed into the visual system to convey information. An image can be an artifact, such as a photograph or other two-dimensiona ...
; then a light-sensitive part of the eye (the
retina
The retina (from la, rete "net") is the innermost, light-sensitive layer of tissue of the eye of most vertebrates and some molluscs. The optics of the eye create a focused two-dimensional image of the visual world on the retina, which then ...
) where the images fall and are processed. The retina makes a connection to the
brain
A brain is an organ that serves as the center of the nervous system in all vertebrate and most invertebrate animals. It is located in the head, usually close to the sensory organs for senses such as vision. It is the most complex organ in a ve ...
via the
optic nerve
In neuroanatomy, the optic nerve, also known as the second cranial nerve, cranial nerve II, or simply CN II, is a paired cranial nerve that transmits visual information from the retina to the brain. In humans, the optic nerve is derived fro ...
. The remaining components of the eye keep it in its required shape, nourish and maintain it, and protect it.
Three types of cells in the retina convert light energy into electrical energy used by the
nervous system
In biology, the nervous system is the highly complex part of an animal that coordinates its actions and sensory information by transmitting signals to and from different parts of its body. The nervous system detects environmental changes ...
: rods respond to low intensity light and contribute to perception of low-resolution, black-and-white images;
cones
A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex.
A cone is formed by a set of line segments, half-lines, or lines conn ...
respond to high intensity light and contribute to perception of high-resolution, coloured images; and the recently discovered
photosensitive ganglion cell
Intrinsically photosensitive retinal ganglion cells (ipRGCs), also called photosensitive retinal ganglion cells (pRGC), or melanopsin-containing retinal ganglion cells (mRGCs), are a type of neuron in the retina of the mammalian eye. The presence ...
s respond to a full range of light intensities and contribute to adjusting the amount of light reaching the retina, to regulating and suppressing the hormone melatonin, and to
entraining
Entrainment may refer to:
* Air entrainment, the intentional creation of tiny air bubbles in concrete
* Brainwave entrainment, the practice of entraining one's brainwaves to a desired frequency
* Entrainment (biomusicology), the synchronization ...
Humans have two eyes, situated on the left and the right of the
face
The face is the front of an animal's head that features the eyes, nose and mouth, and through which animals express many of their emotions. The face is crucial for human identity, and damage such as scarring or developmental deformities may aff ...
. The eyes sit in bony cavities called the
orbits
In celestial mechanics, an orbit is the curved trajectory of an object such as the trajectory of a planet around a star, or of a natural satellite around a planet, or of an artificial satellite around an object or position in space such as a p ...
, in the
skull
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
. There are six
extraocular muscles
The extraocular muscles (extrinsic ocular muscles), are the seven extrinsic muscles of the human eye. Six of the extraocular muscles, the four recti muscles, and the superior and inferior oblique muscles, control movement of the eye and the othe ...
that control eye movements. The front visible part of the eye is made up of the whitish sclera, a coloured
iris
Iris most often refers to:
*Iris (anatomy), part of the eye
*Iris (mythology), a Greek goddess
* ''Iris'' (plant), a genus of flowering plants
* Iris (color), an ambiguous color term
Iris or IRIS may also refer to:
Arts and media
Fictional ent ...
, and the
pupil
The pupil is a black hole located in the center of the Iris (anatomy), iris of the Human eye, eye that allows light to strike the retina.Cassin, B. and Solomon, S. (1990) ''Dictionary of Eye Terminology''. Gainesville, Florida: Triad Publishing ...
. A thin layer called the
conjunctiva
The conjunctiva is a thin mucous membrane that lines the inside of the eyelids and covers the sclera (the white of the eye). It is composed of non-keratinized, stratified squamous epithelium with goblet cells, stratified columnar epithelium ...
sits on top of this. The front part is also called the anterior segment of the eye.
The eye is not shaped like a perfect sphere, rather it is a fused two-piece unit, composed of an anterior (front) segment and the posterior (back) segment. The anterior segment is made up of the cornea, iris and lens. The cornea is transparent and more curved and is linked to the larger posterior segment, composed of the vitreous, retina, choroid and the outer white shell called the sclera. The cornea is typically about in diameter, and 0.5 mm (500 μm) in thickness near its centre. The posterior chamber constitutes the remaining five-sixths; its diameter is typically about . An area termed the limbus connects the cornea and sclera. The iris is the pigmented circular structure concentrically surrounding the centre of the eye, the pupil, which appears to be black. The size of the pupil, which controls the amount of light entering the eye, is adjusted by the iris'
dilator
Dilator or dilatator is a medical term with a number of uses, including:
*A surgical instrument or medical implement used to induce dilation, that is, to expand an opening or passage such as the cervix (see cervical dilator), urethra, esophagus, ...
and sphincter muscles.
Light energy enters the eye through the cornea, through the pupil and then through the lens. The lens shape is changed for near focus (accommodation) and is controlled by the ciliary muscle. Photons of light falling on the light-sensitive cells of the retina ( photoreceptor cones and rods) are converted into electrical signals that are transmitted to the brain by the optic nerve and interpreted as sight and vision.
Size
The size of the eye differs among adults by only one or 2 millimetres. The eyeball is generally less tall than it is wide. The sagittal vertical (height) of a human adult eye is approximately , the transverse horizontal diameter (width) is and the axial anteroposterior size (depth) averages with no significant difference between sexes and age groups. Strong correlation has been found between the transverse diameter and the width of the orbit (r = 0.88). The typical adult eye has an anterior to posterior diameter of , and a volume of .
The eyeball grows rapidly, increasing from about diameter at birth to by three years of age. By age 12, the eye attains its full size.
