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Hickory is a common name for trees composing the genus ''Carya'', which includes around 18 species. Five or six species are native to
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, Indochina, and India ( Assam), as many as twelve are native to the United States, four are found in Mexico, and two to four are native to Canada. A number of hickory species are used for products like edible nuts or wood. Hickories are temperate forest trees with
pinnately Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
compound leaves and large
nuts Nut often refers to: * Nut (fruit), fruit composed of a hard shell and a seed, or a collective noun for dry and edible fruits or seeds * Nut (hardware), fastener used with a bolt Nut or Nuts may also refer to: Arts, entertainment, and media Com ...
. Hickory flowers are small, yellow-green
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in ''Salix''). They contain many, usually unisexual flowers, arranged cl ...
s produced in spring. They are wind-pollinated and self-incompatible. The fruit is a globose or oval nut, long and diameter, enclosed in a four- valved
husk Husk (or hull) in botany is the outer shell or coating of a seed. In the United States, the term husk often refers to the leafy outer covering of an ear of maize (corn) as it grows on the plant. Literally, a husk or hull includes the protective ...
, which splits open at maturity. The nut shell is thick and bony in most species, and thin in a few, notably the pecan (''C. illinoinensis''); it is divided into two halves, which split apart when the seed
germinates Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an flowering plant, angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spor ...
.


Etymology

The name "hickory" derives from a Native American word in an Algonquian language (perhaps Powhatan). It is a shortening of ''pockerchicory'', ''pocohicora'', or a similar word, which may be the name for the hickory tree's nut, or may be a milky drink made from such nuts. The genus name ''Carya'' is grc, κάρυον, ''káryon'', meaning " nut".


Species and classification

The genus ''Carya'' is in the walnut family, Juglandaceae. In the APG system, this family is included in the order Fagales. Several species are known to hybridize, with around nine accepted, named hybrids. Beaked hickory (''Annamocarya sinensis'') is a species formerly classified as ''Carya sinensis'', but now considered its own genus, ''
Annamocarya ''Annamocarya'' is a genus of flowering plants in the family Juglandaceae, containing only one species, ''Annamocarya sinensis'', native to southwestern China (Guangxi, Guizhou, Yunnan) and northern Vietnam. It is related to the hickories, and w ...
''.


Asian hickories

''Carya'' sect. ''Sinocarya'' *''
Carya dabieshanensis Hickory is a common name for trees composing the genus ''Carya'', which includes around 18 species. Five or six species are native to China, Indochina, and India (Assam), as many as twelve are native to the United States, four are found in Mexi ...
'' M.C. Liu – Dabie Shan hickory (may be synonymous with ''C. cathayensis'') *''
Carya cathayensis ''Carya cathayensis'' () ( common name Chinese hickory) is a species of hickory native to China. Peeled and roasted nuts, known as 山核桃仁, are a well-known specialty of Hangzhou Hangzhou ( or , ; , , Standard Mandarin pronunciatio ...
'' Sarg. – Chinese hickory *'' Carya hunanensis'' W.C.Cheng & R.H.Chang – Hunan hickory *'' Carya kweichowensis'' Kuang & A.M.Lu – Guizhou hickory *'' Carya poilanei'' Leroy – Poilane's hickory *''
Carya tonkinensis ''Carya tonkinensis'', the Vietnam hickory, is a species of flowering plant in the genus ''Carya Hickory is a common name for trees composing the genus ''Carya'', which includes around 18 species. Five or six species are native to China, Ind ...
'' Lecomte – Vietnamese hickory


