Heterometrus Laevigatus
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''Heterometrus'', whose members are also known by the collective vernacular name giant forest scorpions, is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of scorpions belonging to the family
Scorpionidae The Scorpionidae (burrowing scorpions or pale-legged scorpions) make up the Taxonomic rank, superfamily Scorpionoidea. The family was established by Pierre André Latreille, 1802. Genera Scorpionidae contains the following genera: * ''Aops'' V ...
. It is distributed widely across
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
southeastern Asia, including
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,
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,
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
,
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(
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), and
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(
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). It is notable for containing some of the largest living
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of scorpions.


Taxonomy

The genus was introduced by
C.G. Ehrenberg Christian Gottfried Ehrenberg (19 April 1795 – 27 June 1876) was a German Natural history, naturalist, zoologist, comparative anatomist, geologist, and microscopy, microscopist. Ehrenberg was an Evangelicalism, evangelist and was considered to ...
(in Hemprich & Ehrenberg, 1828), originally as a
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
of the genus ''
Buthus ''Buthus'' is a genus of scorpion belonging and being eponymous to the family Buthidae. It is distributed widely across northern Africa, including Morocco, Mauritania, Algeria, Tunisia, Libya, Egypt, Senegal, Guinea-Bissau, Nigeria, Sudan, Somali ...
''. It was elevated to genus rank by F. Karsch in 1879. H.W.C. Couzijn (1978, 1981) subdivided the genus into several
subgenera In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
, but F. Kovařík (2004)
synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
ized these subgenera with the nominal genus. In 2020 the genus was reviewed by L. Prendini & S. F. Loria, three of the former subgenera were revalidated and elevated to genera and one valid subgenus was elevated to genus rank, species were transferred to appropriate genera, resulting in 28 new combinations.


Diversity

The content of this genus may vary, depending on the authority. Eight species are known, many of which are quite similar in appearance: *'' Heterometrus glaucus'' (Thorell, 1876) *'' Heterometrus laevigatus'' (Thorell, 1876) *''
Heterometrus laoticus ''Heterometrus laoticus'' or Vietnam forest scorpion, is a scorpion species found in peaty areas of Vietnam and Laos. They can reach lengths of . They are a communal species, but cannibalism has been known to occur, and if caught, they can be ex ...
'' Couzijn, 1981 *'' Heterometrus longimanus'' (Herbst, 1800) *''
Heterometrus petersii ''Heterometrus'', whose members are also known by the collective vernacular name giant forest scorpions, is a genus of scorpions belonging to the family Scorpionidae. It is distributed widely across tropical and subtropical southeastern Asia, ...
'' (Thorell, 1876) *''
Heterometrus silenus ''Heterometrus'', whose members are also known by the collective vernacular name giant forest scorpions, is a genus of scorpions belonging to the family Scorpionidae. It is distributed widely across tropical and subtropical southeastern Asia, ...
'' (Simon, 1884) *''
Heterometrus spinifer ''Heterometrus spinifer'', the Asian forest scorpion, giant blue scorpion, or giant forest scorpion, is a species of scorpions belonging to the family Scorpionidae. Description ''H. spinifer'' can reach a length around . The body is shiny blac ...
'' (Ehrenberg, 1828) *''
Heterometrus thorellii ''Heterometrus'', whose members are also known by the collective vernacular name giant forest scorpions, is a genus of scorpions belonging to the family Scorpionidae. It is distributed widely across tropical and subtropical southeastern Asia, ...
'' (Pocock, 1897)


General characteristics

Members of ''Heterometrus'' are generally large-sized
scorpion Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always end ...
s (100–200 mm or about 4-8  in total length). Coloration is dark in most species, often uniformly brown or black, sometimes with a greenish shine, with brighter-colored
telson The telson () is the posterior-most division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment on accou ...
, walking legs, and/or pedipalp pincers in some species. The scorpions are heavily built with especially powerful and globose pedipalp pionkes, broad
mesosoma The mesosoma is the middle part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the metasoma. It bears the legs, and, in the case of winged insects, the wings. In hymenopterans of t ...
l tergites and a proportionally slender and thin
metasoma The metasoma is the posterior part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the mesosoma. In insects, it contains most of the digestive tract, respiratory system, and circula ...
. The telson is proportionally small and the stinger is often shorter than the vesicle. The
cephalothorax The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cepha ...
and mesosoma are largely devoid of carinae and granulation and the median eyes are situated in a small, lenticular depression on the cephalothorax. Some species are parthenogenic. Orthobothriotaxy type C. Pedipalp femur with three trichobothria and pedipalp patella consists with 19 trichobothria. Pedipalp chela with 26 trichobothria. Retrolateral pedal spurs are absent. Stridulatory organ is located on the opposing surfaces of pedipalp coxa and first leg.


Toxicity

As in other genera of the Scorpionidae, the symptoms from ''Heterometrus'' envenomations are rather mild and no human fatalities are known. The sting causes local pain,
inflammation Inflammation (from la, wikt:en:inflammatio#Latin, inflammatio) is part of the complex biological response of body tissues to harmful stimuli, such as pathogens, damaged cells, or Irritation, irritants, and is a protective response involving im ...
,
oedema Edema, also spelled oedema, and also known as fluid retention, dropsy, hydropsy and swelling, is the build-up of fluid in the body's tissue. Most commonly, the legs or arms are affected. Symptoms may include skin which feels tight, the area ma ...
, swelling, and redness of the skin, lasting for hours to a few days. Plant extracts known in the traditional Thai medicine as natural scorpion venom antidotes are effective as symptomatic treatment of ''H. laoticus'' stings. The
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
heteroscorpine-1 was found the major component of the venom in ''H. laoticus''.


Habitat for the scorpion

Species of ''Heterometrus'' live in vegetated, often forested,
humid Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity depen ...
regions with
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
to
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologic ...
s. As most scorpions, they are predominantly
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
and hide in
burrow An Eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of sh ...
s, below logs, and in leaf litter.


In captivity

Due to their impressive size, low toxicity, and docile behavior, species of ''Heterometrus'' are popular pet scorpions. Unlike many other scorpions, they can be kept in pairs or small groups.


References


External links

* Keeping instructions for ''H. spinifer'' and ''H. laoticus''
Exotic Pets
* Images and descriptions of various species of ''Heterometrus''
Mes Scorpions
{{Taxonbar, from=Q138995 Scorpionidae Scorpion genera Taxa named by Christian Gottfried Ehrenberg