Components
The eye is made up of three coats, or layers, enclosing various anatomical structures. The outermost layer, known as the fibrous tunic, is composed of the
cornea
The cornea is the transparent front part of the eye that covers the iris, pupil, and anterior chamber. Along with the anterior chamber and lens, the cornea refracts light, accounting for approximately two-thirds of the eye's total optical ...
and sclera, which provide shape to the eye and support the deeper structures. The middle layer, known as the vascular tunic or uvea, consists of the choroid,
ciliary body
The ciliary body is a part of the eye that includes the ciliary muscle, which controls the shape of the lens, and the ciliary epithelium, which produces the aqueous humor. The aqueous humor is produced in the non-pigmented portion of the ciliar ...
, pigmented epithelium and
iris
Iris most often refers to:
*Iris (anatomy), part of the eye
*Iris (mythology), a Greek goddess
* ''Iris'' (plant), a genus of flowering plants
* Iris (color), an ambiguous color term
Iris or IRIS may also refer to:
Arts and media
Fictional ent ...
. The innermost is the
retina
The retina (from la, rete "net") is the innermost, light-sensitive layer of tissue of the eye of most vertebrates and some molluscs. The optics of the eye create a focused two-dimensional image of the visual world on the retina, which then ...
, which gets its oxygenation from the blood vessels of the choroid (posteriorly) as well as the retinal vessels (anteriorly).
The spaces of the eye are filled with the
aqueous humour
The aqueous humour is a transparent water-like fluid similar to plasma, but containing low protein concentrations. It is secreted from the ciliary body, a structure supporting the lens of the eyeball. It fills both the anterior and the poster ...
anteriorly, between the cornea and lens, and the
vitreous body
The vitreous body (''vitreous'' meaning "glass-like"; , ) is the clear gel that fills the space between the lens and the retina of the eyeball (the vitreous chamber) in humans and other vertebrates. It is often referred to as the vitreous hu ...
, a jelly-like substance, behind the lens, filling the entire posterior cavity. The aqueous humour is a clear watery fluid that is contained in two areas: the
anterior chamber
The anterior chamber ( AC) is the aqueous humor-filled space inside the eye between the iris and the cornea's innermost surface, the endothelium. Hyphema, anterior uveitis and glaucoma are three main pathologies in this area. In hyphema, blood f ...
between the cornea and the iris, and the
posterior chamber
The posterior chamber is a narrow space behind the peripheral part of the iris, and in front of the suspensory ligament of the lens and the ciliary processes. The posterior chamber consists of small space directly posterior to the iris but anterio ...
between the iris and the lens. The lens is suspended to the ciliary body by the suspensory ligament (
Zonule of Zinn
The zonule of Zinn () (Zinn's membrane, ciliary zonule) (after Johann Gottfried Zinn) is a ring of fibrous strands forming a zonule (little band) that connects the ciliary body with the crystalline lens of the eye. These fibers are sometimes coll ...
), made up of hundreds of fine transparent fibers which transmit muscular forces to change the shape of the lens for accommodation (focusing). The vitreous body is a clear substance composed of water and proteins, which give it a jelly-like and sticky composition.
Structures surrounding the eye
Extraocular muscles
Each eye has seven
extraocular muscles
The extraocular muscles (extrinsic ocular muscles), are the seven extrinsic muscles of the human eye. Six of the extraocular muscles, the four recti muscles, and the superior and inferior oblique muscles, control movement of the eye and the othe ...
located in its
orbit
In celestial mechanics, an orbit is the curved trajectory of an object such as the trajectory of a planet around a star, or of a natural satellite around a planet, or of an artificial satellite around an object or position in space such as ...
. Six of these muscles control the
eye movements
Eye movement includes the voluntary or involuntary movement of the eyes. Eye movements are used by a number of organisms (e.g. primates, rodents, flies, birds, fish, cats, crabs, octopus) to fixate, inspect and track visual objects of inte ...
, the seventh controls the movement of the upper
eyelid
An eyelid is a thin fold of skin that covers and protects an eye. The levator palpebrae superioris muscle retracts the eyelid, exposing the cornea to the outside, giving vision. This can be either voluntarily or involuntarily. The human eye ...
. The six muscles are four recti muscles – the
lateral rectus
Lateral is a geometric term of location which may refer to:
Healthcare
* Lateral (anatomy), an anatomical direction
*Lateral cricoarytenoid muscle
*Lateral release (surgery), a surgical procedure on the side of a kneecap
Phonetics
*Lateral co ...
, the
medial rectus
The medial rectus muscle is a muscle in the orbit near the eye. It is one of the extraocular muscles. It originates from the common tendinous ring, and inserts into the anteromedial surface of the eye. It is supplied by the inferior division of ...
, the
inferior rectus
Inferior may refer to:
* Inferiority complex
* An anatomical term of location
* Inferior angle of the scapula, in the human skeleton
* ''Inferior'' (book), by Angela Saini
* ''The Inferior'', a 2007 novel by Peadar Ó Guilín
See also
*Junior (di ...
, and the
superior rectus
The superior rectus muscle is a muscle in the orbit. It is one of the extraocular muscles. It is innervated by the superior division of the oculomotor nerve (III). In the primary position (looking straight ahead), its primary function is elevation ...
, and two oblique muscles the
inferior oblique
The inferior oblique muscle or obliquus oculi inferior is a thin, narrow muscle placed near the anterior margin of the floor of the orbit. The inferior oblique is one of the extraocular muscles, and is attached to the maxillary bone (origin) and ...
, and the
superior oblique
The superior oblique muscle, or obliquus oculi superior, is a fusiform muscle originating in the upper, medial side of the orbit (i.e. from beside the nose) which abducts, depresses and internally rotates the eye. It is the only extraocular musc ...