North American hickories

''Carya'' sect. ''Carya'' – typical hickories *'' Carya floridana'' Sarg. – scrub hickory *'' Carya glabra'' (Mill.) Sweet – pignut hickory, pignut, sweet pignut, coast pignut hickory, smoothbark hickory, swamp hickory, broom hickory *'' Carya laciniosa'' (Mill.) K.Koch – shellbark hickory, shagbark hickory, bigleaf shagbark hickory, kingnut, big shellbark, bottom shellbark, thick shellbark, western shellbark *''
Carya myristiciformis ''Carya myristiciformis'', the nutmeg hickory, a tree of the Juglandaceae or walnut family, also called swamp hickory or bitter water hickory, is found as small, possibly relict populations across the Southern United States and in northern Mexic ...
'' ( F.Michx.) Nutt. – nutmeg hickory, swamp hickory, bitter water hickory *''
Carya ovalis ''Carya ovalis'', the red hickory or sweet pignut hickory, is a fairly uncommon but widespread hickory native to eastern North America. It is typically found growing in dry, well drained sandy upland ridges and sloped woodlands from southern Onta ...
'' (Wangenh.) Sarg. – red hickory, spicebark hickory, sweet pignut hickory (treated as a synonym of ''C. glabra'' by ''Flora N. Amer.'') *''
Carya ovata ''Carya ovata'', the shagbark hickory, is a common hickory in the Eastern United States and southeast Canada. It is a large, deciduous tree, growing well over tall, and can live more than 350 years. The tallest measured shagbark, located in Sav ...
'' (Mill.) K.Koch – shagbark hickory **''C. o.'' var. ''ovata'' – northern shagbark hickory **''C. o.'' var. ''australis'' – southern shagbark hickory, Carolina hickory (syn. ''C. carolinae-septentrionalis'') *''
Carya pallida ''Carya pallida'', sand hickory, or pale hickory is a species of hickory native to the southeastern United States. It is a perennial, dicotyledonous plant which prefers rocky or sandy habitats. The sand hickory can reach heights of up to 30m, b ...
'' (Ashe) Engl. & Graebn. – sand hickory *'' Carya texana'' Buckley – black hickory *'' Carya tomentosa'' (Poir.) Nutt. – mockernut hickory (syn. ''C. alba'') *
A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species). It is one of the modern descendan ...
'' Carya washingtonensis'' Manchester – Miocene of Kittitas County, Washington ''Carya'' sect. ''Apocarya'' – pecans *''
Carya aquatica ''Carya aquatica'', the bitter pecan or water hickory, is a large tree, that can grow over tall of the Juglandaceae or walnut family. In the American South it is a dominant plant species found on clay flats and backwater areas near streams and r ...
'' (F.Michx.) Nutt. – bitter pecan or water hickory *'' Carya cordiformis'' (Wangenh.) K.Koch – bitternut hickory *'' Carya illinoinensis'' (Wangenh.) K.Koch – pecan *''
Carya palmeri ''Carya palmeri'', the Mexican hickory, is a tree species native to Mexico. It was first described by Wayne Eyer Manning. ''Carya palmeri'' is part of the genus '' Carya'', and the family Juglandaceae. Range ''Carya palmeri'' is endemic to Mexi ...
'' W.E. Manning – Mexican hickory


Ecology

Hickory is used as a food plant by the
larvae A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
of some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
species. These include: * Luna moth (''
Actias luna The Luna moth (''Actias luna'') also known as the American moon moth is a Nearctic moth in the family Saturniidae, subfamily Saturniinae, a group commonly known as giant silk moths. It has lime-green colored wings and a white body. The larvae ( ...
'') *
Brown-tail moth The brown-tail moth (''Euproctis chrysorrhoea'') is a moth of the family Erebidae. It is native to Europe, neighboring countries in Asia, and the north coast of Africa. Descriptions of outbreaks, i.e., large population increases of several years ...
(''Euproctis chrysorrhoea'') * '' Coleophora'' case-bearers, ''C. laticornella'' and ''C. ostryae'' *
Regal moth ''Citheronia regalis'', the regal moth or royal walnut moth, is a North American moth in the family Saturniidae. The caterpillars are called hickory horned devils. The adult (imago) has a wingspan of . The species was first described by Johan Chri ...
s (''Citheronia regalis''), whose caterpillars are known as hickory horn-devils * Walnut sphinx (''Amorpha juglandis'') * The bride (
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
'' Catocala neogama neogama'') * Hickory tussock moth (''Lophocampa caryae'') The hickory leaf stem gall phylloxera (''
Phylloxera Grape phylloxera is an insect pest of commercial grapevines worldwide, originally native to eastern North America. Grape phylloxera (''Daktulosphaira vitifoliae'' (Fitch 1855) belong to the family Phylloxeridae, within the order Hemiptera, bugs ...
caryaecaulis'') also uses the hickory tree as a food source. Phylloxeridae are related to aphids and have a similarly complex life cycle. Eggs hatch in early spring and the
gall Galls (from the Latin , 'oak-apple') or ''cecidia'' (from the Greek , anything gushing out) are a kind of swelling growth on the external tissues of plants, fungi, or animals. Plant galls are abnormal outgrowths of plant tissues, similar to be ...
s quickly form around the developing insects. ''Phylloxera'' galls may damage weakened or stressed hickories, but are generally harmless. Deformed leaves and twigs can rain down from the tree in the spring as
squirrel Squirrels are members of the family Sciuridae, a family that includes small or medium-size rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrels. Squ ...
s break off infected tissue and eat the galls, possibly for the protein content or because the galls are fleshy and tasty to the squirrels. The pecan gall curculio ('' Conotrachelus elegans'') is a true weevil species also found feeding on galls of the hickory leaf stem gall phylloxera. The
banded hickory borer ''Knulliana'' is a genus of longhorn beetles. It is monotypic, being represented by the single species ''Knulliana cincta'', commonly known as the banded hickory borer. The species may be found throughout the eastern half of North America, includ ...
(''Knulliana cincta'') is also found on hickories.