. The seventh muscle is the
levator palpebrae superioris muscle
The levator palpebrae superioris ( la, elevating muscle of upper eyelid) is the muscle in the orbit that elevates the upper eyelid.
Structure
The levator palpebrae superioris originates from inferior surface of the lesser wing of the sphenoid bon ...
. When the muscles exert different tensions, a torque is exerted on the globe that causes it to turn, in almost pure rotation, with only about one millimeter of translation. Thus, the eye can be considered as undergoing rotations about a single point in the centre of the eye.
File:Eye orbit anterior.jpg, Eye and orbit anatomy with motor nerves
File:Lateral orbit nerves.jpg, Image showing orbita with eye and nerves visible (periocular fat removed)
File:Lateral orbit anatomy 2.jpg, Image showing orbita with eye and periocular fat
File:Eye orbit anatomy anterior2.jpg, Normal
anatomy
Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having it ...
of the human eye and orbit, anterior view
Vision
Field of view
The approximate
field of view
The field of view (FoV) is the extent of the observable world that is seen at any given moment. In the case of optical instruments or sensors it is a solid angle through which a detector is sensitive to electromagnetic radiation.
Human ...
of an individual human eye (measured from the fixation point, i.e., the point at which one's gaze is directed) varies by facial anatomy, but is typically 30° superior (up, limited by the brow), 45° nasal (limited by the nose), 70° inferior (down), and 100° temporal (towards the temple). For both eyes combined (
Binocular vision
In biology, binocular vision is a type of vision in which an animal has two eyes capable of facing the same direction to perceive a single three-dimensional image of its surroundings. Binocular vision does not typically refer to vision where an ...
) visual field is approximately 100° vertical and a maximum 190° horizontal, approximately 120° of which makes up the binocular field of view (seen by both eyes) flanked by two uniocular fields (seen by only one eye) of approximately 40 degrees. It is an area of 4.17
steradian
The steradian (symbol: sr) or square radian is the unit of solid angle in the International System of Units (SI). It is used in three-dimensional geometry, and is analogous to the radian, which quantifies planar angles. Whereas an angle in radian ...
s or 13700
square degree
__NOTOC__
A square degree (deg2) is a non- SI unit measure of solid angle. Other denotations include ''sq. deg.'' and (°)2. Just as degrees are used to measure parts of a circle, square degrees are used to measure parts of a sphere. Analogous to ...
s for binocular vision. When viewed at large angles from the side, the iris and pupil may still be visible by the viewer, indicating the person has peripheral vision possible at that angle.
About 15° temporal and 1.5° below the horizontal is the blind spot created by the optic nerve nasally, which is roughly 7.5° high and 5.5° wide.
Dynamic range
The retina has a static contrast ratio of around 100:1 (about 6.5 f-stops). As soon as the eye moves rapidly to acquire a target (
saccade
A saccade ( , French for ''jerk'') is a quick, simultaneous movement of both eyes between two or more phases of fixation in the same direction.Cassin, B. and Solomon, S. ''Dictionary of Eye Terminology''. Gainesville, Florida: Triad Publishi ...
s), it re-adjusts its exposure by adjusting the iris, which adjusts the size of the pupil. Initial dark adaptation takes place in approximately four seconds of profound, uninterrupted darkness; full adaptation through adjustments in retinal rod photoreceptors is 80% complete in thirty minutes. The process is nonlinear and multifaceted, so an interruption by light exposure requires restarting the dark adaptation process over again.
The human eye can detect a luminance from 10−6 cd/m2, or one millionth (0.000001) of a
candela
The candela ( or ; symbol: cd) is the unit of luminous intensity in the International System of Units (SI). It measures luminous power per unit solid angle emitted by a light source in a particular direction. Luminous intensity is analogous t ...
per square meter to 108 cd/m2 or one hundred million (100,000,000) candelas per square meter. (that is it has a range of 1014 cd/m2, or one hundred trillion 100,000,000,000,000, about 46.5 f-stops). This range does not include looking at the midday sun (109 cd/m2) or lightning discharge.
At the low end of the range is the
absolute threshold
In neuroscience and psychophysics, an absolute threshold was originally defined as the lowest level of a stimulus – light, sound, touch, etc. – that an organism could detect. Under the influence of signal detection theory, absolute thresho ...
of vision for a steady light across a wide field of view, about 10−6 cd/m2 (0.000001 candela per square meter). The upper end of the range is given in terms of normal visual performance as 108 cd/m2 (100,000,000 or one hundred million candelas per square meter).
The eye includes a
lens
A lens is a transmissive optical device which focuses or disperses a light beam by means of refraction. A simple lens consists of a single piece of transparent material, while a compound lens consists of several simple lenses (''elements ...
similar to
lenses
A lens is a transmissive optical device which focuses or disperses a light beam by means of refraction. A simple lens consists of a single piece of transparent material, while a compound lens consists of several simple lenses (''elements''), ...
found in optical instruments such as cameras and the same physics principles can be applied. The
pupil
The pupil is a black hole located in the center of the Iris (anatomy), iris of the Human eye, eye that allows light to strike the retina.Cassin, B. and Solomon, S. (1990) ''Dictionary of Eye Terminology''. Gainesville, Florida: Triad Publishing ...
aperture
In optics, an aperture is a hole or an opening through which light travels. More specifically, the aperture and focal length of an optical system determine the cone angle of a bundle of rays that come to a focus in the image plane.
An ...
; the iris is the diaphragm that serves as the aperture stop. Refraction in the
cornea
The cornea is the transparent front part of the eye that covers the iris, pupil, and anterior chamber. Along with the anterior chamber and lens, the cornea refracts light, accounting for approximately two-thirds of the eye's total optical ...
causes the effective aperture (the entrance pupil) to differ slightly from the physical pupil diameter. The entrance pupil is typically about 4 mm in diameter, although it can range from 2 mm () in a brightly lit place to 8 mm () in the dark. The latter value decreases slowly with age; older people's eyes sometimes dilate to not more than 5–6mm in the dark, and may be as small as 1mm in the light.
Eye movement
The visual system in the human brain is too slow to process information if images are slipping across the retina at more than a few degrees per second. Thus, to be able to see while moving, the brain must compensate for the motion of the head by turning the eyes. Frontal-eyed animals have a small area of the retina with very high visual acuity, the
fovea centralis
The fovea centralis is a small, central pit composed of closely packed cones in the eye. It is located in the center of the macula lutea of the retina.
The fovea is responsible for sharp central vision (also called foveal vision), which is ne ...
. It covers about 2 degrees of visual angle in people. To get a clear view of the world, the brain must turn the eyes so that the image of the object of regard falls on the fovea. Any failure to make eye movements correctly can lead to serious visual degradation.
Having two eyes allows the brain to determine the depth and distance of an object, called stereovision, and gives the sense of three-dimensionality to the vision. Both eyes must point accurately enough that the object of regard falls on corresponding points of the two retinas to stimulate stereovision; otherwise, double vision might occur. Some persons with congenitally crossed eyes tend to ignore one eye's vision, thus do not suffer double vision, and do not have stereovision. The movements of the eye are controlled by six muscles attached to each eye, and allow the eye to elevate, depress, converge, diverge and roll. These muscles are both controlled voluntarily and involuntarily to track objects and correct for simultaneous head movements.
Rapid eye movement
Rapid eye movement, REM, typically refers to the
sleep
Sleep is a sedentary state of mind and body. It is characterized by altered consciousness, relatively inhibited sensory activity, reduced muscle activity and reduced interactions with surroundings. It is distinguished from wakefulness by a de ...
stage during which the most vivid dreams occur. During this stage, the eyes move rapidly.
Saccades
Saccades are quick, simultaneous movements of both eyes in the same direction controlled by the frontal lobe of the brain.
Fixational Eye Movements
Even when looking intently at a single spot, the eyes drift around. This ensures that individual photosensitive cells are continually stimulated in different degrees. Without changing input, these cells would otherwise stop generating output.
Eye movements include drift,
ocular tremor
Ocular tremor (ocular microtremor) is a constant, involuntary eye tremor of a low amplitude and high frequency. It is a type of fixational eye movement that occurs in all normal people, even when the eye appears still. The frequency of ocular mic ...
, and microsaccades. Some irregular drifts, movements smaller than a saccade and larger than a microsaccade, subtend up to one tenth of a degree. Researchers vary in their definition of
microsaccade
Microsaccades are a kind of fixational eye movement. They are small, jerk-like, involuntary eye movements, similar to miniature versions of voluntary saccades. They typically occur during prolonged visual fixation (of at least several seconds), ...
s by amplitude. Martin Rolfs states that 'the majority of microsaccades observed in a variety of tasks have amplitudes smaller than 30 min-arc'. However, others state that the "current consensus has largely consolidated around a definition of microsaccades that includes magnitudes up to 1°."
retina
The retina (from la, rete "net") is the innermost, light-sensitive layer of tissue of the eye of most vertebrates and some molluscs. The optics of the eye create a focused two-dimensional image of the visual world on the retina, which then ...
during head movement by producing an eye movement in the direction opposite to head movement in response to neural input from the vestibular system of the inner ear, thus maintaining the image in the centre of the visual field. For example, when the head moves to the right, the eyes move to the left. This applies for head movements up and down, left and right, and tilt to the right and left, all of which give input to the ocular muscles to maintain visual stability.
Smooth pursuit movement
Eyes can also follow a moving object around. This tracking is less accurate than the vestibulo-ocular reflex, as it requires the brain to process incoming visual information and supply feedback. Following an object moving at constant speed is relatively easy, though the eyes will often make saccades to keep up. The smooth pursuit movement can move the eye at up to 100°/s in adult humans.
It is more difficult to visually estimate speed in low light conditions or while moving, unless there is another point of reference for determining speed.
Optokinetic reflex
The Optokinetic reflex (or optokinetic nystagmus) stabilizes the image on the retina through visual feedback. It is induced when the entire visual scene drifts across the retina, eliciting eye rotation in the same direction and at a velocity that minimizes the motion of the image on the retina. When the gaze direction deviates too far from the forward heading, a compensatory saccade is induced to reset the gaze to the centre of the visual field.
For example, when looking out of the window at a moving train, the eyes can focus on a moving train for a short moment (by stabilizing it on the retina), until the train moves out of the field of vision. At this point, the eye is moved back to the point where it first saw the train (through a saccade).
Near response
The adjustment to close-range vision involves three processes to focus an image on the retina.
Vergence movement
When a creature with binocular vision looks at an object, the eyes must rotate around a vertical axis so that the projection of the image is in the centre of the retina in both eyes. To look at a nearby object, the eyes rotate 'towards each other' (
convergence
Convergence may refer to:
Arts and media Literature
*''Convergence'' (book series), edited by Ruth Nanda Anshen
*Convergence (comics), "Convergence" (comics), two separate story lines published by DC Comics:
**A four-part crossover storyline that ...
), while for an object farther away they rotate 'away from each other' (
divergence
In vector calculus, divergence is a vector operator that operates on a vector field, producing a scalar field giving the quantity of the vector field's source at each point. More technically, the divergence represents the volume density of t ...