Evolutionary history

The oldest fossils attributed to ''Carya'' are Cretaceous pollen grains from Mexico and New Mexico. Fossil and molecular data suggest the genus ''Carya'' may have diversified during the Miocene. Modern ''Carya'' first appear in
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
strata 34 million years ago. Recent discoveries of ''Carya'' fruit fossils further support the hypothesis that the genus has long been a member of Eastern North American landscapes, however its range has contracted and Carya is no longer extant west of the Rocky Mountains. Fossils of early hickory nuts show simpler, thinner shells than modern species with the exception of pecans, suggesting that the trees gradually developed defenses to rodent seed predation. During this time, the genus had a distribution across the Northern Hemisphere, but the Pleistocene Ice Age beginning 2 million years ago obliterated it from Europe. The distribution of Carya in North America also contracted and it completely disappeared from the continent west of the Rocky Mountains. Since fossil records show North America as having the largest number of Juglandaceae species, it is likely that the genus originated there and later spread to Europe and Asia.


Fruit

Some fruits are borderline and difficult to categorize. Hickory nuts (''Carya'') and walnuts ('' Juglans'') in the Juglandaceae family grow within an outer husk; these fruits are sometimes considered to be
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
s or drupaceous nuts, rather than true botanical nuts. "Tryma" is a specialized term for such nut-like drupes.


Nutrition

Dried hickory nuts are 3% water, 18% carbohydrates, 13% protein, and 64% fats (table). In a 100 gram (3.5 oz) reference amount, dried hickory nuts supply 657
calorie The calorie is a unit of energy. For historical reasons, two main definitions of "calorie" are in wide use. The large calorie, food calorie, or kilogram calorie was originally defined as the amount of heat needed to raise the temperature of on ...
s, and are a rich source (20% or more of the Daily Value, DV) of several B vitamins and
dietary mineral In the context of nutrition, a mineral is a chemical element required as an essential nutrient by organisms to perform functions necessary for life. However, the four major structural elements in the human body by weight (oxygen, hydrogen, carbon ...
s, especially manganese at 220% DV (table).