).
Pupil constriction
Lenses cannot refract light rays at their edges as well as closer to the centre. The image produced by any lens is therefore somewhat blurry around the edges ( spherical aberration). It can be minimized by screening out peripheral light rays and looking only at the better-focused centre. In the eye, the pupil serves this purpose by constricting while the eye is focused on nearby objects. Small apertures also give an increase in depth of field, allowing a broader range of "in focus" vision. In this way the pupil has a dual purpose for near vision: to reduce spherical aberration and increase depth of field.
Accommodation of the lens
Changing the curvature of the lens is carried out by the
ciliary muscle
The ciliary muscle is an intrinsic muscle of the eye formed as a ring of smooth muscleSchachar, Ronald A. (2012). "Anatomy and Physiology." (Chapter 4) . in the eye's middle layer, uvea (vascular layer). It controls accommodation for viewing ...
s surrounding the lens; this process is known as "accommodation". Accommodation narrows the inner diameter of the ciliary body, which actually relaxes the fibers of the suspensory ligament attached to the periphery of the lens, and also allows the lens to relax into a more convex, or globular, shape. A more convex lens refracts light more strongly and focuses divergent light rays from near objects onto the retina, allowing closer objects to be brought into better focus.
Eye care professionals
The human eye contains enough complexity to warrant specialized attention and care beyond the duties of a general practitioner. These specialists, or
eye care professional
An eye care professional (ECP) is an individual who provides a service related to the eyes or vision. It is any healthcare worker involved in eye care, from one with a small amount of post-secondary training to practitioners with a doctoral level ...
s, serve different functions in different countries. Eye care professionals can have overlap in their patient care privileges. For example, both an ophthalmologist (M.D.) and optometrist (O.D.) are professionals who diagnoses eye disease and can prescribe lenses to correct vision. However, typically only ophthalmologists are licensed to perform surgical procedures. Ophthalmologists may also specialize within a surgical area, such as
cornea
The cornea is the transparent front part of the eye that covers the iris, pupil, and anterior chamber. Along with the anterior chamber and lens, the cornea refracts light, accounting for approximately two-thirds of the eye's total optical ...
,
cataracts
A cataract is a cloudy area in the lens of the eye that leads to a decrease in vision. Cataracts often develop slowly and can affect one or both eyes. Symptoms may include faded colors, blurry or double vision, halos around light, trouble w ...
,
laser
A laser is a device that emits light through a process of optical amplification based on the stimulated emission of electromagnetic radiation. The word "laser" is an acronym for "light amplification by stimulated emission of radiation". The fi ...
,
retina
The retina (from la, rete "net") is the innermost, light-sensitive layer of tissue of the eye of most vertebrates and some molluscs. The optics of the eye create a focused two-dimensional image of the visual world on the retina, which then ...
, or
oculoplastics
Oculoplastics, or oculoplastic surgery, includes a wide variety of surgical procedures that deal with the orbit (eye socket), eyelids, tear ducts, and the face. It also deals with the reconstruction of the eye and associated structures.
Training ...
.
Eye care professionals include:
*
Ocularist An ocularist specializes in the fabrication and fitting of ocular prostheses for people who have lost an eye or eyes due to trauma or illness.{{Cite journal , last1=Khandekar , first1=Rajiv , last2=Changal , first2=Nusrat , last3=AlRasheed , first ...
s
*
Ophthalmologists
Ophthalmology ( ) is a surgical subspecialty within medicine that deals with the diagnosis and treatment of eye disorders.
An ophthalmologist is a physician who undergoes subspecialty training in medical and surgical eye care. Following a medic ...
*
Optometrists
Optometry is a specialized health care profession that involves examining the eyes and related structures for defects or abnormalities. Optometrists are health care professionals who typically provide comprehensive primary eye care.
In the Un ...
*
Opticians
An optician, or ''dispensing optician'', is a technical practitioner who designs, fits and dispenses lenses for the correction of a person's vision. Opticians determine the specifications of various ophthalmic appliances that will give the nec ...
*
Orthoptists
Vision therapy (VT), or behavioral optometry, is an umbrella term for alternative medicine treatments using eye exercises, based around the pseudoscientific claim that vision problems are the true underlying cause of learning disabilities, par ...
Eye irritation has been defined as "the magnitude of any stinging, scratching, burning, or other irritating sensation from the eye". It is a common problem experienced by people of all ages. Related eye symptoms and signs of irritation are discomfort, dryness, excess tearing, itchiness, grating, foreign body sensation, ocular fatigue, pain, soreness, redness, swollen eyelids, and tiredness, etc. These eye symptoms are reported with intensities from mild to severe. It has been suggested that these eye symptoms are related to different causal mechanisms, and symptoms are related to the particular ocular anatomy involved.
Several suspected causal factors in our environment have been studied so far. One hypothesis is that
indoor air pollution
Indoor air quality (IAQ) is the air quality within and around buildings and structures. IAQ is known to affect the health, comfort, and well-being of building occupants. Poor indoor air quality has been linked to sick building syndrome, reduce ...
may cause eye and airway irritation. Eye irritation depends somewhat on destabilization of the outer-eye tear film, i.e. the formation of dry spots on the cornea, resulting in ocular discomfort. Occupational factors are also likely to influence the perception of eye irritation. Some of these are lighting (glare and poor contrast), gaze position, reduced blink rate, limited number of breaks from visual tasking, and a constant combination of accommodation, musculoskeletal burden, and impairment of the visual nervous system. Another factor that may be related is work stress. In addition, psychological factors have been found in multivariate analyses to be associated with an increase in eye irritation among VDU users. Other risk factors, such as chemical toxins/irritants (e.g. amines, formaldehyde, acetaldehyde, acrolein, N-decane, VOCs, ozone, pesticides and preservatives, allergens, etc.) might cause eye irritation as well.
Certain
volatile organic compound
Volatile organic compounds (VOCs) are organic compounds that have a high vapour pressure at room temperature. High vapor pressure correlates with a low boiling point, which relates to the number of the sample's molecules in the surrounding air, a ...
s that are both chemically reactive and airway irritants may cause eye irritation. Personal factors (e.g. use of contact lenses, eye make-up, and certain medications) may also affect destabilization of the tear film and possibly result in more eye symptoms. Nevertheless, if airborne particles alone should destabilize the tear film and cause eye irritation, their content of surface-active compounds must be high. An integrated physiological risk model with blink frequency, destabilization, and break-up of the eye tear film as inseparable phenomena may explain eye irritation among office workers in terms of occupational, climate, and eye-related physiological risk factors.
There are two major measures of eye irritation. One is blink frequency which can be observed by human behavior. The other measures are break up time, tear flow, hyperemia (redness, swelling), tear fluid cytology, and epithelial damage (vital stains) etc., which are human beings' physiological reactions. Blink frequency is defined as the number of blinks per minute and it is associated with eye irritation. Blink frequencies are individual with mean frequencies of < 2–3 to 20–30 blinks/minute, and they depend on environmental factors including the use of contact lenses. Dehydration, mental activities, work conditions, room temperature, relative humidity, and illumination all influence blink frequency. Break-up time (BUT) is another major measure of eye irritation and tear film stability. It is defined as the time interval (in seconds) between blinking and rupture. BUT is considered to reflect the stability of the tear film as well. In normal persons, the break-up time exceeds the interval between blinks, and, therefore, the tear film is maintained. Studies have shown that blink frequency is correlated negatively with break-up time. This phenomenon indicates that perceived eye irritation is associated with an increase in blink frequency since the cornea and conjunctiva both have sensitive nerve endings that belong to the first trigeminal branch.Sibony PA, Evinger C. "Anatomy and physiology of normal and abnormal eyelid position and movement". In: Miller NR, Newman NJ, editors. Walsh & Hoyt's ''Clinical Neuro-ophthalmology''. Baltimore, MD: Williams and Wilkins; 1998. pp. 1509–1592 Other evaluating methods, such as hyperemia, cytology etc. have increasingly been used to assess eye irritation.
There are other factors that are related to eye irritation as well. Three major factors that influence the most are indoor air pollution, contact lenses and gender differences. Field studies have found that the prevalence of objective eye signs is often significantly altered among office workers in comparisons with random samples of the general population. These research results might indicate that indoor air pollution has played an important role in causing eye irritation. There are more and more people wearing contact lens now and dry eyes appear to be the most common complaint among contact lens wearers. Although both contact lens wearers and spectacle wearers experience similar eye irritation symptoms, dryness, redness, and grittiness have been reported far more frequently among contact lens wearers and with greater severity than among spectacle wearers. Studies have shown that incidence of dry eyes increases with age,Seal, D. V., and Mackie, I. A. (1986). "The questionable dry eye as a clinical and biochemical entity". In F. J. Holly (Ed.) ''The preocular tear film – In health, disease, and contact lens wear''. Dry Eye Institute, Lubbock, TX, pp. 41–51. especially among women. Tear film stability (e.g.
tear break-up time Tear break-up time (TBUT) also known as tear film break-up time (TFBUT) is the time taken for the first dry spot to appear on the cornea after a complete blinking, blink. TFBUT measurement is an easy and fast method used to assess the stability of ...
) is significantly lower among women than among men. In addition, women have a higher blink frequency while reading. Several factors may contribute to gender differences. One is the use of eye make-up. Another reason could be that the women in the reported studies have done more VDU work than the men, including lower grade work. A third often-quoted explanation is related to the age-dependent decrease of tear secretion, particularly among women after 40 years of age.
In a study conducted by
UCLA
The University of California, Los Angeles (UCLA) is a public land-grant research university in Los Angeles, California. UCLA's academic roots were established in 1881 as a teachers college then known as the southern branch of the California ...
, the frequency of reported symptoms in industrial buildings was investigated. The study's results were that eye irritation was the most frequent symptom in industrial building spaces, at 81%. Modern office work with use of office equipment has raised concerns about possible adverse health effects. Since the 1970s, reports have linked mucosal, skin, and general symptoms to work with self-copying paper. Emission of various particulate and volatile substances has been suggested as specific causes. These symptoms have been related to sick building syndrome (SBS), which involves symptoms such as irritation to the eyes, skin, and upper airways, headache and fatigue.
Many of the symptoms described in SBS and
multiple chemical sensitivity
Multiple chemical sensitivity (MCS), also known as idiopathic environmental intolerances (IEI), is an unrecognized and controversial diagnosis characterized by chronic symptoms attributed to exposure to low levels of commonly used chemicals. Sym ...
(MCS) resemble the symptoms known to be elicited by airborne irritant chemicals. A repeated measurement design was employed in the study of acute symptoms of eye and respiratory tract irritation resulting from occupational exposure to sodium borate dusts. The symptom assessment of the 79 exposed and 27 unexposed subjects comprised interviews before the shift began and then at regular hourly intervals for the next six hours of the shift, four days in a row. Exposures were monitored concurrently with a personal real time aerosol monitor. Two different exposure profiles, a daily average and short term (15 minute) average, were used in the analysis. Exposure-response relations were evaluated by linking incidence rates for each symptom with categories of exposure.