Uses

Hickory wood is hard, stiff, dense and shock resistant. There are woods stronger than hickory and woods that are harder, but the combination of strength, toughness, hardness, and stiffness found in hickory wood is not found in any other commercial wood. It is used for tool handles, pickaxe handles, bows,
wheel A wheel is a circular component that is intended to rotate on an axle Bearing (mechanical), bearing. The wheel is one of the key components of the wheel and axle which is one of the Simple machine, six simple machines. Wheels, in conjunction wi ...
spokes,
cart A cart or dray (Australia and New Zealand) is a vehicle designed for transport, using two wheels and normally pulled by one or a pair of draught animals. A handcart is pulled or pushed by one or more people. It is different from the flatbed tr ...
s, drumsticks, lacrosse stick handles, golf club shafts (sometimes still called ''hickory stick'', even though made of
steel Steel is an alloy made up of iron with added carbon to improve its strength and fracture resistance compared to other forms of iron. Many other elements may be present or added. Stainless steels that are corrosion- and oxidation-resistant ty ...
or graphite), the bottoms of skis, walking sticks, and for punitive use as a switch (like
hazel The hazel (''Corylus'') is a genus of deciduous trees and large shrubs native to the temperate Northern Hemisphere. The genus is usually placed in the birch family Betulaceae,Germplasmgobills Information Network''Corylus''Rushforth, K. (1999). ...
), and especially as a cane-like hickory stick in schools and use by parents. Paddles are often made from hickory. This property of hickory wood has left a trace in some Native American languages: in Ojibwe, hickory is called ''mitigwaabaak'', a compound of ''mitigwaab'' "bow" and the final ''-aakw'' "hardwood tree". Due to its grain structure, hickory is more susceptible to moisture absorption than other species of wood, and is therefore more prone to shrinkage, warping or swelling with changes in humidity. Baseball bats were formerly made of hickory, but are now more commonly made of ash. Hickory is replacing ash as the wood of choice for Scottish
shinty Shinty ( gd, camanachd, iomain) is a team game played with sticks and a ball. Shinty is now played mainly in the Scottish Highlands and amongst Highland migrants to the big cities of Scotland, but it was formerly more widespread in Scotland, an ...
sticks (also known as camans). Hickory was extensively used for the construction of early aircraft. Hickory is also highly prized for wood-burning stoves and
chiminea A chimenea , also spelled chiminea (from Spanish: ''chimenea'' which derive from French ''cheminée'', "chimney"), is a freestanding front-loading fireplace or oven with a bulbous body and usually a vertical smoke vent or chimney. History Hi ...
s, as its density and high energy content make it an efficient fuel. Hickory wood is also a preferred type for
smoking Smoking is a practice in which a substance is burned and the resulting smoke is typically breathed in to be tasted and absorbed into the bloodstream. Most commonly, the substance used is the dried leaves of the tobacco plant, which have bee ...
cured meats. In the Southern United States, hickory is popular for cooking
barbecue Barbecue or barbeque (informally BBQ in the UK, US, and Canada, barbie in Australia and braai in South Africa) is a term used with significant regional and national variations to describe various cooking methods that use live fire and smoke t ...
, as hickory grows abundantly in the region and adds flavor to the meat. Hickory is sometimes used for wood flooring due to its durability in resisting wear and character. Hickory wood is not noted for rot resistance. A extract from shagbark hickory
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, ...
is also used in an edible syrup similar to
maple syrup Maple syrup is a syrup made from the sap of maple trees. In cold climates, these trees store starch in their trunks and roots before winter; the starch is then converted to sugar that rises in the sap in late winter and early spring. Maple tree ...
, with a slightly bitter, smoky taste. The Cherokee Indians would produce a green dye from hickory bark, which they used to dye cloth. When this bark was mixed with maple bark, it produced a yellow dye pigment. The ashes of burnt hickory wood were traditionally used to produce a strong lye (potash) fit for soapmaking. The nuts of some species are palatable and were used by Cherokee Indians in making Kanuchi soup, while others are bitter and only suitable for animal feed. Shagbark and shellbark hickory, along with pecan, are regarded by some as the finest nut trees. Pecans are the most important nut tree native to North America. When cultivated for their nuts, clonal ( grafted) trees of the same cultivar cannot pollinate each other because of their self-incompatibility. Two or more cultivars must be planted together for successful pollination. Seedlings (grown from hickory nuts) will usually have sufficient genetic variation.


Gallery

File:Carya nuts.jpg, Comparison of North American ''Carya'' nuts File:Hickory nuts 6060.JPG, Ripe hickory nuts ready to fall File:2014-11-02 14 36 58 Hickory foliage during autumn along Woosamonsa Road in Hopewell Township, New Jersey.jpg, Autumn foliage


See also

* Hican * Walnut


References

*Philips, Roger. ''Trees of North America and Europe''. Random House, Inc., New York. , 1979.


External links


''Carya'' images at the Arnold Arboretum of Harvard University Plant Image Database
* Damery, Jonathan
"The ''Carya'' Collection."
''Arnold Arboretum of Harvard University website.'' Accessed 26 May 2020. {{Taxonbar, from=Q142788 * Edible nuts and seeds Extant Oligocene first appearances Native American cuisine of the Southeastern Woodlands Plant dyes Plants used in Native American cuisine Wood