Acute incidence rates for nasal, eye, and
throat irritation
Throat irritation can refer to a dry cough, a scratchy feeling at the back of the throat, a sensation of a lumpy feeling, something stuck at the back of the throat, or possibly a feeling of dust in the throat. The symptoms are unpleasant and usuall ...
, and coughing and breathlessness were found to be associated with increased exposure levels of both exposure indices. Steeper exposure-response slopes were seen when short term exposure concentrations were used. Results from multivariate logistic regression analysis suggest that current smokers tended to be less sensitive to the exposure to airborne sodium borate dust.
Several actions can be taken to prevent eye irritation—
* trying to maintain normal blinking by avoiding room temperatures that are too high; avoiding relative humidities that are too high or too low, because they reduce blink frequency or may increase water evaporation.
* trying to maintain an intact film of tears by the following actions:
:# Blinking and short breaks may be beneficial for VDU users. Increasing these two actions might help maintain the tear film.
:# Downward gazing is recommended to reduce ocular surface area and water evaporation.
:# The distance between the VDU and keyboard should be kept as short as possible to minimize evaporation from the ocular surface area by a low direction of the gaze, and
:#Blink training can be beneficial.
In addition, other measures are proper lid hygiene, avoidance of eye rubbing, and proper use of personal products and medication. Eye make-up should be used with care.
Eye disease
There are many diseases, disorders, and age-related changes that may affect the eyes and surrounding structures.
As the eye ages, certain changes occur that can be attributed solely to the aging process. Most of these anatomic and physiologic processes follow a gradual decline. With aging, the quality of vision worsens due to reasons independent of diseases of the aging eye. While there are many changes of significance in the non-diseased eye, the most functionally important changes seem to be a reduction in pupil size and the loss of accommodation or focusing capability (
presbyopia
Presbyopia is physiological insufficiency of accommodation associated with the aging of the eye that results in progressively worsening ability to focus clearly on close objects. Also known as age-related farsightedness (or age-related long si ...
). The area of the pupil governs the amount of light that can reach the retina. The extent to which the pupil dilates decreases with age, leading to a substantial decrease in light received at the retina. In comparison to younger people, it is as though older persons are constantly wearing medium-density sunglasses. Therefore, for any detailed visually guided tasks on which performance varies with illumination, older persons require extra lighting. Certain ocular diseases can come from
sexually transmitted disease
Sexually transmitted infections (STIs), also referred to as sexually transmitted diseases (STDs) and the older term venereal diseases, are infections that are spread by sexual activity, especially vaginal intercourse, anal sex, and oral ...
s such as herpes and genital warts. If contact between the eye and area of infection occurs, the STD can be transmitted to the eye.
With aging, a prominent white ring develops in the periphery of the cornea called
arcus senilis
Arcus senilis (AS), also known as gerontoxon, arcus lipoides, arcus corneae, corneal arcus, arcus adiposus, or arcus cornealis, are rings in the peripheral cornea. It‘s usually caused by cholesterol deposits, so it may be a sign of high choleste ...
. Aging causes laxity, downward shift of eyelid tissues and atrophy of the orbital fat. These changes contribute to the etiology of several eyelid disorders such as
ectropion
Ectropion is a medical condition in which the lower eyelid turns outwards. It is one of the notable aspects of newborns exhibiting congenital Harlequin-type ichthyosis, but ectropion can occur due to any weakening of tissue of the lower eyelid ...
,
entropion
Entropion is a medical condition in which the eyelid (usually the lower lid) folds inward. It is very uncomfortable, as the eyelashes continuously rub against the cornea causing irritation. Entropion is usually caused by genetic factors. This is d ...
,
dermatochalasis
Dermatochalasis is a medical condition, defined as an excess of skin in the upper or lower eyelid, also known as "baggy eyes." It may be either an acquired or a congenital condition. It is generally treated with blepharoplasty.
Presentation Ass ...
, and ptosis. The vitreous gel undergoes liquefaction (
posterior vitreous detachment
A posterior vitreous detachment (PVD) is a condition of the eye in which the vitreous membrane separates from the retina.
It refers to the separation of the posterior hyaloid membrane from the retina anywhere posterior to the vitreous base (a 3 ...
or PVD) and its opacities — visible as
floater
Floaters or eye floaters are sometimes visible deposits (e.g., the shadows of tiny structures of protein or other cell debris projected onto the retina) within the eye's vitreous humour ("the vitreous"), which is normally transparent, or betwe ...
s — gradually increase in number.
Eye care professional
An eye care professional (ECP) is an individual who provides a service related to the eyes or vision. It is any healthcare worker involved in eye care, from one with a small amount of post-secondary training to practitioners with a doctoral level ...
s, including ophthalmologists and optometrists, are involved in the treatment and management of ocular and vision disorders. A
Snellen chart
A Snellen chart is an eye chart that can be used to measure visual acuity. Snellen charts are named after the Dutch ophthalmologist Herman Snellen, who developed the chart in 1862. Many ophthalmologists and vision scientists now use an improved ...
is one type of
eye chart
__NOTOC__
An eye chart, or optotype, is a chart used to subjectively measure visual acuity. Eye charts are often used by health care professionals, such as optometrists, physicians or nurses, to screen persons for vision impairment. Ophthalmologi ...
used to measure
visual acuity
Visual acuity (VA) commonly refers to the clarity of vision, but technically rates an examinee's ability to recognize small details with precision. Visual acuity is dependent on optical and neural factors, i.e. (1) the sharpness of the retinal ...
. At the conclusion of a complete
eye examination
An eye examination is a series of tests performed to assess vision and ability to focus on and discern objects. It also includes other tests and examinations pertaining to the eyes. Eye examinations are primarily performed by an optometrist, ...
, the eye doctor might provide the patient with an
eyeglass prescription
An eyeglass prescription is an order written by an eyewear prescriber, such as an optometrist, that specifies the value of all parameters the prescriber has deemed necessary to construct and/or dispense corrective lenses appropriate for a patie ...
for
corrective lens
A corrective lens is a lens (i.e. a transmissive optical device) that is typically worn in front of the eye to improve daily vision. The most common use is to treat refractive errors: myopia, hypermetropia, astigmatism, and presbyopia. G ...
es. Some disorders of the eyes for which corrective lenses are prescribed include myopia (
near-sightedness
Near-sightedness, also known as myopia and short-sightedness, is an eye disease where light focuses in front of, instead of on, the retina. As a result, distant objects appear blurry while close objects appear normal. Other symptoms may includ ...
),
hyperopia
Far-sightedness, also known as long-sightedness, hypermetropia, or hyperopia, is a condition of the eye where distant objects are seen clearly but near objects appear blurred. This blurred effect is due to incoming light being focused behind, i ...
(far-sightedness),
astigmatism
Astigmatism is a type of refractive error due to rotational asymmetry in the eye's refractive power. This results in distorted or blurred vision at any distance. Other symptoms can include eyestrain, headaches, and trouble driving at n ...
, and
presbyopia
Presbyopia is physiological insufficiency of accommodation associated with the aging of the eye that results in progressively worsening ability to focus clearly on close objects. Also known as age-related farsightedness (or age-related long si ...
(the loss of focusing range during aging).
Macular degeneration
Macular degeneration is especially prevalent in the U.S. and affects roughly 1.75 million Americans each year. Having lower levels of lutein and zeaxanthin within the macula may be associated with an increase in the risk of age-related macular degeneration. < Lutein and zeaxanthin act as antioxidants that protect the retina and macula from oxidative damage from high-energy light waves. As the light waves enter the eye they excite electrons that can cause harm to the cells in the eye, but they can cause oxidative damage that may lead to macular degeneration or cataracts. Lutein and zeaxanthin bind to the electron free radical and are reduced rendering the electron safe. There are many ways to ensure a diet rich in lutein and zeaxanthin, the best of which is to eat dark green vegetables including kale, spinach, broccoli and turnip greens. Nutrition is an important aspect of the ability to achieve and maintain proper eye health.
zeaxanthin
Zeaxanthin is one of the most common carotenoids in nature, and is used in the xanthophyll cycle. Synthesized in plants and some micro-organisms, it is the pigment that gives paprika (made from bell peppers), corn, saffron, goji ( wolfberries ...
are two major carotenoids, found in the macula of the eye, that are being researched to identify their role in the pathogenesis of eye disorders such as age-related
macular degeneration
Macular degeneration, also known as age-related macular degeneration (AMD or ARMD), is a medical condition which may result in blurred or no vision in the center of the visual field. Early on there are often no symptoms. Over time, however, som ...
and
cataracts
A cataract is a cloudy area in the lens of the eye that leads to a decrease in vision. Cataracts often develop slowly and can affect one or both eyes. Symptoms may include faded colors, blurry or double vision, halos around light, trouble w ...
File:Diagram of human eye without labels.svg, Right eye without labels (horizontal section)
File:Blausen 0388 EyeAnatomy 01.png
File:Blausen 0389 EyeAnatomy 02.png
File:Three Main Layers of the Eye.png, The structures of the eye labeled
File:Three Internal chambers of the Eye.svg, Another view of the eye and the structures of the eye labeled
See also
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Eye colour
Eyes are organs of the visual system. They provide living organisms with vision, the ability to receive and process visual detail, as well as enabling several photo response functions that are independent of vision. Eyes detect light and conv ...
*
Eye development
Eye formation in the human embryo begins at approximately three weeks into embryonic development and continues through the tenth week.Ort, D., David, H."Development of the Eye" Retrieved 22 April 2015. Cells from both the mesodermal and the ect ...
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Eye disease
This is a partial list of human eye diseases and disorders.
The World Health Organization publishes a classification of known diseases and injuries, the International Statistical Classification of Diseases and Related Health Problems, or ICD-10. ...
Hyaloid canal
Hyaloid canal (Cloquet's canal and Stilling's canal) is a small transparent canal running through the vitreous body from the optic nerve disc to the lens. It is formed by an invagination of the hyaloid membrane, which encloses the vitreous body ...
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Iris recognition
Iris recognition is an automated method of biometric identification that uses mathematical pattern-recognition techniques on video images of one or both of the irises of an individual's eyes, whose complex patterns are unique, stable, and can ...
*
Knobloch syndrome
Knobloch syndrome is a rare genetic disorder presenting severe eyesight problems and often a defect in the skull. It was named after the ophthalmologist William Hunter Knobloch (1926 - 2005), who first described the syndrome in 1971. A usual occur ...
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Lacrimal caruncle
The lacrimal caruncle, or caruncula lacrimalis, is the small, pink, globular nodule at the inner corner (the medial canthus) of the eye. It consists of tissue types of neighbouring eye structures. It may suffer from lesions and allergic inflammati ...
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Rheum
Rheum (; from Greek: ῥεῦμα ''rheuma'' 'a flowing, rheum') is a thin mucus naturally discharged from the eyes, nose, or mouth, often during sleep (cf. mucopurulent discharge).Amodio, Aime"Where Do Eye Boogers Come From?" Families.com bl ...
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Spectral sensitivity
Spectral sensitivity is the relative efficiency of detection, of light or other signal, as a function of the frequency or wavelength of the signal.
In visual neuroscience, spectral sensitivity is used to describe the different characteristics o ...